After Han et al., 2012, Rozadilla et al., 2016, and Herne et al., 2019
Note:
Subfamily-names for three subgroups of Hypsilophodontids; Othnielinae, Zephyrosaurinae, and Hypsilophodontinae were originally given by me in the very first versions of the Archive (“Life as we know It”) in 1995.
As the Archive was taken properly online in early 2000's, these pages were only reformatted, but not heavily revisited.
After that the Archive has grown more and more influential in general, and unfortunately these "unoffical" temporary names have escaped to the wider paleofield.
Thereby I have to formally renounce these three subfamily-names and I strongly recommend that people stop using them.
Especially as at least one of these groups has been formally named, as Thescelosauridae: Orodrominae Brown, Evans, Ryan & Russell, 2013
My only excuse of creating this taxonomic fumble is that I was young and stupid 3rd year student, with only preliminary knowledge of rules and principles of taxonomy, and had no clue how this would effect people.
--Mikko Haaramo, Helsinki, 2019-03-21
<==o †Ornithopoda Marsh, 1881b sensu Sereno, 1986 |--o †Orodrominae Brown, Evans, Ryan & Russell, 2013 [“Zephyrosaurines” [informal]] | |?- †Koreanosaurus boseongensis Huh, Lee, Kim, Lim & Godefroit, 2011 | |-- †Oryctodromeus cubicularis Varricchio, Martin & Katsura, 2007 | `--+-- †Albertadromeus syntarsus Brown, Evans, Ryan & Russell, 2013 | |-- †Orodromeus makelai Horner & Weishampel, 1988 | `-- †Zephyrosaurus schaffi Sues, 1980 `--+-- †Parksosaurus warreni (Parks, 1926a) Sternberg, 1937 [Thescelosaurus warreni Parks, 1926a] [Parksosauridae Bucholtz, 2002] `==o †“Elasmaria” Calvo, Porfiri, & Novas, 2007 [paraphyletic? (Herne et al., 2019)] |?- †Gasparinisaura cincosaltensis Coria & Salgado, 1996 `--+?- †Galleonosaurus dorisae Herne, Nair, Evans & Tait, 2019; E.Cret. Aust. |?- †Diluvicursor pickeringi Herne, Tait, Weisbecker, Hall, Nair, Cleeland & Salisbury, 2018; E.Cret. Aust. |?- †Leaellynasaura amicagraphica Rich & Rich, 1989 |?- †Qantassaurus intrepidus Rich & Vickers-Rich, 1999 |?- †Anabisetia saldiviai Coria & Calvo, 2002 |?- †Macrogryphosaurus gondwanicus Calvo, Porfiri & Novas, 2007 |?- †Morrosaurus antarcticus Rozadilla, Agnolin, Novas, Aranciaga Rolando, Motta, Lirio & Isasi, 2016 (published online 2015) |?- †Notohypsilophodon comodorensis Martínez, 1998 |?- †Telenkauen santacrucensis Novas, Cambiaso & Ambrosio, 2004 |?- †Trinisaura santamartaensis Coria, Moly, Reguero, Santillana & Marenssi, 2013 `--o †Clypeodonta |-- †Hypsilophodon foxii Huxley, 1869 `--+-- †Kangnasaurus coetzeei Haughton, 1915 |--o †Thescelosaurus Gilmore, 1913 [incl.? Bugenasaura Galton, 1995] [Thescelosauridae Sternberg, 1937] | |-- †T. neglectus Gilmore, 1913 [T. edmontonensis Sternberg] | |-- †T. garbanii Morris, 1976 [Buganasaura infernalis Galton, 1995; Bugenasaura garbanii (Morris) Galton, 1995] | `-- †T. assiniboiensis Brown, Boyd, & Russell, 2011 [“T.” sp. (Morris, 1976) (SDSM 7210)] `--o †Iguanodontoidea Dollo, 1888 sensu Weishampel & Heinrich, 1992 `--o †Iguanodontia Dollo, 1888 sensu Weishampel, & Heinrich, 1992, and McDonald et al., 2010 |--o †Tenontosaurus Ostrom, 1970 [Tenontosauridae] | |-- †T. tilletti Ostrom, 1970 | |-- †T. dossi Winkler, Murry & Jacobs, 1997 | `-- †T. sp. `--+--o †Rhabdodontidae Weishampel, Jianu, Csiki & Norman, 2003 | |?