After P.A. Willis, 1993, Brochu, 1997, 2000, 2006, 2007, 2010, and McAliley, R.E. Willis, Ray., White, Brochu & Densmore III, 2006 (fig. 6A)
<==o Crocodylinae Cuvier, 1807 (true crocodiles; aitokrokotiilit) |-- †“Crocodylus” megarhinus Andrews, 1906 [incl. “Crocodylus” articeps Andrews, 1906] |?- †Mekosuchinae Willis, Molnar & Scanlon, 1993 sensu Willis, 1993 `--+?- †Aldabrachampsus dilophus Brochu, 2006; Quat. WIOc. |--o Osteolaeminae sensu Brochu, 2007 (kääpiökrokotiilit) | |-- †Rimasuchus lloidi (Tchernov, 1986) [“Crocodylus” lloidi (Tchernov, 1986)] M.Mioc. NEAf. | |--+-- †“Crocodylus” pigotti Tchernov & Van Couvering, 1978 | | `--o †Euthecodon Fourtau, 1918 | | |-- †E. arambourgii Ginsburg and Buffetaut, 1978 | | |-- †E. nitriae Fortau, 1920 | | `-- †E. brumpti Joleaud, 1920 | `--+-- †Voay robustus (Grandidier & Vaillant, 1872) Brochu, 2007 [Crocodylus robustus Grandidier & Vaillant, 1872] (madagascarinkrokotiili; Madagascar horned crocodile) | `--o Osteolaemus Cope, 1861 | |-- O. osborni (Schmidt, 1919) Wermuth & Mertens, 1961 [Osteoblepharon osborni Schmidt, 1919] (Congo Dwarf Crocodile) | `-- O. tetraspis Cope, 1861 (West African Dwarf Crocodile; kääpiökrokotiili) `--o Crocodylinae Cuvier, 1807 sensu Brochu, 2007 [Crocodylus sensu lato] (true crocodiles; aitokrokotiilit) |?- †Crocodylus? checchiai |?- †Crocodylus bambolii Ristori, 1890; L.Mioc. SEu. |-- Mecistops cataphractus (Cuvier, 1825) Gray 1844 sensu McAliley et al., 2006 [Crocodylus cataphractus Cuvier, 1825] [incl. †C. ndokoensis] (African slender-snouted crocodile; panssarikrokotiili) `==o Crocodylus Laurenti, 1768 Mio. R. Af., Plio. R. NA. Car., Pleist. R. EAs. Au. (krokotiilit; crocodiles) [paraphyletic] |-- †C. anthropophagus Brochu, Njau, Blumenschine & Densmore, 2010; Plioc.-E.Pleist. EAf. |-- †C. palaeindicus [C. sivalensis] |-- †C. thorbjarnarsoni Brochu & Storrs, 2012 |--+-- C. suchus Geoffroy Saint-Hilaire, 1807 [C. niloticus Laurenti, 1768 in partim; Crocodylus niloticus suchus Wermuth & Mertens, 1977] (West African crocodile, länsiafrikankrokotiili) | `--+-- C. niloticus Laurenti, 1768 [C. madagascariensis Granadier, 1872 (nomen nudum)] (nile crocodile; niilinkrokotiili) | `--+-- †C. falconensis Scheyer, Aguilera, Delfino, Fortier, Carlini, Sánchez, Carrillo-Briceño, Quiroz & Sánchez-Villagra, 2013 | `--+-- C. rhombifer Cuvier, 1807 [†C. antilliensis] (kuubankrokotiili; cuban crocodile) | `--+-- C. acutus Cuvier, 1807 (amerikankrokotiili; american crocodile) | `--+-- C. intermedius Graves, 1819 (orinagonkrokotiili; orinago crocodile) | `-- C. moreletii Duméril & Bibron, 1851 (kyhmykrokotiili) `--+-- C. palustris Lesson, 1831 (suokrokotiili; Indian Mugger) `--+-- C. siamensis Schneider, 1801 [†C. ossifragus Dubois, 1908] (siaminkrokotiili) `--+-- C. porosus Meyer, 1795 [incl. C. nathani Longman, 1924] Plio. Pleist. R. Aust., R. SEAs. (suistokrokotiili; saltwater crocodile, saltie) |-- C. johnstoni Krefft, 1873; Pleist. R. Aust. (australiankrokotiili; Johnson's crocodile) |-- C. mindorensis Schmidt, 1935 (filippiinienkrokotiili) `-- C. novaeguineae Schmidt, 1928 (uudenguineankrokotiili)
Reference(s):
- Brochu, C. A. 1997: A review of “Leidyosuchus” (Crocodyliformes, Eusuchia) from the Cretaceous through Eocene of North America.
–Journal of Vertebrate Paleontology: vol. 17, #4, pp. 679-697 - Brochu, C. A., 2000: Phylogenetic relationships and devergence timing of Crocodylus based on morphology and the fossil record.
–Copeia: Vol. 100, #3, pp. 657-673 - Brochu, C. A., 2006: A new miniature horned crocodile from thw Quaternary of Aldabra Atoll, western Indian Ocean.
–Copeia: Vol. 2006 (106), #2, pp. 149-158 - Brochu, C. A., 2007: Morphology, relationships, and biogeographical significance of an extinct horned crocodile (Crocodylia, Crocodylidae) from the Quaternary of Madagascar.
–Zoological Journal of the Linnean Society: Vol. 150, #4, pp. 835-863 [doi: 10.1111/j.1096-3642.2007.00315.x] - Brochu, C. A., Njau, J., Blumenschine, R. J. & Densmore III, L. D., 2010: A new horned crocodile from the Plio-Pleistocene hominid sites at Olduvai Gorge, Tanzania.
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