After Tsuji & Müller, 2009
<==o Procolophonomorpha Romer, 1964 sensu Lee, 1995 [Ankyramorpha deBraga & Reitz, 1996] |?- †Australothyris smithi Modesto, Scott & Reisz, 2009; M.Per. SAf. `--+-- †Procolophonoidea Romer, 1956 [Procolophonia Seeley, 1888; Procolophoniformes Lee, 1995] `--+--o †Nycteroleteridae Romer, 1956 [incl. Rhipaeosauridae Chudinov, 1955] | |-- †Leptoropha novojilovi Chudinov, 1955 | |-- †Macroleter poezicus Tverdochlebova & Ivachnenko, 1984 | |--o †Rhipaeosaurus Efremov, 1940 | | |-- †R. talonophorus Chudinov, 1955 | | `-- †R. tricuspidens Efremov, 1940 | `--+-- †Bashkyroleter bashkyricus (Efremov, 1940) Ivakhnenko, 1997 [=? Nycteroleter bashkyricus] | |--o †Nycteroleter Efremov, 1938 | | |-- †N. kassini Tchudinov, 1955 | | |-- †N. ineptus Efremov, 1938 | | `-- †N. ultimus Tchudinov &: Vjuschkov, 1955 | `--+-- †“Bashkyroleter” mesensis Ivakhnenko, 1997 | `-- †Emeroleter levis Ivakhnenko, 1997 `-- †Pareiasauria Seeley, 1888 sensu Tsuji & Müller, 2009 [Hallucicrania Lee, 1995; Pareiasauriformes Lee, 1996]
After Lee, 1993, 1996, 1997, and Debraga & Reisz, 1996<==o Procolophonomorpha |?- †Australothyris |--o †Lanthanosuchoidea | |--o †Acleistorhinidae | | |?- †Colobomycter | | `-- †Acleistorhinus | `--o †Lanthanosuchidae | |-- †Lanthaniscus | |-- †Lanthanosuchus | `-- †Chalcosaurus `--o Procolophonomorpha |--o †Nyctiphruretia | |?- †Tokosaurus | |-- †Nyctiphruretus | `--o †Nycteroleteridae | |?- †Rhipaeosaurus | |-- †Nycteroleter | `-- †Macroleter `--o Procolophonia |-- †Procolophonoidea Romer, 1956 `-- Hallucicrania |
Procolophonomorpha after Modesto, Scott & Reisz, 2009:<==o Procolophonomorpha |-- †Australothyris smithi `--o †Ankyramorpha |-- †Lanthanosuchoidea `--+-- †Bolosauridae |-- †Nyctiphruretia `--+-- †Nyctiphruretia `-- Procolophonia Seeley, 1888 [incl. Testudinata?] |
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