After Xu et al., 2006, Butler, Upchurch & Norman, 2008, Han et al., 2012, Baron, Norman & Barrett, 2016, Salgado et al., 2017, and Herne et al., 2019
<==o †Ornithischia Seeley, 1888 [Predentata Marsh] (bird-hipped dinosaurs; nokkasaurit, linnunlanteiset dinosaurit) |?- †Pisanosaurus mertii Casamiquela, 1967 [recently treated as a Silesaurid (Agnolìn & Rozadilla, 2017)] `==o †“Fabrosauridae” Galton, 1972 [paraphyletic grade of basal ornithischians] |-- †Eocursor parvus Butler, Smith & Norman, 2007 |?-o †Gongbusaurus Dong, Zhou, & Zhang, 1983 | |?- †G. shiyii Dong, Zhou, & Zhang, 1983 | `-- †G. wucaiwanensis |?- †Xiaosaurus dashanpensis Dong & Tang, 1983; M.Jur. EAs. `--+-- †Heterodontosauridae Kuhn, 1966 [non Romer, 1966] `--o †Genasauria Sereno, 1986 |--o †Thyreophora Nopsca, 1915 (shield-carriers; kilpeäkantavat) | |?- †Fabrosaurus australis Ginsburg, 1964 sensu Sereno, 1991 [Lesothosaurus diagnosticus Galton, 1978; Lesothosaurus diagnisticus Ferigolo & Langer, 2006: 3 (lapsus calami); Stormbergia dangershoeki Butler, 2005 sensu Knoll et al., 2009] [paraphyletic Fabrosauridae] | |?- †Tatisaurus oehleri (Simmons, 1965) Simmons, 1965 [Scelidosaurus oehleri Simmons, 1965] | `--+--+-- †Laquintasaura venezuelae Barrett, Butler, Mundil, Scheyer, Irmis & Sanchez-Villagra, 2014 sensu Baron, Norman & Barrett, 2016 | | `-- †Scutellosaurus lawleri Colbert, 1981 | `--o Thyreophoroidea Nopsca, 1915 [Enoplosauria] | |-- †Emausaurus ernsti Haubold, 1990 | `--o Eurypoda Sereno, 1986 [Stegosauria + Ankylosauria] | |-- †Stegosauria Marsh, 1877 (stegosaurs; laattaselät) | `--o †Ankylosauromorpha sensu Carpenter, 2001 (Armored-plated dinosaurs) | |-- †Scelidosaurus harrisonii Owen, 1861 | `-- †Ankylosauria Osborn, 1923a (Armored-plated dinosaurs; panssariselät) `--o †Neornithischia Cooper, 1985 |?- †Stormbergia dangershoeki Butler, 2005 [= Lesothosaurus diagnosticus Galton, 1978 sensu Knoll et al., 2009] [paraphyletic Fabrosauridae] `--+-- †Agilisaurus louderbacki Peng, 1992 [paraphyletic Fabrosauridae] `--+--+-- †Hexinlusaurus multidens (He & Cai, 1983) Barrett, Butler & Knoll, 2005 [Yandusaurus multidens He & Cai, 1983; Agilisaurus multidens (He & Cai) Peng, 1990; Othnielia multidens (He & Cai) Paul, 1996]; M.Jur. EAs. | `-- †Yandusaurus hongheensis He, 1979 `--o †Ornithopoda Marsh, 1881b sensu Salgado, Canudo, Garrido, Moreno-Azanza, Martínez, Coria & Gasca, 2017 |-- †Kulindadromeus zabaikalicus Godefroit, Sinitsa, Dhouailly, Bolotsky, Sizov, McNamara, Benton & Spagna, 2014 [incl.? Daurosaurus olovus Alifanov & Saveliev, 2014; Kulindapteryx ukureica Alifanov & Saveliev, 2014] |-- †Isaberrysaura mollensis Salgado, Canudo, Garrido, Moreno-Azanza, Martínez, Coria & Gasca, 2017 `--+--o ?“Othnieliines” [informal] | |?- †Drinker nisti Bakker, Siegwarth & Filla, 1990 [= Drinker copei?] | |?- †Othnielia rex (Marsh, 1877b) Galton, 1977b [Nanosaurus rex Marsh, 1877b] (nomen dubium) | `-- †Othnielosaurus consors (Marsh, 1894) Galton, 2007 [Laosaurus cursors Marsh, 1894] [incl.? †Drinker nisti Bakker, Siegwarth & Filla, 1990; = Drinker copei?] `--o †Cerapoda Cooper, 1985 sensu Barrett et al., 2005 [Neornithischia sensu Sereno, 1996] |--o †Marginocephalia Sereno, 1986 (marginal-skulled; rajakalloiset) | |?- †Micropachycephalosaurus hongtuyanensis Dong, 1978 sensu Butler & Zhao, 2009 [Cerapoda incertae sedis] | |-- †Pachycephalosauria (Sternberg, 1945) Marya´nska & Osmólska, 1974 (thick-headed dinosaurs; paksupäät) | `-- †Ceratopsia Marsh, 1890 (horned dinosaurs; sarvinaamat) `--o †Ornithopoda Marsh, 1881b sensu Han, Barrett, Butler & Xu, 2012 (bird-feet; linnunjalat) |--o †Jeholosauridae Han, Barrett, Butler & Xu, 2012 | |-- †Haya griva Makovicky, Kilbourne, Sadleir & Norell, 2011 | `--+-- †Jeholosaurus shangyuanensis Xu, Wang & You, 2000 | `-- †Changchunsaurus parvus Zan, Chen, Jin & Li, 2005 `-- †Ornithopoda Marsh, 1881b sensu Herne, Nair, Evans & Tait, 2019 (bird-feet; linnunjalat)
