After Meylan & Ganko, 1988 [from Meylan & Ganko, 1997, Meylan, 2001], Parham & Hutchison, 2003, and Parham, 2005
<==o Cryptodira Dumeril & Bibron, 1835 [non Cope, 1871] [Pan-Cryptodira Joyce, Parham & Gauthier, 2004; Cryptodiramorpha Lee, 1995 sensu Parham & Hutchison, 2003] (pystytaivuttajat) |-- †Kayentachelys aprix Gaffney, Hutchison, Jenkins & Meeker, 1987; E.Jur, SWNAm. [Kayentachelyidae Gaffney, Hutchison, Jenkins & Meeker, 1987] |-- †Indochelys spatulata Datta, Manna, Ghosh & Das, 2000 [Indochelyidae Datta, Manna, Ghosh & Das, 2000] |?- †Camerochelys vilanovai Pérez-García & Murelaga, 2013 `--o Selmacryptodira Gaffney, Hutchinson, Jenkins & Meeker, 1987 |-- †Paracryptodira [=? Pleurosternoidea Hay, 1930] `--o Eucryptodira Gaffney, 1975a sensu Gaffney, 1984 [Daiocryptodira Milner, 2004] |--+-- †Plesiochelyidae Baur, 1888 [non Rütimeyer, 1873?] [incl.? Thalassemydidae Zittel, 1889] | `--o †Eurysternidae Dollo, 1886 | |-- †Idiochelys fitzingeri Meyer, 1839 | |-- †Solnhofia parsonsi Gaffney, 1975b | |?- †Palaeomedusa testa Meyer, 1860 [Eurysternum crassipes Wagner 1859; Thalassemys marina Fraas 1903] | `--o †Eurysternum von Meyer, 1839 [Achelonia; Acichelys; Aplax; Euryaspis; Hydropelta; Palaeomedusa] | |-- †E. wagleri Meyer, 1839 | |-- †E. radianus (Wagner, 1861) | |-- †E. ignoratum Bräm, 1965 | `?- †E.? neuquinum Fernandez & Fuente, 1988 `--+?= †Xinjiangchelyidae Nesov, 1990 `--o Centrocryptodira |?- †“Sinemys” wuerhoensis (Yeh, 1973) |?- †Meiolaniformes Sterli & de la Fuente, 2012 `--+?- †Osteopygis emarginatus Cope, 1868; U.Cret.-Pal. NA. [taxonomic history] |?- †“Macrobaenidae” Sukhanov, 1964 [monophyly undemonstrated, also treated as paracryptodires] |?-o †Sinemydidae Yeh, 1963 sensu Zhou & Rabi, 2015 [Macrobaenidae Sukhanov, 1964 in partim | |?- †Tienfucheloides undutus Nesov, 1978 | |?- †“Sinemys” wuerhoensis Yeh, 1973 | |--+--o †Kirgizemys Nesov & Khozatskii, 1973 [Nessov & Khozatsky] | | | |-- †K. exaratus Nesov & Khozatskii, 1978 | | | `-- †“K.” dmitrievi Nesov & Khozatskii, 1981 | | `--+-- †Judithemys sukhanovi Parham & Hutchison, 2003 | | `-- †Changmachelys bohlini Brinkman, Yuan, Q. Ji, D. Ji, & You, 2013 | |-- ̸Xiaochelys ningchengensis Zhou & Rabi, 2015; E.Cret. NEAs. | `--+--+-- †Liaochelys jianchangensis Zhou, 2010 | | `--o †Ordosemys Brinkman & Peng, 1994 | | |?- †O. brinkmania | | |?- †O. perforata | | |-- †O. liaoxiensis (Ji, 1995) [Manchurochelys liaoxiensis Ji, 1995] | | `-- †O. leios Brickman & Peng, 1993 | `--+-- †Manchurochelys manchoukuoensis Endo & Shikama, 1932 | `--+-- †Dracochelys bicuspis Gaffney & Yah, 1992 [Gaffney & Ye], L. Cret CAs. | `--o †Sinemys Wiman, 1930 | |-- †S. lens Wiman, 1930 | `-- †S. gamera Brinkman & Peng, 1993 `--+-- †Judithemys sukhanovi Parham & Hutchison, 2003 |--o †Hangaiemys Sukhanov & Narmandakh, 1974 [a macrobaenid sensu Sukhanov, 2000] | |?- †H. kansuensis (Bohlin, 1953) [“Osteopygis” kansuensis Bohlin, 1953] | |-- †H. hoburensis Sukhanov & Narmandakh, 1974 | `-- †“H. leptis” [Sukhanov & Narmandakh, in press] [nomen nudum] `-- Polycryptodira [Cryptodira sensu Parham & Hutchison, 2003] (Recent cryptodirans; nykypystytaivuttajat)
