After Lee, 1993, 1996, and 1997 (fig. 21), and Tsuji & Müller, 2009
<==o Pareiasauroidea Nopcsa, 1928 |?-o †Sclerosauridae [maybe aberrant procolophonids (Sues & Baird, 1998, deBraga, 2003)] | |?- †Basileosaurus freyi Wiedersheim, 1879 | `-- †Sclerosaurus armatus v. Meyer, 1859 [Aristodesmus] `--o Pareiasauria Seeley, 1888 [Pareiasauridae Cope, 1896 sensu lato] `==o †Bradysaurus Watson, 1914b [Brachypareia; Bradysuchus; Koalemasaurus; Platyoropha] [Bradysauridae von Huene, 1948] U.Perm. |?- †Bradysaurus rogersi (Broom, 1912) [Propappus rogersi Broom, 1912; Brachypareia rogersi (Broom, 1912); Bradysaurus rogersi (Broom, 1912); Pareiasaurus serridens Owen, 1876] |?- †Bradysaurus acutirostris [Koalemasaurus] |?- †Bradysaurus whaitsi |-- †Bradysaurus baini (Seeley, 1892) Watson, 1914b [Pareiasaurus baini Seeley, 1892; incl. Platyoropha broomi Haughton & Boonstra, 1929] `--+-- †“Bradysaurus” seeleyi Haughton & Boonstra, 1929 [incl. Bradysaurus vanderbyli Haughton & Boonstra, 1929] `--o Velosauria Lee, 1994 sensu Lee, 1997 |-- †Nochelosaurus alexandri Haughton & Boonstra, 1929 [Nochelesaurus (lapsus calami)] U.Perm. `--+-- †Embrithosaurus schwarzi Watson, 1914 [Dolichopareia, Nochelesaurus] `--+-- †Bunostegos akokanensis Sidor, Blackburn & Gado, 2003; U.Perm. WCAf. `--+-- †Deltavjatia vjatkensis (Hartmann-Weinberg, 1937) Lebedev in Ivakhnenko, 1987 [taxonomic history] U.Perm. EEu. `--o Velosauria Lee, 1994 sensu Jalil & Janvier, 2005 `--o †Pareiasauridae Cope, 1896 sensu stricto? [incl. Elginiidae Cope, 1895] |?- †Pareiasauria gen. et sp. indet. 1 [Argana Basin, Marocco] [Jalil & Janvier, 2005] |?- †Pareiasauria gen. et sp. indet. 2 [Argana Basin, Marocco] [Jalil & Janvier, 2005] |-- “Welgevonden pareiasaur” `--+--+-- †Shansisaurus xuecunensis Cheng, 1980 [Huanghesaurus liulinensis Gao, 1983] U.Perm. EAs. | `--+-- †Shihtienfenia permica Young & Yeh, 1963; U.Perm. CAs. | `--o †Pareiasuchus Broom & Haughton, 1913; U.Perm. SAf. | |-- †P. peringueyi Broom & Haughton, 1913 | `-- †P. nasicornis Haughton & Boonstra, 1929 `--+-- †Sanchuansaurus pygmaeus Gao, 1989 (sedis mutabilis); U.Perm. CAs. |?- †Obirkovia gladiator Bulanov & Jashina, 2005 |--o †Pareiasaurus Owen, 1876; U.Perm. [Pareiasuchus Lydekker, 1890; Propappus Seeley, 1888; ?Pareiasuchus Broom & Haughton, 1913] | |?- †P. henneni Case, 1917 | |?- †P. pinnatus Olson & Broom, 1937 | |?- †P. russouwi Broom, 1913 | |?- †P. strubeni | `-- †P. serridens Owen, 1876 [Propappus serridens (Owen, 1876); Propappus omocratus Seeley, 1888; Pareiasaurus omocratus (Seeley, 1888); Pareiasaurus minor Seeley, 1892; Anthodon gregoryi Broom, 1930; Propappus rogersi Broom, 1912; Brachypareia rogersi (Broom, 1912); Bradysaurus rogersi (Broom, 1912); Pareiasaurus pinnatus Olson & Broom, 1937; Pareiasaurus rubidgei Broom, 1940; Anthodon nesemanni Broom, 1940] `--+-- †Scutosaurus karpinskii (Amalitskii, 1922) Hartmann-Weinberg, 1930 [Amalitzkia; Proelginia permiana]; U.Perm. EEu. `--+-- †Arganaceras vacanti Jalil & Janvier, 2005; U.Perm. NWAf. |-- †Parasaurus geinitzi von Meyer, 1857 (non Marsh, 1885) U.Perm. |-- “Kupferschiefer pareiasaur”; U.Perm. |--o †Elginia Newton, 1893; U.Perm. WEu. | |?- †E. gordoni | `-- †E. mirabilis Newton, 1893 `--o Pumiliopareiasauria Lee |-- †Nanoparia luckhoffi Broom, 1936 [Nanopareia (lapsus calami)]; U.Perm. |-- †Provelosaurus americanus (Araújo, 1985) [Pareiasaurus americanus Araújo, 1985] U.Perm. SAm. `--+--o †Anthodon Owen, 1876; U.Perm. | |?- †A. gregoryi Broom, 1930 [=? Pareiasaurus serridens Owen, 1876] | `-- †A. serrarius Owen, 1876 `-- †Pumilopareia pricei Broom & Robinson, 1948 [“Nanoparia” pricei]; U.Perm.
