Coleoptera

After “The Tree of Life” (1994-2000), Beutel & Haas, 2000, Grimaldi & Engel, 2005, and Béthoux, 2009

<==o Coleoptera Linnaeus, 1758 sensu de Geer, 1774 (beetles; kovakuoriaiset)
   |?- †Sylvacoelus sharovi Perm. EEu.
   |--o †Protocoleoptera
   |  `--o †Tschekardocoleidae Rohdendorf, 1944
   |     `-- †Moravocoleus
   `--o Coleoptera sensu Grimaldi & Engel, 2005
      `==o “Archecoleoptera” [paraphyletic stem-beetles]
         |?- †Notocupoides triassicus Trias. EEu.
         |?- †Schizocoleidae
         |?-+-- †Taldycupedidae
         |  `-- †Tricoleidae
         |?- †Asiocoleidae
         |--o †Permocupedidae
         |  `-- †Permocupoides sojanensis Perm. EEu.
         `--+-- †Rhombocoleidae
            `--+-- †Triabocupedidae
               `--+-- Archostemata (alkukuoriaiset)
                  `--o “Neocoleoptera”
                     |-- Adephaga (petokuoriaiset; ground beetles, tiger beetles, whirligigs, etc.)
                     `--+-- Myxophaga (rihmansyöjät) [maybe derivied basal polyphagans (Grimaldi & Engel, 2005)]
                        `--+?-o †Ademosytidae
                           |  `-- †Ademosyne
                           `-- Polyphaga (erilaisruokaiset; scarabs, fireflies, ladybirds, weevils, etc.)

Reference(s):

Béthoux, O., 2009: The earliest beetle identified.
–Journal of Paleontology: Vol. 83, #6, pp. 931-937
Beutel, R. G. & Haas, F., 2000: Phylogenetic relationships of the suborders of Coleoptera (Insecta).
–Cladistics: Vol. 16, #1, pp. 103-141
Grimaldi, D. A. & Engel, M. S., 2005: Evolution of the Insects.
–Cambridge University Press, New York, 2005, xv-755
Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968
Maddison, D. R., 1994: Coleoptera – beetles. (Updated: 2000-09-11)
–iNet: The Tree of Life Project: http://tolweb.org/tree?group=Coleoptera&contgroup=Endopterygota; [visited: 2005-06-23]
Maddison, D. R., 1996: Ommatidae. (Updated: ????)
–iNet: The Tree of Life Project: http://tolweb.org/tree?group=Ommatidae&contgroup=Archostemata; [visited: 2005-06-23]
Maddison, D. R., 1996: Cupedidae. (Updated: ????)
–iNet: The Tree of Life Project: http://tolweb.org/tree?group=Ommatidae&contgroup=Archostemata; [visited: 2005-06-23]
Parker, S. P. (ed.), 1982: Synopsis and classification of living organisms. Vols. 1 & 2
–McGrew-Hill Book Company

Mikko's Phylogeny Archive

Coleoptera – beetles

After “The Tree of Life” (1994-2000), Beutel & Haas, 2000, Grimaldi & Engel, 2005, and Béthoux, 2009

Reference(s):