After Clark, 1994, Ortega, Gasparini, Buscalioni & Calvo, 2000, and Turner & Calvo, 2005
Family -level taxonomy of all crocodylomorphs [Crocodylia sensu lato]
Alternative phylogeny of Crocodyliformes after Sereno, Larsson, Sidor & Gado, 2001
Alternative phylogeny of Crocodyliformes after Tykoski, Rowe, Ketcham & M. W. Colbert, 2002
Alternative phylogeny of Crocodyliformes after Clark, 1994 and Pol & Norell, 2004
Alternative phylogeny of Crocodyliformes after Clark, 1994, and Carvalho, Ribeiro & Avilla, 2004
<==o Crocodyliformes Hay, 1930 `==o †Protosuchia Mook, 1934 [paraphyletic] |-- †Eopneumatosuchus colberti Crompton & Smith, 1980 |?- †Microchampsa scutata Young, 1951 [nomen dubium] `--+?- †Artzosuchus brachicephalus Efimov, 1983 [Artzosuchidae Efimov, 1983] |?- Protosuchia genera et species indet.1 [North America, Kayenta Formation] |?- †Hadongsuchus acerdentis |-- †Orthosuchus strombergi Nash, 1975 [?Orthosuchidae] `--+?- †Platyognathus hsui Young, 1944 [Platyognathidae] |?-o Edentosuchidae | |?- Protosuchia genera et species indet.2 [North America, Kayenta Formation] | |?- Protosuchia genera et species indet.3 [North America, Kayenta Formation, UCMP unnumbered] | `-- †Edentosuchus tienshanensis Young, 1973 `==o †Protosuchidae sensu lato |-- †Dianosuchus changchiawaensis Young, 1982 |-- †Erythrochampsa longipes (Broom, 1904) Haughton, 1924 [Notochampsa longipes Broom, 1904] |-- †Notochampsa istedana Broom 1904 [Nothochampsa (lapsus calami)] [Notochampsidae Haughton, 1924] |-- †Stegomosuchus longipes Emerson & Loomis, 1904 |?- †Tagarosuchus kulemzini Efimov, 1999 `--+--o †Protosuchidae Brown, 1934 sensu Wu, Brinkman & Lu, 1994 | |-- †Hemiprotosuchus leali Bonaparte, 1972 | `--o †Protosuchus Brown, 1934 [Lesothosuchus Whetstone & Whybrow, 1983; Baroqueosuchus Busbey & Gow, 1984] | |-- †P. sp.1 | |-- †P. sp.2 | |-- †P. micmac Sues, Shubin, Olsen & Amaral, 1996 | |-- †P. richardsoni (Brown, 1933) | `-- †P. haughtoni (Whetstone & Whybrow, 1983) `--+?- †“Las Hoyas crocodyliform” `--+--o †Gobiosuchidae Osmólska, 1972 [moved to Protosuchia by Wu, Sues & Dong, 1997] | |-- †Zaraasuchus shepardi Pol & Norell, 2004b | `--o †Gobiosuchus Osmólska, 1972 [Gobiosuchidae] Osmólska, 1972 | |-- †G. kielanae Osmólska, 1972 | `-- †G. parvus Efimov, 1983 `--+--+-- †Zosuchus davidsoni Pol & Norell, 2004 | `--+--o †Sichuanosuchus Peng, 1995; U. Jur. ?L. Cret., EAs. | | |-- †S. huidongensis Peng, 1995 | | `-- †S. shuhanensis Wu, Sues & Dong, 1997 | `--o †Shantungosuchus Young, 1961 | |-- †S. chuhsienensis Young, 1961 | |-- †S. brachycephalus Young, 1982 [posthumous] | `-- †S. hangjinensis Wu, Brinkman & Lu, 1994 |--o †Shartegosuchidae Efimov, 1988 | |?- †Kyasuchus saevi Efimov & Leshchinskiy, 2000 | |--o †Nominosuchus Efimov, 1996 | | |-- †N. matutinus Efimov, 1996 | | `-- †N. arcanus Kurzanov, Efimov & Gubin, 2003 | `--+-- †Fruitachampsa callisoni Clark, 2012; L.Jur. WNAm.(“Fruita crocodyliform”) | `--+-- †Shartegosuchus asperopalatum Efimov, 1988 | `-- †Adzhosuchus fuscus Efimov, Gubin & Kurzanov, 2000 `--+-- †Hsisosuchus chungkingensis (Young & Chow, 1953) [Hsisosuchidae] `--o Mesoeucrocodylia Whetstone & Whybrow, 1983 |-- †Notosuchia Gasparini, 1971 sensu Turner & Sertich, 2010 [incl. Ziphosuchia Ortega, Gasparini, Buscalioni & Calvo, 2000] `--+--o †Trematochampsidae | |-- †Amargasuchus minor Chiappe, 1988 | |?- ?†Baharijodon carnosauroides Kuhn, 1936 | |-- †Caririsuchus camposi Kellner, 1987 | `--o †Trematochampsa Buffetaut, 1974 | |-- †T. taqueti Buffetaut, 1974 | `-- †T. oblita Buffetaut & Taquet, 1979 `-- Neosuchia sensu Clark, 1994 (modern crocodiles; modernit krokotiilit)
Mesoeucrocodylia incertae sedis: [= Mesocrocodiles not yet reviewed]
†Bottosaurus Agassiz, 1849
†B.harlani
†B. perrugosus
†B. tuberculatus
†B. perrugosus
†Dianchungosaurus lufengensis Yang, 1982a
†Entradasuchus spinosus Hunt & Lockley, 1995
†Lisboasaurus estesi Seiffert, 1970 sensu Buscalioni, Pérez-Moreno & Evans, 1996 [non-archosaurian archosauromorphan? (a protorosaurian?)]
