After Clark, 1994, Ortega, Gasparini, Buscalioni & Calvo, 2000, and Turner & Calvo, 2005

Family -level taxonomy of all crocodylomorphs [Crocodylia sensu lato]

Alternative phylogeny of Crocodyliformes after Sereno, Larsson, Sidor & Gado, 2001

Alternative phylogeny of Crocodyliformes after Tykoski, Rowe, Ketcham & M. W. Colbert, 2002

Alternative phylogeny of Crocodyliformes after Clark, 1994 and Pol & Norell, 2004

Alternative phylogeny of Crocodyliformes after Clark, 1994, and Carvalho, Ribeiro & Avilla, 2004

<==o Crocodyliformes Hay, 1930
   `==o †Protosuchia Mook, 1934 [paraphyletic]
      |-- †Eopneumatosuchus colberti Crompton & Smith, 1980
      |?- †Microchampsa scutata Young, 1951 [nomen dubium]
      `--+?- †Artzosuchus brachicephalus Efimov, 1983 [Artzosuchidae Efimov, 1983]
         |?- Protosuchia genera et species indet.1 [North America, Kayenta Formation]
         |?- †Hadongsuchus acerdentis
         |-- †Orthosuchus strombergi Nash, 1975 [?Orthosuchidae]
         `--+?- †Platyognathus hsui Young, 1944 [Platyognathidae]
            |?-o Edentosuchidae
            |  |?- Protosuchia genera et species indet.2 [North America, Kayenta Formation]
            |  |?- Protosuchia genera et species indet.3 [North America, Kayenta Formation, UCMP unnumbered]
            |  `-- †Edentosuchus tienshanensis Young, 1973
            `==o †Protosuchidae sensu lato
               |-- †Dianosuchus changchiawaensis Young, 1982
               |-- †Erythrochampsa longipes (Broom, 1904) Haughton, 1924 [Notochampsa longipes Broom, 1904]
               |-- †Notochampsa istedana Broom 1904 [Nothochampsa (lapsus calami)] [Notochampsidae Haughton, 1924]
               |-- †Stegomosuchus longipes Emerson & Loomis, 1904
               |?- †Tagarosuchus kulemzini Efimov, 1999
               `--+--o †Protosuchidae Brown, 1934 sensu Wu, Brinkman & Lu, 1994
                  |  |-- †Hemiprotosuchus leali Bonaparte, 1972
                  |  `--o †Protosuchus Brown, 1934 [Lesothosuchus Whetstone & Whybrow, 1983; Baroqueosuchus Busbey & Gow, 1984]
                  |     |-- †P. sp.1
                  |     |-- †P. sp.2
                  |     |-- †P. micmac Sues, Shubin, Olsen & Amaral, 1996
                  |     |-- †P. richardsoni (Brown, 1933)
                  |     `-- †P. haughtoni (Whetstone & Whybrow, 1983)
                  `--+?- †“Las Hoyas crocodyliform”
                     `--+--o †Gobiosuchidae Osmólska, 1972 [moved to Protosuchia by Wu, Sues & Dong, 1997]
                        |  |-- †Zaraasuchus shepardi Pol & Norell, 2004b
                        |  `--o †Gobiosuchus Osmólska, 1972 [Gobiosuchidae] Osmólska, 1972
                        |     |-- †G. kielanae Osmólska, 1972
                        |     `-- †G. parvus Efimov, 1983
                        `--+--+-- †Zosuchus davidsoni Pol & Norell, 2004
                           |  `--+--o †Sichuanosuchus Peng, 1995; U. Jur. ?L. Cret., EAs.
                           |     |  |-- †S. huidongensis Peng, 1995
                           |     |  `-- †S. shuhanensis Wu, Sues & Dong, 1997
                           |     `--o †Shantungosuchus Young, 1961
                           |        |-- †S. chuhsienensis Young, 1961
                           |        |-- †S. brachycephalus Young, 1982 [posthumous]
                           |        `-- †S. hangjinensis Wu, Brinkman & Lu, 1994
                           |--o †Shartegosuchidae Efimov, 1988
                           |  |?- †Kyasuchus saevi Efimov & Leshchinskiy, 2000
                           |  |--o †Nominosuchus Efimov, 1996
                           |  |  |-- †N. matutinus Efimov, 1996
                           |  |  `-- †N. arcanus Kurzanov, Efimov & Gubin, 2003
                           |  `--+-- †Fruitachampsa callisoni Clark, 2012; L.Jur. WNAm.(“Fruita crocodyliform”)
                           |     `--+-- †Shartegosuchus asperopalatum Efimov, 1988
                           |        `-- †Adzhosuchus fuscus Efimov, Gubin & Kurzanov, 2000
                           `--+-- †Hsisosuchus chungkingensis (Young & Chow, 1953) [Hsisosuchidae]
                              `--o Mesoeucrocodylia Whetstone & Whybrow, 1983
                                 |-- †Notosuchia Gasparini, 1971 sensu Turner & Sertich, 2010 [incl. Ziphosuchia Ortega, Gasparini, Buscalioni & Calvo, 2000]
                                 `--+--o †Trematochampsidae
                                    |  |-- †Amargasuchus minor Chiappe, 1988
                                    |  |?- ?†Baharijodon carnosauroides Kuhn, 1936
                                    |  |-- †Caririsuchus camposi Kellner, 1987
                                    |  `--o †Trematochampsa Buffetaut, 1974
                                    |     |-- †T. taqueti Buffetaut, 1974
                                    |     `-- †T. oblita Buffetaut & Taquet, 1979
                                    `-- Neosuchia sensu Clark, 1994 (modern crocodiles; modernit krokotiilit)