After Tsuji et al., 2013<==o Pareiasauria |-- †“Bradysaurus” seeleyi `--+-- †Nochelesaurus |-- †Bradysaurus baini `--+-- †Embrithosaurus `--+-- †Bunostegos `--+--+-- †Deltavjatia | `-- †Parasaurus `--o Velosauria |--o Pumiliopareiasauria | |-- †Nanoparia | |-- †Provelosaurus | `--+-- †Anthodon | `-- †Pumilopareia `--o †Pareiasauridae |--+-- †Shansisaurus | `--+-- †Shihtienfenia | `-- †Pareiasuchus `--+-- †Sanchuansaurus |-- †Pareiasaurus |-- †Scutosaurus |-- †Arganaceras |-- †Obirkovia `-- †Elginia Reference(s):
|
After Lee, 1993, 1996, and 1997 (fig. 21), and Jalil & Janvier, 2005<==o Hallucicrania |?-o †Lathanosuchoidea | |--o †Acleistorhinidae | | |-- †Colobomycter | | `-- †Acleistorhinus | `--o †Lanthanosuchidae | |-- †Chalcosaurus | |-- †Lanthaniscus | `-- †Lanthanosuchus `--o Pareiasauroidea |?-o †Sclerosauridae | |?- †Basileosaurus | `-- †Sclerosaurus `--o Pareiasauria |-- †“Bradysaurus” seeleyi `--+-- †Bradysaurus baini `--o Velosauria |-- †Nochelesaurus `--+-- †Embrithosaurus `--+-- †Bunostegos `--+-- †Deltavjatia `--o Velosauria |--o †Pareiasauridae | |--+-- †Shansisaurus | | `--+-- †Shihtienfenia | | `-- †Pareiasuchus | `--+?- †Sanchuansaurus (sedis mutabilis) | |-- †Pareiasaurus | `--+-- †Scutosaurus | `--+-- †Arganaceras | |-- †Parasaurus | |-- “Kupferschiefer pareiasaur” | `-- †Elginia `--+-- †Pareiasauria gen. et sp. indet. 1 [Argana Basin, Marocco] [Jalil & Janvier, 2005] `--o Pumiliopareiasauria |?- †Nanoparia |-- †Provelosaurus `--+-- †Anthodon `--+-- †Pumilopareia `-- Chelonia |
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Jalil, N.-E. & Janvier, P., 2005: Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d'Argana, Maroc.
[The Late Permian pareiasaurs (Amniota, Parareptilia) from the Argana basin, Marocco].
–Geodiversitas: Vol. 27, #1, pp. 35-132 - Lee, M. S. Y., 1993: The origin of the turtle body plan: Bridging a famous morphological gap.
–Science: Vol. 261, 24 September, 1716-1720 - Lee, M. S. Y., 1996: Correlated progression and the origin of turtles.
–Nature: Vol. 379, 29 February, pp. 812-815 - Lee, M. S. Y., 1997: Pareiasaur phylogeny and the origin of turtles.
–Zoological Journal of the Linnean Society: Vol. 120, pp. 197-280 - Lee, M. S. Y., 2000: The Russian pareiasaurs. 71-85
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000, xxxix-696 - Meylan, P. A. & Ganko, E., 1997: Higher Relationships of Testudines – The extinct and living turtles.
–iNet: The Tree of Life (proposed page): http://www.eckerd.edu/turtles/testudines.html - Sues, H.-D. & Baird, D., 1998: Procolophonidae (Reptilia: Parareptilia) from the Upper Triassic Wolfville Formation of Nova Scotia, Canada.
–Journal of Vertebrate Paleontology: Vol. 18, #3, pp. 525-532 - Sukhanov, V. R., 2000: Mesozoic turtles of Middle and Central Asia. 309-367
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000, xxxix-696 - Tsuji, L. A. & Müller, J., 2009: Assembling the history of the Parareptilia: phylogeny, diversification, and a new definition of the clade.
–Fossil Record: Vol. 12, #1, pp. 71-81 [doi: 10.1002/mmng.200800011] - Tsuji, L. A., Sidor, C. A., Steyer, J. S., Smith, R. M. H., Tabor, N. J. & Ide, O., 2013: The vertebrate fauna of the Upper Permian of Niger—VII. Cranial anatomy and relationships of Bunostegos akokanensis (Pareiasauria).
–Journal of Vertebrate Paleontology: Vol. 33, #4, pp. 747-763 [doi: 10.1080/02724634.2013.739537]