- †Matheronodon provincialis Godefroit, Garcia, Gomez, Stein, Cincotta, Lefèvre & Valentin, 2017 | |?-o †Mochlodon Seeley, 1881 | | |-- †M. suessi (Bünzel, 1871) [Iguanodon suessi Bünzel, 1871] | | `-- †M. vorosi | |-- †Muttaburrasaurus langdoni Bartholomai & Molnar, 1981 sensu McDonald, Kirkland, DeBlieux, Madsen, Cavin, Milner & Panzarin, 2010 | |--o †Rhabdodon Matheron, 1869 | | |-- †R. priscus Matheron, 1869 | | |-- †R. septimanicus Buffetaut & Le Loeuff, 1991 | | `-- †R sp. [Villeveyrac Rhabdodon] [Weishampel, Jianu, Csiki & Norman, 2003] | `--o †Zalmoxes Weishampel, Jianu, Csiki & Norman, 2003 [Mochlodon] | |-- †Z. robustus (Nopsca, 1902) Weishampel, Jianu, Csiki & Norman, 2003 [Mochlodon robustus Nopsca, 1902] | `-- †Z. shqiperorum Weishampel, Jianu, Csiki & Norman, 2003 `-- †Iguanodontia Dollo, 1888 sensu Sereno, 1986
Ornithopoda incertae sedis:
†Hypsilophodontidae genera et species indet. [Japan]
†Atlascopcosaurus loadsi Rich & Rich, 1989
†Fulgurotherium australe Huene, 1932 (nomen dubium)
†Laosaurus gracilis Marsh, 1878b (nomen dubium)
†Nanosaurus agilis Marsh, 1877b (nomen dubium)
†Siluosaurus zhanggiani Dong, 1997
Alternative phylogenies of Ornithopoda:
After Herne, Nair, Evans & Tait, 2019<==o †Ornithopoda |-- †Parksosaurus `--+-- †Macrogryphosaurus |-- †Telenkauen |-- †Gasparinisaura |-- †Leaellynasaura |-- †Diluvicursor |-- †Anabisetia |-- †Galleonosaurus |-- †Qantassaurus `--o †Clypeodonta |-- †Hypsilophodon `--o †Iguanodontia |--o †Rhabdodontidae | |-- †Rhabdodon | `--+-- †Mochlodon | `-- †Zalmoxes `--+-- †Muttaburrasaurus `--+-- Tenontosaurus `--o †Dryomorpha |-- †Owenodon |--+-- †Dysalotosaurus | `-- †Dryosaurus `-- †Ankylopollexia Reference(s):
|
After Rozadilla, Agnolin, Novas, Aranciaga Rolando, Motta, Lirio & Isasi, 2016<==o †Ornithopoda |-- †Hypsilophodon `--+-- †Thescelosaurus `--+--o †Elasmaria | |-- †Gasparinisaura | `--+-- †Morrosaurus | |-- †Trinisaura | |-- †Macrogryphosaurus | |-- †Notohypsilophodon | |-- †Telenkauen | `-- †Anabisetia `--+-- †Parksosaurus `--+-- †Kangnasaurus `--+-- †Rhabdodontidae `--+-- †Tenontosaurus `--+-- †Dryosauridae `-- †Ankylopollexia Reference(s):
|
After Brown, Evans, Ryan & Russell, 2013<==o †Ornithopoda |--o †Thescelosauridae | |--o †Orodrominae | | |--+-- †Oryctodromeus | | | `-- TMP 2008.045.002 | | `--+-- †Albertadromeus | | |-- †Orodromeus | | `-- †Zephyrosaurus | `--o †Thescelosaurinae | |-- †Parksosaurus | `--+-- †Haya | |-- †Jeholosaurus | |-- †Changchunsaurus | `-- †Thescelosaurus `--+-- †Hypsilophodon `--+-- †Gasparinisaura `--+-- †Tenontosaurus `--+-- †Rhabdodontidae `--+-- †Dryosauridae `-- †Ankylopollexia Reference(s):