Mesoeucrocodylia incertae sedis: [= Mesocrocodiles not yet reviewed]

Bottosaurus Agassiz, 1849
            †B.harlaniB. perrugosusB. tuberculatusB. perrugosusDianchungosaurus lufengensis Yang, 1982a
        †Entradasuchus spinosus Hunt & Lockley, 1995
        †Lisboasaurus estesi Seiffert, 1970 sensu Buscalioni, Pérez-Moreno & Evans, 1996 [non-archosaurian archosauromorphan? (a protorosaurian?)]
        †Lusitanisuchus mitracostatus (Seiffert, 1970) Schwarz & Fechner, 2004 [Lisboasaurus mitracostatus Seiffert, 1970]
        †Khoratosuchus jintasakuli Lauprasert et al., 2009

    †Cunampaiidae Cracraft, 1968
        †Cunampaia simplex Rusconi, 1946 [originally descriped as a prhorusrhacoid bird!!]

Alternative phylogenies of Crocodyliformes:

After Holliday & Gardner, 2012

Holliday & Gardner (2012) used Sereno & Larsson's datamatrix, with including new coding for Aegisuchus.

<==o Crocodyliformes
   |-- †Zosuchus
   `--+-- †Hsisosuchus
      `--o Mesoeucrocodylia
         |--o Thalattosuchia
         |  `--+-- †Pelagosuchus
         |     `--+-- †Teleosauridae
         |        `-- †Metriorhynchidae
         `--o Metasuchia
            |--o †Notosuchia
            |  |--+==Araripesuchus
            |  |  `--+-- †Uruguaysuchus
            |  |     `--+-- †Simosuchus
            |  |        `-- †Anatosuchus
            |  `--+-- †Baurusuchus
            |     `--+--+-- †Mariliasuchus
            |        |  `-- †Comahuesuchus
            |        `--+-- †Malawisuchus
            |           |-- †Notosuchus
            |           `-- †Sphagesaurus
            `--o Neosuchia
               |--o †Sebecia
               |  |-- †Uberabasuchus
               |  `--+-- †Hamadasuchus
               |     `--+-- †Sebecus
               |        `--+-- †Stolokrosuchus
               |           `--+-- †Peirosaurus
               |              `-- †Lomasuchus
               `--+--o †Mahajangasuchidae
                  |  |-- †Mahajangasuchus
                  |  `-- †Kaprosuchus
                  `--+--o †Pholidosauridae
                     |  |-- †Goniopholis
                     |  `--+--+-- †Dyrosaurus
                     |     |  `-- †Terminonaris
                     |     `--+-- †Sarcosuchus
                     |        `-- †Pholidosaurus
                     `--+-- †Laganosuchus
                        `--+-- †Bernissartia
                           `-- Eusuchia

Reference(s):

  • Holliday, C. M. & Gardner, N. M., 2012: A New Eusuchian Crocodyliform with Novel Cranial Integument and Its Significance for the Origin and Evolution of Crocodylia.
    †Plos One: Vol. 7, #1, pp. e30471 [doi: 10.1371/journal.pone.0030471]

