After Brochu, 1997, Salisbury, Molnar, Frey & Willis, 2006, and Holliday & Gardner, 2012
Family -level taxonomy of all crocodylomorphs [Crocodylia sensu lato]
<==o Eusuchia Huxley, 1875 sensu Salisbury, Molnar, Frey & Willis., 2006 |-- †Isisfordia duncani Salisbury, Molnar, Frey & Willis, 2006; E.Cret. EAust. `--o Eusuchia sensu Brochu, 1997 |--o †Allodaposuchus Nopsca, 1928; U.Cret. CEu. | |-- †A. palustris Blanco, Puértolas-Pascual, Marmi, Vila & Sellés, 2014 | `--+-- †A. subjuniperus Puértolas-Pascual, Canudo & Moreno-Azanza, 2014 | |-- †Arenysuchus gascabadiolorum | `--+-- †A. precedens Nopsca, 1928 | `-- †A. hulki Blanco, Fortuny, Vicente, Luján, García-Marçà & Sellés, 2015 |--o †Hylaeochampsidae Owen, 1875 sensu Ösi, Clark & Weishampel, 2007 | |-- †Hylaeochampsa vectiana Owen, 1874; E.Cret. WEu. | `-- †Iharkutosuchus makadii Ösi, Clark & Weishampel, 2007; L.Cret. CEu. |?-o †Planocraniidae Li, 1976 | |--o †Planocrania Li, 1976; Pal. EAs. | | |-- †P. hengdongensis Li, 1984 | | `-- †P. datangensis Li, 1976 | `--o †Boverisuchus Kuhn, 1938 sensu Brochu, 2012 [Pristichamphus Gervais, 1853 (in partim); Weigeltisuchus Kuhn, 1938] | |-- †B. magnifrons Kuhn, 1938 [Pristichamphus magnifrons; Weigeltisuchus geiseltalensis Kuhn, 1938] | `-- †B. vorax (Troxell, 1925) [Crocodylus vorax Troxell, 1925; Pristichampsus vorax (Troxell, 1925) Langston, 1975] `--o Crocodylia Laurenti, 1768 [non Gmelin, 1788] sensu Brochu, 1997 (Recent crocodiles; nykykrokotiilit) |--o †Aegyptosuchidae Kuhn, 1936 | |-- †Aegyptosuchus peyeri Stromer, 1933; L.Cret. NEAf. | `-- †Aegisuchus witmeri Holliday & Gardner, 2012; L.Cret. NEAf. |--+--o †Borealosuchus Brochu, 1997; L.Cret. Pal. NA. [Leidyosuchus] | | |-- †B. sternbergii (Gilmore, 1910) [Leidyosuchus sternbergii Gilmore, 1910] | | `--+-- †B. griffithi Wu, Brinkman, & Fox, 2001 | | `--+-- †B. formidabilis (Erickson, 1976) [Leidyosuchus formidabilis Erickson, 1976] | | `--+-- †B. acutidentatus (Sternberg, 1932) [Leidyosuchus acutidentatus Sternberg, 1932] | | `-- †B. wilsoni (Mook, 1959) [= Diplocynodon stuckeri Mook, 1960] | `--o Gavialoidea Hay, 1930 sensu Brochu, 1997 | |== “Thoracosaurs” [paraphyletic basal gavialoids, how ever, see Lee & Yates (2018)] | `-- Gavialidae Hay, 1930 (gavials; graviaalit) `--o Brevirostres von Zittel, 1890 sensu Holliday & Gardner, 2012 |?-o †Pristichamphus Gervais, 1853 [Pristichampsidae Kuhn, 1968] | |-- †P. rollinatii (Gray, 1831) [Crocodilus rollianti Gray, 1831; Pristichamphus rollinati (Gray) Gervais, 1853] (nomen dubium) | |-- †P. birjukovi Efimov, 1988 | `-- †P. kuznetzovi Efimov, 1988 `--o Brevirostres von Zittel, 1890 sensu Brochu, 1997 |-- Alligatoroidea Gray, 1844 sensu Brochu, 1997 (alligaattorinsukuiset krokotiilit, alligaattorit ja kaimaanit; alligators & caimans) `-- Crocodyloidea Cuvier, 1807 sensu Brochu, 1997 (krokotiilinsukuiset krokotiilit)
Eusuchia incertae sedis:
†Leidyosuchus riggsi Schmidt, 1938; Pal. NA.