|
After Han, Barrett, Butler & Xu, 2012<==o †Ornithopoda |--+-- †Orodromeus | `-- †Zephyrosaurus `--+--o †Jeholosauridae | |-- †Haya | `--+-- †Jeholosaurus | `-- †Changchunsaurus `--+-- †Hypsilophodon `--+-- †Thescelosaurus `--+-- †Telenkauen |--+-- †Parksosaurus | `-- †Gasparinisaura `--+-- †Rhabdodontidae `--+-- †Tenontosaurus `--+-- †Dryosauridae `-- †Ankylopollexia Reference(s):
|
After Boyd, Brown, Scheetz & Clarke, 2009 [based on datamatrix by Varricchio et al. (2007)]<==o †Ornithopoda |-- †Othnielosaurus `--+--+--+-- †Parksosaurus | | `-- †Thescelosaurus | `--+-- †Oryctodromeus | `--+-- †Orodromeus | `-- †Zephyrosaurus `--+-- †Hypsilophodon `--+-- †Gasparinisaura `--+-- †Tenontosaurus `--+-- †Rhabdodontidae `--+-- †Dryosauridae `-- †Ankylopollexia Reference(s):
|
After Weishampel, & Heinrich, 1992<==o †Ornithopoda |--o †Hypsilophodontidae | |-- †Thescelosaurus | `--+--+-- †Othnielia | | `-- †Yandusaurus | `--+--+-- †Orodromeus | | `-- †Zephyrosaurus | `--+-- †Hypsilophodon | `-- †Parksosaurus `--+-- †Tenontosaurus `--+-- †Dryosauridae `-- †Ankylopollexia Reference(s):
|
Reference(s):
- Barrett, P. M., 2000: The dinosaur fauna from the Earliest Cretaceous Tetori Group of Central Honshu, Japan.
–Journal of Vertebrate Paleontology: Vol. 20, #3, suppl. to #3, 28A - Boyd, C. A., Brown, C. M., Scheetz, R. D. & Clarke, J. A., 2009: Taxonomic Revison of the Basal Neornithischian Taxa Thescelosaurus and Bugenasaura.
–Journal of Vertebrate Paleontology: Vol. 29, #3, pp. 758-770 [doi: 10.1671/039.029.0328] - Brown, C. M., Evans, D. C., Ryan, M. J. & Russell, A. P., 2013: New Data on the Diversity and Abundance of Small-Bodied Ornithopods (Dinosauria, Ornithischia) from the Belly River Group (Campanian) of Alberta.
–Journal of Vertebrate Paleontology: Vol. 33, #3, pp. 495-520 [doi: 10.1080/039.033.0517] - Butler, R. J., Upchurch, P. & Norman, D. B., 2008: The phylogeny of the ornithischian dinosaurs.
–Journal of Systematic Palaeontology: Vol. 6, #1, pp. 1-40 [doi: 10.1017/S1477201907002271] - Calvo, J., Porfiri, J. & Novas, F., 2007: Discovery of a new ornithopod dinosaur from the Portezuelo Formation (Upper Cretaceous), Neuquén, Patagonia, Argentina.
–Arquivos do Museu Nacional: Vol. 65, pp. 471-483 - Coria, R. A., Moly, J. J., Reguero, M., Santillana, S. & Marenssi, S., 2013: A new ornithopod (Dinosauria; Ornithischia) from Antarctica.
–Cretaceous Research: Vol. 41, pp. 186-193 [doi: 10.1016/j.cretres.2012.12.004] - Godefroit, P., Garcia, G., Gomez, B., Stein, K., Cincotta, A., Lefèvre, U. & Valentin, X., 2017: Extreme tooth enlargement in a new Late Cretaceous rhabdodontid dinosaur from Southern France.
–Scientific Reports: Vol. 7, #1, Art.nr. 13098 [doi: 10.1038/s41598-017-13160-2] - Han, F.-L., Barrett, P. M., Butler, R. J. & Xu, X., 2012: Postcranial Anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China.
