After McKenna & Bell, 1997

<==o †Pantolesta McKenna, 1975 [taxonomic history]
   |--o †Simidectes Stock, 1933 [taxonomic history] M.Eoc. NA.
   |  |-- †S. magnus (Peterson, 1919) [taxonomic history]
   |  |-- †S. medius (Peterson, 1919) [taxonomic history]
   |  `-- †S. merriami (Stock, 1933b) [taxonomic history]
   |-- †Pantolestidae Cope, 1884 [taxonomic history]
   `--+--o ?†Paroxyclaenidae [formerly root-ungulates]
      |  |?- †Dulcidon gandaensis [a dichobunid??]
      |  |--o †Paroxyclaeninae Weitzel, 1933
      |  |  |?- †Kiinkerishella zaysanica Gabunia & Birjukov, 1978; Eoc. As. [Paroxyclaenidae incertae sedis (McKenna & Bell, 1997)]
      |  |  |--o †Paroxyclaenus Teilhard de Chardin, 1922; ?L.-U.Eoc. Eu.
      |  |  |  `-- †P. lemuroides
      |  |  |-- †Spaniella Crusafont Pairó & D.E. Russell, 1967; L.Eoc. Eu.
      |  |  |--o †Vulpavoides Matthes, 1952 [incl.? Pugiodens??; †Russellites Van Valen, 1965] M.Eoc. Eu.
      |  |  |  |-- †E. simplicidens (van Valen, 1965)
      |  |  |  `-- †E. vanvaleni
      |  |  |-- †Pugiodens Matthes, 1952 [=? Vulpavoides] M.Eoc. Eu.
      |  |  `--o †Kopidodon Weitzel, 1933; M.Eoc. Eu.
      |  |     `-- †K. macrognathus
      |  `--o †Merialinae D.E. Russell & Godinot, 1988
      |     |?- †Oboia argillaceus Gabunia, 1989; L.Olig. As. [Pantolestidae: Pantolestinae sensu McKenna & Bell, 1997]
      |     |-- †Merialus martinae D.E. Russell & Godinot, 1988; L.Eoc. Eu.
      |     `-- †Euhookeria hopwoodi (Cray, 1973) D.E. Russell & Godinot, 1988; U.Eoc. Eu.
      `?-o †Ptolemaiidae Osborn, 1908; U. Eoc. NEAf. [recently treated as basal Afrotherians]
         |-- †Qarunavus meyeri Simons & Gingerich, 1974; U.Eoc. NEAf.
         |--o †Cleopatrodon Bown & Simons, 1987; U. Eoc. NEAf.
         |  |-- †C. ayeshae Bown & Simons, 1987
         |  `-- †C. major Bown & Simons, 1987
         `--o †Ptolemaia Osborn, 1908 [non †Ptolemaia McCutcheon & Wilson, 1961, a fossil rugose coral (Cnidaria: Anthozoa:Rugosa)]; U. Eoc. NEAf.
            |-- †P. lyonsi Osborn, 1908
            `-- †P. grangeri Bown & Simons, 1987

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