†Limnosaurus sp. (nomen dubium)
†Helmstedtisuchus freyi Schleich and Windolf, 1994; Eoc. CEu. [a scombrid fish]
†Dolichochampsa minima U. Cret. SA. [Dolichochampsidae]
After Lee & Yates, 2018<==o Eusuchia |--+-- †Allodaposuchus | `-- †Lohuecosuchus `--+--+--o †Borealosuchus | | |-- †B. sternbergii | | `--+-- †B. formidabilis | | `--+-- †B. acutidentatus | | `-- †B. wilsoni | `--o “Thoracosaurs” | |-- †Eothoracosaurus mississippiensis | `--+--+-- †Thoracosaurus neocesariensis | | `-- †Thoracosaurus macrorhynchus | `--+-- †Argochampsa krebsi | `--+-- †Eogavialis africanus (sic) | `--+-- †Eosuchus lerichei | `-- †Eosuchus minor `--+--+-- †Planocrania hengdongensis | `--+-- †Planocrania datangensis | `-- †Boverisuchus vorax `--o Crocodylia |-- Alligatoroidea `--o Crocodyloidea |-- †Prodiplocynodon langi `--+-- †“Asiatosuchus” germanicus `--+-- †“Crocodylus” affinis `--+-- †“Crocodylus” depressifrons] `--+--+-- †“Crocodylus” acer | `-- †Brachyuranochampsa eversolei `--+--+-- †Asiatosuchus clarkae | `-- †Mekosuchinae `--o Longirostres |-- Gavialidae `-- Crocodylidae sensu stricto Reference(s):
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After Blanco, Fortuny, Vicente, Luján, García-Marçà & Sellés, 2015<==o Eusuchia |--o †Hylaeochampsidae | |-- †Acynodon iberoccitanus | `--+-- †Hylaeochampsa vectiana | |-- †Iharkutosuchus makadii | `-- †Acynodon adriacus `--o Crocodylia |-- Gavialoidea `--+-- †Planocrania hengdongensis |--+-- †Planocrania datangensis | `--o †Pristichamphus | |-- †P. vorax | `-- †P. geiseltalensis |--o †Borealosuchus | |-- †B. sternbergii | `--+-- †B. formidabilis | `--+-- †B. acutidentatus | `-- †B. wilsoni |--o †Allodaposuchus | |-- †A. palustris | `--+-- †A. subjuniperus | |-- †Arenysuchus gascabadiolorum | `--+-- †A. precedens | `-- †A. hulki `--o Brevirostres |-- Alligatoroidea `-- Crocodyloidea Reference(s):
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After Brochu, 1997, and Salisbury, Molnar, Frey & Willis, 2006<==o Eusuchia |-- †Isisfordia `--o Eusuchia sensu Brochu, 1997 |-- †Allodaposuchus |--o †Hylaeochampsidae | |-- †Hylaeochampsa | `-- †Iharkutosuchus `--o Crocodylia sensu Brochu, 1997 |--+-- †Borealosuchus | `-- Gavialoidea `--+-- †Pristichamphus `--o Brevirostres |-- Alligatoroidea `-- Crocodyloidea |
Reference(s):
- Blanco, A., Fortuny, J., Vicente, A., Luján, À. H., García-Marçà, J. A. & Sellés, A. G., 2015: A new species of Allodaposuchus (Eusuchia, Crocodylia) from the Maastrichtian (Late Cretaceous) of Spain: phylogenetic and paleobiological implications.
–PeerJ: Vol. 3, pp. e1171 [doi: 10.7717/peerj.1171] - Blanco, A., Puértolas-Pascual, E., Marmi, J., Vila, B. & Sellés, A. G., 2014: Allodaposuchus palustris sp. nov. from the Upper Cretaceous of Fumanya (South-Eastern Pyrenees, Iberian Peninsula): Systematics, Palaeoecology and Palaeobiogeography of the Enigmatic Allodaposuchian Crocodylians.