–Journal of Vertebrate Paleontology: Vol. 32, #6, pp. 1370-1395 [doi: 10.1080/02724634.2012.694385] - Herne, M. C., Nair, J. P., Evans, A. R. & Tait, A. M., 2019: New small-bodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian-Antarctic rift system, with revision of Qantassaurus intrepidus Rich and Vickers-Rich, 1999.
–Journal of Paleontology: in press (pp. 1-42) [doi: 10.1017/jpa.2018.95] - Herne, M. C., Tait, A. M., Weisbecker, V., Hall, M., Nair, J. P., Cleeland, M. & Salisbury, S. W., 2018: A new small-bodied ornithopod (Dinosauria, Ornithischia) from a deep, high-energy Early Cretaceous river of the Australian–Antarctic rift system.
–PeerJ: Vol. 5, Art.nr. e4113 [doi: 10.7717/peerj.4113] - Keesey, T. M., 1996-2000: The Dinosauricon
–iNet: The Dinosauricon: http://dinosauricon.com/ - Martinez, R. D., Lamanna, M. C., Smith, J. B., Dodson, P. & Luna, M., 2000: Dinosaurs from the Bajo Barreal Formation, Central Patagonia.
–Journal of Vertebrate Paleontology: Vol. 20, #3, suppl. to #3, 56A - McDonald, A. T., Kirkland, J. I., DeBlieux, D. D., Madsen, S. K., Cavin, J., Milner, A. R. C. & Panzarin, L., 2010: New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs.
–Plos One: Vol. 5, #11, pp. e14075 (1-35) [doi: 10.1371/journal.pone.0014075] - Novas, F. E., Cambiaso, A. V. & Ambrosio, A., 2004: A new basal iguanodontian (Dinosauria, Ornithischia) from the Upper Cretaceous of Patagonia.
–Ameghiniana: Vol. 41, #1, pp. 75-82 - Rozadilla, S., Agnolin, F. L., Novas, F. E., Aranciaga Rolando, A. M., Motta, M. J., Lirio, J. M. & Isasi, M. P., 2016: A new ornithopod (Dinosauria, Ornithischia) from the Upper Cretaceous of Antarctica and its palaeobiogeographical implications.
–Cretaceous Research: Vol. 57, pp. 311-324 [doi: 10.1016/j.cretres.2015.09.009] - Sues, H.-D., & Norman, D. B., 1990: Hypsilophodontidae, Tenontosaurus, Dryosauridae. 498-509
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990, xvi-733 - Varricchio, D. D., Martin, A. J. & Katsura, Y., 2007: First trace and body fossil evidence of a burrowing, denning dinosaur.
–Proceedings of the Royal Society of London: Biological series: #FirstCite Early Online Publishing, [DOI: 10.1098/rspb.2006.0443] [abstract only] - Weishampel, D. B. & Heinrich, R. E., 1992: Systematics of Hypsilophodontidae and basal Iguanodontia (Dinosauria: Ornithopoda).
–Historical Biology: Vol. 6, pp. 159-184 [doi: 10.1080/10292389209380426] - Weishampel, D. B., Jianu, C.-M., Csiki, Z. & Norman, D. B., 2003: Osteology and phylogeny of Zalmoxes (n. g.), an unusual Euornithopod dinosaur from the latest Cretaceous of Romania.
–Journal of Systematic Palaeontology: Vol. 12, #2, pp. 65-123
[from iNet abstract:] - Winkler, D. A., Murry, P. A. & Jacobs, L. L., 1997: A new species of Tenontosaurus (Dinosauria: Ornithopoda) from the Early Cretaceous of Texas.
–Journal of Vertebrate Paleontology: Vol. 17, #2, pp. 330-348 - Zan, S.-Q., Chen, J., Jin, L.-Y. & Li, T. 2005: A primitive ornithopod from the Early Cretaceous Quantou Formation of central Jilin, China.
–Vertebrata PalAsiatica 43:182-193 [from a correpondence in Dinosaur Mailing List]