Ornithischia incertae sedis:
†Alocodon kuehnei Thulborn, 1973 [nomen dubium, a tooth-taxa]
†Crosbysaurus harrisae Heckert, 2004; U.Trias. SWNA. [tooth-taxa]
†Galtonia gibbidens (Cope, 1878) Hunt & Lucas, 1994 [Thecodontosaurus gibbidens Cope, 1878] U.Trias. SWNA. [tooth-taxa]
†Krzyzanowskisaurus hunti (Heckert, 2002) Heckert, 2005 [Revueltosaurus hunti Heckert] U.Trias. SWNA. [tooth-taxa] [possibly jr. synon. of †Revueltosaurus callenderi Hunt, 1989 (Archosauria: Pseudosuchia: ?Suchia)]
†Lucianosaurus wildi Hunt & Lucas, 1994; U.Trias. SWNA. [tooth-taxa]
†Lusitanosaurus liasicus Lapparent & Zbyszewski, 1957
†Pekinosaurus olseni Hunt & Lucas, 1994; U.Trias. SWNA. [tooth-taxa]
†Phyllodon henkeli Thulborn, 1973
†Protecovasaurus lucasi Heckert, 2004; U.Trias. SWNA. [tooth-taxa]
†Taveirosaurus costai Antunes & Sigogneau-Russell, 1991 [tooth -taxa]
†Technosaurus smalli Chattarjee, 1984; U.Trias. SWNA. [tooth-taxa]
†Tecovasaurus murryi Hunt & Lucas, 1994; U.Trias. SWNA. [tooth-taxa]
†Trimucrodon cuneatus Thulborn, 1973 [tooth -taxa]
Alternative phylogenies of Basal Ornithischians
After Herne, Nair, Evans & Tait, 2019 (fig. 25/2)<==o †Ornithischia |-- †Heterodontosauridae `--+-- †Eocursor `--+--o †Thyreophora | |-- †Scutellosaurus | `--+-- †Emausaurus | `--+-- †Scelidosaurus | `--+-- †Stegosauria | `-- †Ankylosauria `--o †Neornithischia |-- †Lesothosaurus `--+-- †Agilisaurus `--+--+-- †Hexinlusaurus | `-- †Yandusaurus `--+--+-- Nanosaurus | `--+--o †Jeholosauridae | | |-- †Haya | | `--+-- †Jeholosaurus | | `-- †Changchunsaurus | `--o †Thescelosauridae | |--+-- †Orodromaeus | | `-- †Koreanosaurus | `--+-- †Zephyrosourus | `--+-- †Yueosaurus | `-- †Thescelosaurus `--o †Cerapoda |--o †Marginocephalia | |-- †Pachycephalosauria | `-- †Ceratopsia `--o †Ornithopoda |-- †Parksosaurus `--+-- †Macrogryphosaurus |-- †Telenkauen |-- †Gasparinisaura |-- †Leaellynasaura |-- †Diluvicursor |-- †Anabisetia |-- †Galleonosaurus |-- †Quantassaurus `--o †Clypeodonta |-- †Hypsilophodon `-- †Iguanodontia Reference(s):
|
After Salgado, Canudo, Garrido, Moreno-Azanza, Martínez, Coria & Gasca, 2017<==o †Ornithischia |-- †Pisanosaurus `--+-- †Eocursor |-- †Scutellosaurus |-- †Laquintasaura |-- †Stormbergia |-- †Lesothosaurus |-- †Heterodontosauridae |--o †Thyreophora | |-- †Emausaurus | `--+-- †Scelidosaurus | `--+-- †Stegosauria | `-- †Ankylosauria `--o †Neornithischia |-- †Agilisaurus `--+-- †Hexinlusaurus `==o †Ornithopoda |-- †Kulindadromeus |-- †Isaberrysaura `--+-- †Hypsilophodon |-- †Yandusaurus |-- †Othnieliosaurus |-- †Jeholosaurus |-- †Zephyrosourus |-- †Changchunsaurus |-- †Haya |-- †Orodromaeus |-- †Thescelosaurus `--+--o †Marginocephalia | |-- †Pachycephalosauria | `-- †Ceratopsia `--+-- †Talenkauen |--+-- †Parksosaurus | `-- †Gasparinisaura `-- †Iguanodontia Reference(s):
|