–Plos One: Vol. 9, #12, pp. e115837 [doi: 10.1371/journal.pone.0115837] - Brochu, C. A. 1997: A review of “Leidyosuchus” (Crocodyliformes, Eusuchia) from the Cretaceous through Eocene of North America.
–Journal of Vertebrate Paleontology: vol. 17, #4, pp. 679-697 - Brochu, C. A., 2000: Borealosuchus (Crocodylia) from the Paleocene of Big Bend National Park, Texas.
–Journal of Paleontology: Vol. 74, #1, pp. 181-187 - Brochu, C. A., 2012: Phylogenetic relationships of Palaeogene ziphodont eusuchians and the status of Pristichampsus Gervais, 1853.
–Earth and Environmental Science Transactions of the Royal Society of Edinburgh: Vol. 103, #3-4, pp. 521-550 [doi: doi:10.1017/S1755691013000200] - Buscalioni, A. D., Ortega, F. & Vasse, D., 1997: New crocodiles (Eusuchia: Alligatoroidea) from the Upper Cretaceous of southern Europe.
–Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science: Vol. 325, #7, pp. 525-530 [doi: http://dx.doi.org/10.1016/S1251-8050(97)89872-2] - Buscalioni, A. D., Ortega, F., Weishampel, D. B. & Jianu, C. M., 2001: A revision of the crocodyliform Allodaposuchus precedens from the Upper Cretaceous of the Hateg Basin, Romania. Its relevance in the phylogeny of Eusuchia.
–Journal of Vertebrate Paleontology: Vol. 21, #1, pp. 74-86 - Clark, J. M. & Norell, M. A., 1992: The Early Cretaceous crocodylomorph Hylaeochampsa vectiana from the Wealden of the Isle of Wight.
–American Museum Novitates: #3032, pp. 1-19 - Holliday, C. M. & Gardner, N. M., 2012: A New Eusuchian Crocodyliform with Novel Cranial Integument and Its Significance for the Origin and Evolution of Crocodylia.
–Plos One: Vol. 7, #1, Art.nr. e30471, pp. (1-13) [doi: 10.1371/journal.pone.0030471] - Hua, S. & Jouve, S., 2004: A primitive marine gavialoid from the Palaeocene of Marocco.
–Journal of Vertebrate Paleontology: Vol. 24, #2, pp. 341-350 - Salisbury, S. W., Molnar, R. E., Frey, E. & Willis, P. M. A., 2006: The origin of modern crocodyliforms: new evidence from the Cretaceous of Australia.
–Proceedings of the Royal Society of London: Biological series: Vol. 273, pp. 2439-2448 - Lee, M. S. Y. & Yates, A. M., 2018: Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil record.
–Proceedings of the Royal Society B: Biological Sciences: Vol. 285, #1881, Art.nr. 20181071, pp. 1-10 [doi: 10.1098/rspb.2018.1071] - Storrs, G. W. & Efimov, M. B., 2000: Mesozoic crocodyliforms of north-central Eurasia. 402-419
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000 xxxix-696 - Zonneveld, J.-P., Gunnell, G. F. & Bartels, W. S., 2000: Early Eocene fossil vertebrates from the southwestern Green River basin, Lincoln and Uinta Counties, Wyoming.
–Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 369-386 - Wu, X.-C., Brinkman, D. B. & Fox, R. C., 2001: A new crocodylian (Archosauria) from the basal Paleocene of the Red Deer River Valley, southern Alberta.
–Canadian Journal of Earth Sciencies: Vol. 38, #12, pp. 1689-1704 - Ösi, A., Clark, J. M. & Weishampel, D. B., 2007: First report on a new basal eusuchian crocodyliform with multicusped teeth from the Upper Cretaceous (Santonian) of Hungary.
–Neues Jahrbuch für Geologie und Paläontologie, Abh.: Vol. 243, #2, pp. 169-177 [doi: 10.1127/0077-7749/2007/0243-0169]