†Lusitanisuchus mitracostatus (Seiffert, 1970) Schwarz & Fechner, 2004 [Lisboasaurus mitracostatus Seiffert, 1970]
†Khoratosuchus jintasakuli Lauprasert et al., 2009
†Cunampaiidae Cracraft, 1968
†Cunampaia simplex Rusconi, 1946 [originally descriped as a prhorusrhacoid bird!!]
Alternative phylogenies of Crocodyliformes:
After Holliday & Gardner, 2012Holliday & Gardner (2012) used Sereno & Larsson's datamatrix, with including new coding for Aegisuchus. <==o Crocodyliformes |-- †Zosuchus `--+-- †Hsisosuchus `--o Mesoeucrocodylia |--o Thalattosuchia | `--+-- †Pelagosuchus | `--+-- †Teleosauridae | `-- †Metriorhynchidae `--o Metasuchia |--o †Notosuchia | |--+== †Araripesuchus | | `--+-- †Uruguaysuchus | | `--+-- †Simosuchus | | `-- †Anatosuchus | `--+-- †Baurusuchus | `--+--+-- †Mariliasuchus | | `-- †Comahuesuchus | `--+-- †Malawisuchus | |-- †Notosuchus | `-- †Sphagesaurus `--o Neosuchia |--o †Sebecia | |-- †Uberabasuchus | `--+-- †Hamadasuchus | `--+-- †Sebecus | `--+-- †Stolokrosuchus | `--+-- †Peirosaurus | `-- †Lomasuchus `--+--o †Mahajangasuchidae | |-- †Mahajangasuchus | `-- †Kaprosuchus `--+--o †Pholidosauridae | |-- †Goniopholis | `--+--+-- †Dyrosaurus | | `-- †Terminonaris | `--+-- †Sarcosuchus | `-- †Pholidosaurus `--+-- †Laganosuchus `--+-- †Bernissartia `-- Eusuchia Reference(s):
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After O'Connor et al., 2010<==o Crocodyliformes |--+-- †Orthosuchus | `--+--+-- †Hemiprotosuchus | | `-- †Protosuchus | `--+-- “Kayenta Edentosuchus-like form” | `-- †Edentosuchus `--+--+-- †Zaraasuchus | `-- †Gobisuchus `--+--+-- †Zosuchus | `--+-- †Sichuanosuchus | `-- †Shantungosuchus `--+-- †Fruitachampsa (“Fruita form”) `--+-- †Hsisosuchus `--o Mesoeucrocodylia |--o †Notosuchia | |--+-- †Araripesuchus wegeneri | | `--+-- †Araripesuchus buitreraensis | | `--+-- †Araripesuchus gomesii | | `-- †Araripesuchus patagonicus | `--+-- †Uruguaysuchus | `--+--+-- †Libycosuchus | | `--+-- †Malawisuchus | | |-- †Candidodon | | `--+-- †Mariliasuchus | | `--+-- †Pakasuchus | | `-- †Adamantinasuchus | `--+-- †Simosuchus | `--+-- †Notosuchus | `--+--+-- †Chimaerasuchus | | `-- †Sphagesaurus | `--+-- †Comahuesuchus | `--+-- †Baurusuchus | `--+-- †Bretesuchus | `-- †Iberosuchus `--+--+-- †Lomasuchus | `-- †Peirosaurus `--+-- †Mahajangasuchus `--+-- †Anatosuchus `-- Neosuchia Reference:
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After Carvalho, Ribeiro & Avilla, 2004<==o Gondwanasuchia |--o †Chimaerasuchidae | |-- †Chimaerasuchus | `-- †Simosuchus `--o †Notosuchimorpha |-- †Stolokrosuchus `--o †Notosuchiformes |-- †Uruguaysuchus `--o †Terriasuchia |-- †Comahuesuchus `--o †Notosuchia |--o †Ziphosuchia | |--o †Notosuchidae | | `-- †Notosuchus | `--o †Baurusuchoidea | |-- †Baurusuchidae Price, 1945 | `-- †Sebecidae `--o †Peirosaurimorpha |--o †Candidodontidae | |-- †Mariliasuchus | `-- †Candidodon `--o †Peirosauriformes |-- †Araripesuchus `--o †Peirosauroidea |--o †Itasuchidae | |-- †Itasuchus | `-- †Malawisuchus `--o †Peirosauridae |-- †Peirosaurus Price, 1955 `--o †Lomasuchinae |-- †Lomasuchus `--o †Mahajangasuchini |-- †Mahajangasuchus `-- †Uberabasuchus Reference(s):