After Baron, Norman & Barrett, 2016<==o †Ornithischia |-- †Pisanosaurus `--+-- †Heterodontosauridae `--+-- †Eocursor `--+--o †Thyreophora | |--+-- †Laquintasaura | | `-- †Scutellosaurus | `--+-- †Emausaurus | `--+-- †Scelidosaurus | `--+-- †Stegosauria | `-- †Ankylosauria `--+-- †Lesothosaurus [incl. †Stormbergia] `--o †Neornithischia |-- †Agilisaurus `--+-- †Hexinlusaurus `--+-- †Othnieliosaurus `--+-- †Ornithopoda `--o †Marginocephalia |-- †Pachycephalosauria `-- †Ceratopsia Reference(s):
|
|
After Godefroit, Sinitsa, Dhouailly, Bolotsky, Sizov, McNamara, Benton & Spagna, 2014 (based on matrix of Han et al., 2012<==o †Ornithischia |-- †Pisanosaurus `--+-- †Heterodontosauridae `--+-- †Eocursor `--+-- †Lesothosaurus |--o †Thyreophora | |-- †Scutellosaurus | `--+-- †Emausaurus | `--+-- †Scelidosaurus | `--+-- †Stegosauria | `-- †Ankylosauria `--o †Neornithischia |-- †Stormbergia `--+-- †Agilisaurus `--+-- †Hexinlusaurus `--+-- †Kulindadromeus `--o †Cerapoda |-- †Ornithopoda `--o †Marginocephalia |-- †Pachycephalosauria `-- †Ceratopsia Reference(s):
|
After Han, Barrett, Butler & Xu, 2012<==o †Ornithischia |-- †Pisanosaurus `--+-- †Lesothosaurus |-- †Eocursor |-- †Heterodontosauridae |--o †Thyreophora | |-- †Scutellosaurus | `--+-- †Emausaurus | `--+-- †Scelidosaurus | `--+-- †Stegosauria | `-- †Ankylosauria `--o †Neornithischia |-- †Stormbergia `--+-- †Agilisaurus `--+-- †Hexinlusaurus `--+-- †Othnielosaurus `--o †Cerapoda |-- †Ornithopoda `--o †Marginocephalia |-- †Pachycephalosauria `-- †Ceratopsia Reference(s):
|
After Pol, Rauhut & Becerra, 2011<==o †Ornithischia |-- †Pisanosaurus |-- †Echinodon `--+-- †Heterodontosauridae `--+-- †Eocursor `--o †Genasauria |-- †Thyreophora `--+-- †Lesothosaurus `--+-- †Stormbergia `--+-- †Agilisaurus `--+-- †Hexinlusaurus `--+-- †Orodromeus `-- †Cerapoda Reference(s):
|
|
After Butler, Upchurch & Norman, 2008 (fig. 3)<==o †Ornithischia |-- †Pisanosaurus `--+-- †Heterodontosauridae `--+--o †Thyreophora | |-- †Lesothosaurus | `--+-- †Scutellosaurus | `--+-- †Emausaurus | `--+-- †Scelidosaurus | `--+-- †Stegosauria | `-- †Ankylosauria `--+-- †Stormbergia `--+-- †Agilisaurus `--+-- †Hexinlusaurus `--+-- †Othnielia `--+-- †Ornithopoda `--o †Marginocephalia |-- †Pachycephalosauria `-- †Ceratopsia Reference(s):
|
After Sereno, 1986 [via Butler et al., 2016]<==o †Ornithischia |-- †Lesothosaurus `--o †Genasauria |--o †Thyreophora | `--+-- †Scutellosaurus | `--+-- †Scelidosaurus | `--+-- †Stegosauria | `-- †Ankylosauria `--o †Cerapoda |--o †Ornithopoda | |-- †Heterodontosauridae | `-- †Euornithopoda `--o †Marginocephalia |-- †Pachycephalosauria `-- †Ceratopsia Reference(s):
|
After Sereno, 1984 [via Butler et al., 2008]<==o †Ornithischia |-- †Lesothosaurus `--+--o †Ornithopoda | |-- †Heterodontosauridae | `--+-- †Hypsilophodon | `--+-- †Camptosaurus | `--+-- †Tenontosaurus | `--+-- †Iguanodon | `--+-- †Ouranosaurus | `-- †Hadrosauridae `--o †Thyreophora |--+-- †Stegosauridae | `--+-- †Ankylosauridae | `-- †Nodosauridae `--+-- †Pachycephalosauridae `--+-- †Psittacosaurus `--+-- †Leptoceratops `--+-- †Bagaceratops `--+-- †Protoceratops `--+-- †Montanaceratops `-- †Ceratopidae |
After Norman, 1984 [via Butler et al., 2008]<==o †Ornithischia |--o †Thyreophora | |-- †Stegosauria | `--o †Ankylosauria | |-- †Ankylosauridae | `--+-- †Nodosauridae | `-- †Scelidosauridae `--+?- †Heterodontosauridae -pos1 `--o †Ornithopoda |--+-- †Fabrosauridae | `?- †Heterodontosauridae -pos2 `--+?- †Pachycephalosauria -pos1 `--+-- †Hypsilophodontidae `--+--+?- †Pachycephalosauria -pos2 | `-- †Ceratopsia `--+-- †Dryosauridae `--+-- †Iguanodontidae `-- †Hadrosauridae |
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