After O'Connor et al., 2010

<==o Crocodyliformes
   |--+-- †Orthosuchus
   |  `--+--+-- †Hemiprotosuchus
   |     |  `-- †Protosuchus
   |     `--+-- “Kayenta Edentosuchus-like form”
   |        `-- †Edentosuchus
   `--+--+-- †Zaraasuchus
      |  `-- †Gobisuchus
      `--+--+-- †Zosuchus
         |  `--+-- †Sichuanosuchus
         |     `-- †Shantungosuchus
         `--+-- †Fruitachampsa (“Fruita form”)
            `--+-- †Hsisosuchus
               `--o Mesoeucrocodylia
                  |--o †Notosuchia
                  |  |--+-- †Araripesuchus wegeneri
                  |  |  `--+-- †Araripesuchus buitreraensis
                  |  |     `--+-- †Araripesuchus gomesii
                  |  |        `-- †Araripesuchus patagonicus
                  |  `--+-- †Uruguaysuchus
                  |     `--+--+-- †Libycosuchus
                  |        |  `--+-- †Malawisuchus
                  |        |     |-- †Candidodon
                  |        |     `--+-- †Mariliasuchus
                  |        |        `--+-- †Pakasuchus
                  |        |           `-- †Adamantinasuchus
                  |        `--+-- †Simosuchus
                  |            `--+-- †Notosuchus
                  |               `--+--+-- †Chimaerasuchus
                  |                  |  `-- †Sphagesaurus
                  |                  `--+-- †Comahuesuchus
                  |                     `--+-- †Baurusuchus
                  |                        `--+-- †Bretesuchus
                  |                           `-- †Iberosuchus
                  `--+--+-- †Lomasuchus
                     |  `-- †Peirosaurus
                     `--+-- †Mahajangasuchus
                        `--+-- †Anatosuchus
                           `-- Neosuchia

Reference:

  • O'Connor, P. M., Sertich, J. J. W., Stevens, N. J., Roberts, E. M., Gottfried, M. D., Hieronymus, T. L., Jinnah, Z. A., Ridgely, R., Ngasala, S. E. & Temba, J., 2010: The evolution of mammal-like crocodyliforms in the Cretaceous Period of Gondwana.
    –Nature: Vol. 466, #7307, pp. 748-751 [doi: 10.1038/nature09061]

After Carvalho, Ribeiro & Avilla, 2004

<==o Gondwanasuchia
   |--o †Chimaerasuchidae
   |  |-- †Chimaerasuchus
   |  `-- †Simosuchus
   `--o †Notosuchimorpha
      |-- †Stolokrosuchus
      `--o †Notosuchiformes
         |-- †Uruguaysuchus
         `--o †Terriasuchia
            |-- †Comahuesuchus
            `--o †Notosuchia
               |--o †Ziphosuchia
               |  |--o †Notosuchidae
               |  |  `-- †Notosuchus
               |  `--o †Baurusuchoidea
               |     |-- †Baurusuchidae Price, 1945
               |     `-- †Sebecidae
               `--o †Peirosaurimorpha
                  |--o †Candidodontidae
                  |  |-- †Mariliasuchus
                  |  `-- †Candidodon
                  `--o †Peirosauriformes
                     |-- †Araripesuchus
                     `--o †Peirosauroidea
                        |--o †Itasuchidae
                        |  |-- †Itasuchus
                        |  `-- †Malawisuchus
                        `--o †Peirosauridae
                           |-- †Peirosaurus Price, 1955
                           `--o †Lomasuchinae
                              |-- †Lomasuchus
                              `--o †Mahajangasuchini
                                 |-- †Mahajangasuchus
                                 `-- †Uberabasuchus

Reference(s):

  • Carvalho, I. d. S., Ribeiro, L. C. B. & Avilla, L. d. S., 2004: Uberabasuchus terrificus sp. nov., a new Crocodylomorpha from the Bauru Basin (Upper Cretaceous), Brazil.
    –Gondwana Research: Vol. 7, #4, pp. 975-1002

After Tykoski, Rowe, Ketcham & M. W. Colbert, 2002

<==o Crocodyliformes
   |-- †Eopneumatosuchus
   `--+?-o †Artzosuchidae
      |  `-- †Artzosuchus
      |-- †Protosuchia
      `--+?-o †Shartegosuchidae
         |  |?- †Kyasuchus
         |  |-- †Nominosuchus
         |  `-- †Shartegosuchus
         `--o Mesoeucrocodylia
            |?- †Fruitachampsa
            |--o †Thalattosuchia
            |  |-- †Pelagosuchus
            |  `--+-- †Teleosauridae
            |     `-- †Metriorhynchidae
            `--+?-o †Gobiosuchidae
               |  |-- †“Las Hoyas crocodyliform”
               |  |-- †Zaraasuchus
               |  `-- †Gobiosuchus
               |?- †Simosuchus
               |?- †Lisboasaurus
               |--o †Hsisosuchidae
               |  |?- †Doratodon
               |  `-- †Hsisosuchus
               `--+-- †Ziphosuchia
                  `--+-- Neosuchia
                     `--+-- †Araripesuchus
                        `--+--o †Trematochampsidae
                           |  |?- †Baharijodon
                           |  |?- †Amargasuchus
                           |  |?- †Caririsuchus
                           |  `-- †Trematochampsa
                           `--+-- †Mahajangasuchus
                              `--o †Peirosauridae
                                 |-- †Stolokrosuchus
                                 `--o †Peirosauridae
                                    |-- †Peirosaurus
                                    `--o †Lomasuchinae
                                       |-- †Lomasuchus
                                       `--o †Mahajangasuchini
                                          |-- †Mahajangasuchus
                                          `-- †Uberabasuchus