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After Tykoski, Rowe, Ketcham & M. W. Colbert, 2002<==o Crocodyliformes |-- †Eopneumatosuchus `--+?-o †Artzosuchidae | `-- †Artzosuchus |-- †Protosuchia `--+?-o †Shartegosuchidae | |?- †Kyasuchus | |-- †Nominosuchus | `-- †Shartegosuchus `--o Mesoeucrocodylia |?- †Fruitachampsa |--o †Thalattosuchia | |-- †Pelagosuchus | `--+-- †Teleosauridae | `-- †Metriorhynchidae `--+?-o †Gobiosuchidae | |-- †“Las Hoyas crocodyliform” | |-- †Zaraasuchus | `-- †Gobiosuchus |?- †Simosuchus |?- †Lisboasaurus |--o †Hsisosuchidae | |?- †Doratodon | `-- †Hsisosuchus `--+-- †Ziphosuchia `--+-- Neosuchia `--+-- †Araripesuchus `--+--o †Trematochampsidae | |?- †Baharijodon | |?- †Amargasuchus | |?- †Caririsuchus | `-- †Trematochampsa `--+-- †Mahajangasuchus `--o †Peirosauridae |-- †Stolokrosuchus `--o †Peirosauridae |-- †Peirosaurus `--o †Lomasuchinae |-- †Lomasuchus `--o †Mahajangasuchini |-- †Mahajangasuchus `-- †Uberabasuchus Reference(s):
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After Sereno & Larsson, 2009<==o Crocodyliformes |-- †Zosuchus `--+-- †Hsisosuchus `--o Mesoeucrocodylia |--o Thalattosuchia | `--+-- †Pelagosuchus | `--+-- †Teleosauridae | `-- †Metriorhynchidae `--o Metasuchia |--o †Notosuchia | |== †“Araripesuchus” | |--+-- †Uruguaysuchus | | `--+-- †Simosuchus | | `-- †Anatosuchus | `--+-- †Baurusuchus | `--+--+-- †Mariliasuchus | | `-- †Comahuesuchus | `--+-- †Malawisuchus | |-- †Notosuchus | `-- †Sphagesaurus `--o Neosuchia |--o †Sebecia | |-- †Uberabasuchus | `--+-- †Hamadasuchus | `--+-- †Sebecus | `--+-- †Stolokrosuchus | `--+-- †Peirosaurus | `-- †Lomasuchus `--+--o †Mahajangasuchidae | |-- †Mahajangasuchus | `-- †Kaprosuchus `--+--o †Pholidosauridae | |-- †Goniopholis | `--+-- †Dyrosaurus | `--+-- †Terminonaris | `--+-- †Sarcosuchus | `-- †Pholidosaurus `--+-- †Laganosuchus `--+-- †Bernissartia `-- Eusuchia Reference(s):
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After Clark, 1994 (fig. 5.1 & tablet 5.1c):<==o Crocodyliformes |-- †Eopneumatosuchus |-- †Gobisuchus |-- †Orthosuchus |--o †Protosuchidae | |-- “Kayenta Edentosuchus-like form” | `--o †Protosuchinae | |-- †Hemiprotosuchus | `-- †Protosuchus `--o Mesoeucrocodylia |-- †Fruitachampsa `--+-- †Baurusuchus |-- †Notosuchus `--+-- †Libycosuchus `--+-- †Sebecus `--+-- †Araripesuchus `--o Neosuchia |--o †Atoposauridae | +-- †Alligatorium | `-- †Theriosuchus `--+-- †Goniopholis |-- †Bernissartia |-- Eusuchia `--+-- †Eutretauranosuchus `--+--o †Dyrosauridae | |-- †Sokotosuchus | `-- †Dyrosuchus `--+-- †Pholidosaurus `--o †Thalattosuchia |-- †Pelagosaurus `--+-- †Metriorhynchidae `-- †Teleosauridae Reference(s):
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Reference(s):
- Agnolin, F. L. & Pais, D. F., 2006: Revisión de Cunampaia simplex Rusconi, 1956 (Crocodylomorpha, Mesoeucrocodylia: non Aves) del Terciario Inferior de mendoza, Argentina.
