After Arbour & Evans, 2017, and Brown, Henderson, Vinther, Fletcher, Sistiaga, Herrera & Summons, 2017
<==o †Ankylosauria (Brown, 1908) Osborn, 1923 |--o †Nodosauridae Marsh, 1890a [incl. Polacanthidae Wieland, 1911 (in partim)] (nodosaurs; nodulipanssariselät) | |--+-- †Sauroplites scutiger Bohlin, 1953 | | |-- †Mymoorapelta maysi Kirkland & Carpenter, 1994 | | `-- †Dongyangopelta yangyanensis Chen, Zheng, Azuma, Shibata, Lou, Q. Jin & X. Jin, 2013 | `--+-- †Gastonia burgei Kirkland, 1998 | `--+-- †Gargoyleosaurus parkpinorum Carpenter, Miles, & Cloward, 1998 [Gargoyleosaurus parkpini Carpenter, Miles, & Cloward, 1998] | `--+--o †Polacanthus Owen, 1865 non Huxley, 1867 | | |-- †P. foxii Owen, 1865 non Hulke, 1881 | | `?- †P. rudgwickensis Blows, 1996 | `--+-- †Peloroplites cedrimontanus Carpenter, Bartlett, Bird & Barrick, 2008 | `--+-- †Sauropelta edwardsorum Ostrom, 1970a [emend. ????] [Sauropelta edwardsi Ostrom, 1970a] | `--+-- †Nodosaurus textilis Marsh, 1889b [Stegopelta landerensis Williston, 1905] | `--+-- †Niobrarasaurus coleii (Mehl, 1936) Carpenter, Dilkes & Weishampel, 1995 [Hierosaurus coleii Mehl, 1936] | `--+-- †Tatankacephalus cooneyorum WL Parsons & KM Parsons, 2009; E.Cret. WNAm. | `--+-- †Silvisaurus condrayi Eaton, 1960 | `--+-- †Amanantarx ramaljonesi Carpenter et al., 1999 | |--+-- †Hungarosaurus tormai Ősi, 2005 | | `--+-- †Europelta carbonensis Kirkland, Alcalá, Loewen, Espílez, Mampel & Wiersma, 2013 | | `--+-- †Borealopelta markmitchelli Brown, Henderson, Vinther, Fletcher, Sistiaga, Herrera & Summons, 2017 | | `-- †Pawpawsaurus campbelli Lee, 1996 | `--+--o †Struthiosaurinae Nopcsa, 1923 | | |-- †Stegopelta landerensis Williston, 1905 | | `--o †Struthiosaurus Bunzel, 1871 [taxonomic history] | | |-- †S. austriacus Bunzel, 1871 [taxonomic history] | | |-- †S. transylvanicus Nopcsa, 1915 [taxonomic history] | | `-- †S. languedocensis Carcia & Pereda Suberbiola, 2003 [taxonomic history] | `--+-- †Panoplosaurus mirus Lambe, 1919 | `==o †Edmontonia Sternberg, 1928 [ Chassternbergia Bakker, 1988; Denversaurus Bakker, 1988] | |-- †E. longiceps Sternberg, 1928 | |-- †E. rugosidens (Gilmore, 1930) [Paleoscincus rugosidens Gilmore, 1930; Chassternbergia rugosidens (Gilmore, 1930) Bakker, 1988] | `-- †Denversaurus schlessmani Bakker, 1988 [Edmontonia schlessmani (Bakker, 1988)] `--o †Ankylosauridae Brown, 1908 sensu Carpenter, 2001 [Syrmosauridae Maleev, 1952; incl. Polacanthidae Wieland, 1911 (in partim)] (nuijahäntäpanssariselät) |-- †Kunbarrasaurus ieversi Leahey, Molnar, Carpenter, Witmer & Salisbury, 2015 [Minmi sp. Molnar, 1996a] `--+-- †Hylaeosaurus armatus Mantell, 1833 [Synonyms] `--+-- †Liaoningosaurus paradoxus Xu, Wang & You, 2001 |-- †Crichtonpelta benxiensis (Lü, Ji, Gao & Li, 2007) Arbour &am; Currie, 2015 [Crichtonsaurus benxiensis Lü, Ji, Gao & Li, 2007] |-- †Chuanqilong chaoyangensis Han, Zheng, Hu, Xu & Barrett, 2014 |-- †Cederpelta bilbayhallorum Carpenter, Kirkland, Burge & Bird, in Carpenter, 2001 |-- †Aletopelta coombsi Ford & Kirkland, in Carpenter, 2001 `--+--o Shamosaurinae Tumanova, 1983 | |-- †Gobisaurus domoculus Vickaryous, Russell, Currie & Zhao, 2001 [incl. Zhongyuansaurus luoyangensis Xu, Lu, Zhang, Jia, Hu, Zhang, Wu & Ji, 2007; Arbour & Currie, 2015] | `-- †Shamosaurus scutatus Tumanova, 1983 `--o †Ankylosaurinae Brown, 1908 |-- †Zhongyuansaurus luoyangensis Xu, Lu, Zhang, Jia, Hu, Zhang, Wu & Ji, 2007 `--+--o †Pinacosaurus Gilmore, 1930 [Syrmosaurus Maleev, 1952a] | |-- †P. grangeri Gilmore, 1933b [Pinacosaurus ninghsiensis Young, 1935b; Syrmosaurus viminocaudus Maleev, 1952a; Syrmosaurus viminicaudus Maleev, 1954] | `-- †P. mephistocephalus Godefroit, Pereda Suberbiola & Dong, 1999 `--+--+-- †Tsagantegia longicranialis Tumanova, 1993 | `--+-- †Talarurus plicatospineus Maleev, 1952b | `-- †Nodocephalosaurus kirtlandensis Sullivan, 1999 `--+--+-- †Saichania chulsanensis Marya´nska, 1977 [Tianzhenosaurus youngi Pang & Cheng, 1998; Shanxia tienzhenensis Barrett, You, Upchurch & Burton, 1998 (Arbour &am; Currie, 2015)] | `--+-- †Zaraapelta nomadis Arbour, Currie & Badamgarav, 2014 | `-- †Tarchia kielanae Marya´nska, 1977 [Minotaurasaurus ramachandrani CA Miles & CJ Miles, 2009 (Arbour &am; Currie, 2015)] L.Cret. CAs. `--o †Ankylosaurini Brown, 1908 sensu Arbour & Currie, 2015 |-- †Ziapelta sanjuanensis Arbour, Burns, Sullivan, Lucas, & Cantrell, 2014 |-- †Euoplocephalus tutus (Lambe, 1902) Lambe, 1910 [Synonyms] `--+-- †Zuul crurivastator Arbour & Evans, 2017 |--o †Scolosaurus Nopcsa, 1928 [Oohkotokia Penkalski, 2014] | |-- †S. cutleri Nopcsa, 1928 [Oohkotokia horneri Penkalski, 2014] | `-- †S. thronus Penkalski, 2018 |-- †Dyoplosaurus acutosquameus Parks, 1924 |--o †Anodontosaurus lambei Sternberg, 1929 | |-- †A. inceptus Penkalski, 2018 | `-- †A. lambei Sternberg, 1929 `-- †Ankylosaurus magniventris Brown, 1908
Ankylosauria incertae sedis & nomina dubia:
†Crichtonsaurus bohlini Dong, 2002; nomen dudium (Arbour &am; Currie, 2015)
†Loricosaurus sculatus von Huene, 1929 [Titanosaurid scutes]
†Paleoscincus africanus Broom, 1910
†Paleoscincus asper Lambe, 1902
†Paleoscincus costatus Leidy, 1856
†Paleoscincus latus Marsh, 1892 [Pachycephalosaurid tooth]
†Stegosaurides excavatus Bohlin, 1953
†Stegosaurus madagascariensis Piveteau, 1926 [Ankylosauria indet.] (Maidment et al., 2008)
“Polacanthidae”
†Hoplitosaurus marshi (Lucas, 1901) Lucas, 1902a [Stegosaurus marshi Lucas, 1901; Polacanthus marshi (Lucas, 1901)]
†Nodosauridae
†Anoplosaurus curtonotus Seeley, 1879
†Antarctopelta oliveroi Salgado & Gasparini, 2006
†Propanoplosaurus marylandicus Stanford, Weishampel & Deleon, 2011
†Texasetes pleurohalio Coombs, 1995
†Taohelong jinchengensis Yang, You, Li & Hong, 2013
†Priconodon crassus Marsh, 1888
†Liaoningosaurus paradoxus Xu, Wang & You, 2001
†Dracopelta zbyszewskii Galton, 1980a
†Hierosaurus sternbergii Wieland, 1909
†Sarcolestes leedsi Lydekker, 1893c
†Glyptodontopelta mimus Ford, 2000 [Edmontonia australis Ford, 2000]
†Nodosaurus ischyrus (Seeley, 1881)
Ankylosauridae
†Acanthopholis eucercus Seeley, 1869
†Acanthopholis horrida Huxley, 1867 [Syngonosaurus Seeley, 1879; Eucercosaurus Seeley, 1879]
†Acanthopholis macrocercus Seeley, 1869; in partim [=? Acanthopholis horrida Huxley, 1867]