After Zhou & Rabi, 2015<==o Pan-Cryptodira |-- †Plesiochelyidae `--+-- †Eurysternidae `--+--+-- †Xinjiangchelyidae | `--o †Sinemydidae | |--+-- †Kirgizemys | | `--+-- †Judithemys | | `-- †Changmachelys | `--+-- ̸Xiaochelys | `--+--+-- †Liaochelys | | `-- †Ordosemys | `--+-- †Manchurochelys | `--+-- †Dracochelys | `-- †Sinemys `--o Cryptodira |-- Chelydridae (snapping turtles; näykkijäkilpikonnat) `--o Procoelocryptodira |--o Chelonioidea Bauer, 1893 (sea turtles; merikilpikonnat) | |-- “Chelonioinea” [Cheloniidae sensu lato; Cheloniidae sensu Hirayama, 1997; Pancheloniidae Joyce, Parham & Gauthier, 2004] (karettikilpikonnansukuiset merikilpikonnat) | `-- Dermochelyoidea Baur, 1888 (merinahkakilpikonnansukuiset merikilpikonnat; letherback turtle-like turtles) `--o Chelomacryptodira |?- †Planetochelys |?- Trionychoidea (pehmeäkilpikonnat) `?- Testudinoidea |-- †Kayentachelys aprix Gaffney, Hutchison, Jenkins & Meeker, 1987; E.Jur, SWNAm. [Kayentachelyidae Gaffney, Hutchison, Jenkins & Meeker, 1987] |-- †Indochelys spatulata Datta, Manna, Ghosh & Das, 2000 [Indochelyidae Datta, Manna, Ghosh & Das, 2000] |?- †Camerochelys vilanovai Pérez-García & Murelaga, 2013 `--o Selmacryptodira Gaffney, Hutchinson, Jenkins & Meeker, 1987 |-- †Paracryptodira [=? Pleurosternoidea Hay, 1930] `--o Eucryptodira Gaffney, 1975a sensu Gaffney, 1984 [Daiocryptodira Milner, 2004] |--+-- | `--o †Eurysternidae Dollo, 1886 | |-- †Idiochelys fitzingeri Meyer, 1839 | |-- | |?- †Palaeomedusa testa Meyer, 1860 [Eurysternum crassipes Wagner 1859; Thalassemys marina Fraas 1903] | `--o †Eurysternum von Meyer, 1839 [Achelonia; Acichelys; Aplax; Euryaspis; Hydropelta; Palaeomedusa] | |-- †E. wagleri Meyer, 1839 | |-- †E. radianus (Wagner, 1861) | |-- †E. ignoratum Bräm, 1965 | `?- †E.? neuquinum Fernandez & Fuente, 1988 `--+?= `--o Centrocryptodira |?- †“Sinemys” wuerhoensis (Yeh, 1973) |?- †Meiolaniformes Sterli & de la Fuente, 2012 `--+?- †Osteopygis emarginatus Cope, 1868; U.Cret.-Pal. NA. [taxonomic history] |?- †“Macrobaenidae” Sukhanov, 1964 [monophyly undemonstrated, also treated as paracryptodires] |?-o †Sinemydidae Yeh, 1963 sensu Zhou & Rabi, 2015 [Macrobaenidae Sukhanov, 1964 in partim | |?- †Tienfucheloides undutus Nesov, 1978 | |?- †“Sinemys” wuerhoensis Yeh, 1973 | |--+--o †Kirgizemys Nesov & Khozatskii, 1973 [Nessov & Khozatsky] | | | |-- †K. exaratus Nesov & Khozatskii, 1978 | | | `-- †“K.” dmitrievi Nesov & Khozatskii, 1981 | | `--+-- †Judithemys sukhanovi Parham & Hutchison, 2003 | | `-- †Changmachelys bohlini Brinkman, Yuan, Q. Ji, D. Ji, & You, 2013 | |-- ̸Xiaochelys ningchengensis Zhou & Rabi, 2015; E.Cret. NEAs. | `--+--+-- †Liaochelys jianchangensis Zhou, 2010 | | `--o †Ordosemys Brinkman & Peng, 1994 | | |?- †O. brinkmania | | |?- †O. perforata | | |-- †O. liaoxiensis (Ji, 1995) [Manchurochelys liaoxiensis Ji, 1995] | | `-- †O. leios Brickman & Peng, 1993 | `--+-- †Manchurochelys manchoukuoensis Endo & Shikama, 1932 | `--+-- †Dracochelys bicuspis Gaffney & Yah, 1992 [Gaffney & Ye], L. Cret CAs. | `--o †Sinemys Wiman, 1930 | |-- †S. lens Wiman, 1930 | `-- †S. gamera Brinkman & Peng, 1993 `--+-- †Judithemys sukhanovi Parham & Hutchison, 2003 |--o †Hangaiemys Sukhanov & Narmandakh, 1974 [a macrobaenid sensu Sukhanov, 2000] | |?- †H. kansuensis (Bohlin, 1953) [“Osteopygis” kansuensis Bohlin, 1953] | |-- †H. hoburensis Sukhanov & Narmandakh, 1974 | `-- †“H. leptis” [Sukhanov & Narmandakh, in press] [nomen nudum] `-- Polycryptodira [Cryptodira sensu Parham & Hutchison, 2003] (Recent cryptodirans; nykypystytaivuttajat) |
Reference(s):
- Brickman, D. B., Stadtman, K. & Smith, D., 2000: New material of Dinochelys whitei Gaffney, 1979, from the Dry Mesa Quarry (Morrison Formation, Jurassic) of Colorado.
–Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 269-274 - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Datta, P. M., Manna, P., Ghosh, S. C. & Das, D. P., 2000: The first Jurassic turtle from India.
–Palaeontology: Vol. 43, #1, pp. 99-109 - Gaffney, E. S., Hutchison, J. H., Jenkins Jr, F. A. & Meeker, L. J., 1987: Modern turtle origins: the oldest known Cryptodire.
–Science: Vol. 237, #4812, pp. 289-291 [doi: 10.1126/science.237.4812.289] - Gaffney, E. S., Rich, T. H. V., Vickers-Rich, P., Constantine, A., Vacca, R. & Kool, L., 2007: Chubutemys, a new eucryptodiran turtle from the Early Cretaceous of Argentina, and the relationships of the Meiolaniidae.
–American Museum Novitates: #3599, pp. 1-35 - Joyce, W. G., 2000: The first complete skeleton of Solnhofia parsonsi (Cryptodira, Eurysternidae) from the Upper Jurassic of Germany and its taxonomic implications.
–Journal of Paleontology: Vol. 74, #4, pp. 684-700 - Meylan, P. A., 2001–: Testudines – Turtles, tortoises and terrapins.
–iNet: The Tree of Life Project: http://www.tolweb.org/tree?group=Testudines&contgroup=Amniota - Meylan, P. A. & Ganko, E., 1997: Higher Relationships of Testunides – The extinct and living turtles.
–iNet: The Tree of Life Project: [not available] - Meylan, P. A. & Ganko, E., 1997: †Kayentachelys.
–iNet: The Tree of Life Project: [not available] - Meylan, P. A. & Ganko, E., 1997: †Plesiochelyidae.
–iNet: The Tree of Life Project: [not available] - Meylan, P. A. & Ganko, E., 1997: †Meiolaniidae.
–iNet: The Tree of Life Project: [not available] - Parham, J. F., 2005: A reassessment of the referral of sea turtle skulls to the genus Osteopygis (Late Cretaceous, New Jersey, USA).
–Journal of Vertebrate Paleontology: Vol. 25, #1, pp. 71-77 - Parham, J. F. & Hutchison, J. H., 2003: A new eucryptodiran turtle from the Late Cretaceous of North America (Dinosaur Provincial Park; Alberta, Canada).
–Journal of Vertebrate Paleontology: Vol. 23, #4, pp. 783-798 - Pérez-García, A., 2012: High diversity of pancryptodiran turtles in the Lower Cretaceous of Europe.
–Cretaceous Research: Vol. 36, pp. 67-82 [doi: 10.1016/j.cretres.2012.02.004] - Pérez-García, A. & Murelaga, X., 2013: Camerochelys vilanovai gen. et sp. nov., a new pan-cryptodiran turtle in the Early Cretaceous of the Iberian Range (Spain).
–Cretaceous Research: Vol. 41, pp. 143-149 [doi: 10.1016/j.cretres.2012.12.002] - Sterli, J. & de la Fuente, M. S., 2012: New evidence from the Palaeocene of Patagonia (Argentina) on the evolution and palaeo-biogeography of Meiolaniformes (Testudinata, new taxon name).
–Journal of Systematic Palaeontology: Vol. 11, #7, pp. 835-852 [doi: 10.1080/14772019.2012.708674] - Sterli, J. & Joyce, W. G., 2007: The cranial anatomy of the Early Jurassic turtle Kayentachelys aprix.
–Acta Palaeontologica Polonica: Vol. 52 #4, pp. 675-694 - Sukhanov, V. R., 2000: Mesozoic turtles of Middle and Central Asia. 309-367
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000, xxxix-696 - Tong, H., Ji, S.-A. & Ji, Q., 2004: Ordosemys (Testudines: Cryptodira) from the Yixian Formation of Liaoning Province, Northeastern China: New Specimens and Systematic Revision.
–American Museum Novitates: #3438, pp. 1-20 [doi: 10.1206/0003-0082(2004)438<0001:OTCFTY>2.0.CO;2] - Zhou, C.-F. & Rabi, M., 2015: A sinemydid turtle from the Jehol Biota provides insights into the basal divergence of crown turtles.
–Scientific Reports: Vol. 5, art. nr. 16299 [doi: 10.1038/srep16299]