–Revista del Museo Argentino de Ciencias Naturales: Vol. 8, #1, pp. 35-40 - Barrett, P. M. & Xing, X. U., 2005: A reassessment of Dianchungosaurus lufengensis Yang, 1982a, an enigmatic reptile from the lower Lufeng Formation (lower Jurassic) of Yunnan Province, People's Republic of China.
–Journal of Paleontology: Vol. 79, #5, pp. 981-986 [doi: 10.1666/0022-3360(2005)079[0981:ARODLY]2.0.CO;2] - Buckley, G. A., 2001: A skull of Mahajangasuchus insignis (Crocodyliformes) from the Upper Cretaceous of Madagascar.
–Journal of Vertebrate Paleontology: Vol. 21, #3, Supplement to #3, pp. A36 - Buscalioni, A. D., Pérez-Moreno, B. P. & Evans, S. E., 1996: The Upper Jurassic maniraptoran theropod Lisboasaurus estesi (Guimarota, Portugal), reinterpreted as a crocodylomorph.
–Journal of Vertebrate Paleontology: Vol. 16, #2, pp. 358-362 - Clark, J. M. 1994: Patterns of evolution in Mesozoic Crocodyliformes. 84-97.
in Fraser, N. C. & Sues, H.-D. (eds.), 1994: In the Shadow of the Dinosaurs - Early Mesozoic Tetrapods.
–Cambridge University Press, Cambridge, New York, x-435 - Hunt, A. P. & Lockley, M. G., 1995: A nonmarine tetrapod from the Middle Jurassic of the United States: A primitive crocodyliform from the Entrada Sandstone of Eastern Utah.
–Journal of Vertebrate Paleontology: Vol. 15, #3, pp. 554-560 - Leardi, J. M. & Pol, D., 2009: The first crocodyliform from the Chubut Group (Chubut Province, Argentina) and its phylogenetic osition within basal Mesoeucrocodylia.
–Cretaceous Research: Vol. 30, pp. 1376-1386 [doi: 10.1016/j.cretres.2009.08.002] - Luo, Z, & Wu, X-C., 1994: The small tetrapods of the Lower Lufeng Formation, Yunnan, China. 251-270
in Fraser, N. C. & Sues, H.-D. (eds.), 1994: In the Shadow of the Dinosaurs - Early Mesozoic Tetrapods.
–Cambridge University Press, Cambridge, New York, x-435 - Ortega, F., Gasparini, Z., Buscalioni, A. D. & Calvo, J. O., 2000: A new species of Araripesuchus (Crocodylomorpha: Mesoeucrocodylia) from the Lower Cretaceous of Patagonia (Argentina).
–Journal of Vertebrate Paleontology: Vol. 20, #1, pp. 57-76 - Pol, D. & Norell, M. A., 2004a: A new crocodyliform from Zos Canyon, Mongolia.
–American Museum Novitates: #3445, pp. 1-36 - Pol, D. & Norell, M. A., 2004b: A new gobiosuchid crocodyliform taxon from the Cretaceous of Mongolia.
–American Museum Novitates: #3458, pp. 1-31 - Schwarz, D. & Fechner, R., 2004: Lusitanisuchus, a new generic name for Lisboasaurus mitracostatus (Crocodylomorpha: Mesoeucrocodylia), with a description of new ramains from the Upper Jurassic (Kimmeridgian) and Lower Cretaceous (Berriasian) of Portugal.
–Canadian Journal of Earth Sciencies: Vol. 41, pp. 1259-1271 - Sereno, P. C. & Larsson, H. C. E., 2009: Cretaceous crocodyliforms from the Sahara.
–Zookeys: Vol. 28, pp. 1-143 [doi: 10.3897/zookeys.28.325] - Storrs, G. W. & Efimov, M. B., 2000: Mesozoic crocodyliforms of north-central Eurasia. 402-419
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000 xxxix-696 - Turner, A. H. & Sertich, J. J. W., 2010: Phylogenetic history of Simosuchus clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar.
–Society of Vertebrate Paleontology Memoir 10: Journal of Vertebrate Paleontology: Vol. 30, Supplement to #6, pp. 177-236 [doi: 10.1080/02724634.2010.532348] - Wu, X.-C., Sues, H.-D. & Dong, Z.-M., 1997: Sichuanosuchus shuhanensis, a new ?Early Cretaceous Protosuchian (Archosauria: Crocodyliformes) from Sichuan (China), and the monophyly of Protosuchia.
–Journal of Vertebrate Paleontology: Vol. 17, #1, pp. 89-103