Reference(s):

  • Tykoski, R. S., Rowe, T. B., Ketcham, R. A. & Colbert, M. W., 2002: Calsoyasuchus valliceps, a new crocodyliform from the Early Jurassic Kayenta Formation of Arizona.
    –Journal of Vertebrate Paleontology: Vol. 22, #3, pp. 593-611

After Sereno & Larsson, 2009

<==o Crocodyliformes
   |-- †Zosuchus
   `--+-- †Hsisosuchus
      `--o Mesoeucrocodylia
         |--o Thalattosuchia
         |  `--+-- †Pelagosuchus
         |     `--+-- †Teleosauridae
         |        `-- †Metriorhynchidae
         `--o Metasuchia
            |--o †Notosuchia
            |  |== †“Araripesuchus”
            |  |--+-- †Uruguaysuchus
            |  |  `--+-- †Simosuchus
            |  |     `-- †Anatosuchus
            |  `--+-- †Baurusuchus
            |     `--+--+-- †Mariliasuchus
            |        |  `-- †Comahuesuchus
            |        `--+-- †Malawisuchus
            |           |-- †Notosuchus
            |           `-- †Sphagesaurus
            `--o Neosuchia
               |--o †Sebecia
               |  |-- †Uberabasuchus
               |  `--+-- †Hamadasuchus
               |     `--+-- †Sebecus
               |        `--+-- †Stolokrosuchus
               |           `--+-- †Peirosaurus
               |              `-- †Lomasuchus
               `--+--o †Mahajangasuchidae
                  |  |-- †Mahajangasuchus
                  |  `-- †Kaprosuchus
                  `--+--o †Pholidosauridae
                     |  |-- †Goniopholis
                     |  `--+-- †Dyrosaurus
                     |     `--+-- †Terminonaris
                     |        `--+-- †Sarcosuchus
                     |           `-- †Pholidosaurus
                     `--+-- †Laganosuchus
                        `--+-- †Bernissartia
                           `-- Eusuchia

Reference(s):

  • Sereno, P. C. & Larsson, H. C. E., 2009: Cretaceous crocodyliforms from the Sahara.
    –Zookeys: Vol. 28, pp. 1-143 [doi: 10.3897/zookeys.28.325]

After Clark, 1994 (fig. 5.1 & tablet 5.1c):

<==o Crocodyliformes
   |-- †Eopneumatosuchus
   |-- †Gobisuchus
   |-- †Orthosuchus
   |--o †Protosuchidae
   |  |-- “Kayenta Edentosuchus-like form”
   |  `--o †Protosuchinae
   |     |-- †Hemiprotosuchus
   |     `-- †Protosuchus
   `--o Mesoeucrocodylia
      |-- †Fruitachampsa
      `--+-- †Baurusuchus
         |-- †Notosuchus
         `--+-- †Libycosuchus
            `--+-- †Sebecus
               `--+-- †Araripesuchus
                  `--o Neosuchia
                     |--o †Atoposauridae
                     |  +-- †Alligatorium
                     |  `-- †Theriosuchus
                     `--+-- †Goniopholis
                        |-- †Bernissartia
                        |-- Eusuchia
                        `--+-- †Eutretauranosuchus
                           `--+--o †Dyrosauridae
                              |  |-- †Sokotosuchus
                              |  `-- †Dyrosuchus
                              `--+-- †Pholidosaurus
                                 `--o †Thalattosuchia
                                    |-- †Pelagosaurus
                                    `--+-- †Metriorhynchidae
                                       `-- †Teleosauridae

Reference(s):

  • Clark, J. M. 1994: Patterns of evolution in Mesozoic Crocodyliformes. 84-97.
    in Fraser, N. C. & Sues, H.-D. (eds.), 1994: In the Shadow of the Dinosaurs – Early Mesozoic Tetrapods.
    –Cambridge University Press, Cambridge, New York, x-435

Reference(s):