After “The Tree of Life” (1994-2000), Beutel & Haas, 2000, Grimaldi & Engel, 2005, and Beutel, Balke & Steiner Jr., 2006
Note:
Only those species and genera dealed by Beutel et al., 2006 are presented. I will try to update more compleat species list in the future. If you find this lack of information annoying, you're welcome to contribute.
<==o Adephaga (petokuoriaiset; ground beetles, tiger beetles, whirligigs, etc.) |?-o †Coptoclavidae [polyphyletic] | |--o †Charonoscaphinae | | |-- †Charonoscapha | | `-- †Charonoscaphidia | |--o †Coptoclavinae | | `--o †Coptoclava | | `-- †C. longipoda | `--o †Necronectinae [polyphyletic] | |-- †Exedia | `-- †Stygeonectes |?- †Liadytidae |?- †Colymbothetidae |?- †Jurodidae |?- †Eodromeinae `--+--o Gyrinidae (hopeasepät, pintapyöriöt) | |?- †Permosialis | |?- †Triadogyrus | |?- †Anagyrinus | |?- †Angarogyrus | |--o Spanglerogyrinae | | `-- Spanglerogyrus albiventris Folkerts | `--o Gyrininae | |--o Gyrinini | | |-- Aulonocyrus concinnus Klug | | `-- Gyrinus argentinus Mouchamps | `--+--o Enhydrini [paraphyletic] [non Enhydrini, the sea otters] | | `-- Enhydrus | `--+-- Dineutus assimilis Kirby | `--+-- Andogyrus colombicus Régimart | |-- Macrogyrus | `--+-- Gyretes | `--o Orectochilini | |-- Orectochilus villosus Müller | `-- Orectogyrus `--+--+?- †Triaplidae | `--o Haliplidae (vesijuoksijat) | |-- Peltodytes caesus (Duftschmidt) | `--+-- Brychius elevatus (Panzer) | `--+-- Haliplus lineatocollis (Marsham) | |-- Haliplus mucronatus Stephens | |-- (Liaphlus) | |-- (Paraliaphlus) | |-- (Neoaphlus) | `--+-- Apteraliplus parvulus Roberts | `-- Algophilus |--+--o Trachypachidae | | |-- †Psacodromeus guttata U.Jur. CAs. | | |== six other extinct genera | | |--o Trachypachus | | | `-- T. holmbergi Mannerheim | | `--o Systolosoma | | `-- S. breve Solier | `--+--o Rhysodidae | | |-- Omoglymmius hamatus LeConte | | `-- Clinidium americanum | `--o Carabidae [incl. Brachinidae; Cicindelidae; Omophronidae; Paussidae; Pseudomorphidae; and ?Trachypachidae] (maakiitäjäiset) | |?- Gehringia olympiaca Darlington | |?- Nebria brevicollis | |?- Notiophilus biguttatus | |--+-- Bembidion | | `-- Pterostichus | |--+-- Carabus | | |-- Omophron limbatum Olivier | | `-- Metrius contractus Eschscholtz | |--o Paussinae | | |-- Paussini | | |-- Metrini | | `--o Brachinini | | `-- Brachinus (bombardier beetle) | `--+-- Harpalinae | |-- Cicindelinae (tiger beetles) | |-- Pseudomorphinae | |-- Carabinae | `-- Omophroninae `--o Dytiscoidea |--+-- Meru phyllisae Spangler & Steiner [Meruidae] | `--o Noteridae [incl. Phreatodytidae] | |--o Phreatodytinae [Phreatodytidae] | | `-- Phreatodytes | `--o Noterinae | |-- Notomicrus gracilipes Sharp | `--+-- Hydrocoptus subvittatus Motschulsky | `--+-- Pronoterus obscuripennis Fleutiaux & Salle | `--+-- Synchortus sparsus Sharp | |-- Mesonoterus addendus Blatchley | |--o Noterus | | |-- N. clavicornis DeGeer | | `-- N. crassicornis Müller | `--+-- Renotus deyrollei (Sharp) | |-- Siolius bicolor J. Balfour-Browne | |-- Suphis inflatus (LeConte) | `--+-- Hydrocanthus | |-- Canthydrus luctuosus (Aubé) | `-- Suphisellus bicolor (Say) `--+--o Aspidytidae | `--o Aspidytes | |-- A. niobe Ribera et al. | `-- A. wrasei Balke, Ribera & Beutel `--+--o Amphizoidae | `--o Amphizoa | |-- A. lecontei Matthews | `-- A. insolens LeConte `--+--o Dytiscidae (sukeltajakuoriaiset, sukeltajat) | |-- †Angaragabus | |== numerous Recent genera | |-- Liopterus haemorrhoidalis | `--+-- Dytiscus marginalis | `--o Agabus | |-- A. bipustulatus | `-- A. guttatus `--+--o †Parahygrobiidae | `-- †Parahygrobia natans U.Jur. NAs. `--o Hygrobiidae [Pelobiidae] `--o Hygrobia |-- H. hermanni `-- H. australasiae
Reference(s):
- Beutel, R. G., Balke, M. & Steiner Jr., W. E., 2006: The systematic position of Meruidae (Coleoptera, Adephaga) and the phylogeny of the smaller aquatic adephagan beetle families.
–Cladistics: Vol. 22, #1, pp. 102-131 - Beutel, R. G. & Haas, F., 2000: Phylogenetic relationships of the suborders of Coleoptera (Insecta).
–Cladistics: Vol. 16, #1, pp. 103-141 - Grimaldi, D. A. & Engel, M. S., 2005: Evolution of the Insects.
–Cambridge University Press, New York, 2005, xv-755 - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968 - Maddison, D. R., 1994: Coleoptera – beetles. (Updated: 2000-09-11)
–iNet: The Tree of Life Project: http://tolweb.org/tree?group=Coleoptera&contgroup=Endopterygota; [visited: 2005-06-23] - Parker, S. P. (ed.), 1982: Synopsis and classification of living organisms. Vols. 1 & 2
–McGrew-Hill Book Company