†Acanthopholis platypus Seeley, 1869; in partim [=? Acanthopholis horrida Huxley, 1867]
†Acanthopholis stereocercus Seeley, 1869; in partim
†Ahshislepelta minor Burns & Sullivan, 2011
†Amtosaurus magnus Kurzanov (Kuzanov) & Tumanova, 1978
†Anoplosaurus major Seeley, 1879; in partim
†Bissektipelta archibaldi (Averianov, 2002) [Amtosaurus archibaldi Averianov, 2002]
†Brachypodosaurus gravis Chakravarti, 1934
†Chassternbergia rugosidens Bakker, 1988
†Cryptodraco eumerus (Seeley, 1869)
†Dyoplosaurus giganteus Maleev, 1956 [Tarchia gigantea (Maleev, 1956) Marya´nska, 1977] nomen dubium (Arbour, Currie & Badamgarav, 2014)
†Glyptodontopelta mimus Ford, 2000
†“Hanwulosaurus» nomen nudum
†Hieshansaurus pachycephalus Bohlin, 1953
†Lametasaurus indicus Matleyi, 1923
†Maleevus disparoserratus (Maleev, 1952a) Tumanova, 1987 [Syrmosaurus disparoserratus Maleev, 1952a]
†Minmi paravertebra Molnar, 1980 (nomen dubium)
†Onychosaurus hungaricus Nopcsa, 1902
†Polacanthoides ponderosus Nopcsa, 1929
†Peishansaurus philemys Bohlin, 1953 [Pachycephalosaurid jaw fragment]
†Pinacosaurus ninghsiensis Young, 1935
†Priodontognathus phillipsi (Seeley, 1869) [Carpenter states year as 1969, I think not...]
†Regnosaurus northamptoni Mantell, 1848
†Rhadinosaurus alcinus Seeley, 1881
†Rhodanosaurus ludgunensis Nopcsa, 1929 [Struthiosaurus ludgunensis (Nopcsa, 1929)]
†Sangonghesaurus Chao, 1983
†Sarcolestes leedsi Lydekker, 1893
†Syngonosaurus macrocercus Seeley, 1879
†Telarurus disparoserratus Maleev, 1952
†Tianchisaurus nedegoatpeferima Dong, 1993 [Jurassosaurus nedegoapeferkimoruma Dong, vide Holden, 1993; Tianchiasaurus nedegoapeferima Dong, 1993, emend. Dong, 1994] (Jurassic Park ankylosaur)
†Tianzhenosaurus youngi Peng & Cheng, 1998 sensu Lü, Ji, Gao & Li, 2007
Alternative phylogenies of Ankylosauria:
After Arbour & Currie, 2016:<==o †Ankylosauria |-- †Nodosauridae `--o †Ankylosauridae |-- †Minmi `--+-- †Liaoningosaurus `--+-- †Cederpelta `--+-- †Gobisaurus `--+-- †Shamosaurus `--+-- †Zhongyuansaurus |-- †Tsagantegia |--+-- †Shanxia | `--+-- †“Crichtonsaurus” benxiensis [Crichtonpelta] | `--+-- †Dyoplosaurus | `-- †Pinacosaurus mephistocephalus `--+--+-- †Ankylosaurus | `-- †Euoplocephalus `--+--+-- †Minotaurasaurus | `-- †Pinacosaurus grangeri `--+-- †Nodocephalosaurus `--+--+-- †Talarurus | `-- †Tianzhenosaurus `--+-- †Saichania `-- †Tarchia |
After Thompson et al., 2012 (via Arbour & Currie, 2016):<==o †Ankylosauria |-- †Nodosauridae `--o †Ankylosauridae |-- †Minmi `--+-- †Liaoningosaurus `--+-- †Cederpelta `--+-- †Gobisaurus `--+-- †Shamosaurus `--+-- †Zhongyuansaurus |-- †Tsagantegia |--+-- †Shanxia | `--+-- †“Crichtonsaurus” benxiensis [Crichtonpelta] | `--+-- †Dyoplosaurus | `-- †Pinacosaurus mephistocephalus `--+--+-- †Ankylosaurus | `-- †Euoplocephalus `--+--+-- †Minotaurasaurus | `-- †Pinacosaurus grangeri `--+-- †Nodocephalosaurus `--+--+-- †Talarurus | `-- †Tianzhenosaurus `--+-- †Saichania `-- †Tarchia |
After W. L. Parsons & K. M. Parsons, 2009<==o Ankylosauria |--o Nodosauridae | |-- †Sauropelta | |-- †Silvisaurus | `--+-- †Pawpawsaurus | `--+-- †Panoplosaurus | `-- †Edmontonia `--o †Ankylosauridae |-- †Cederpelta `--+-- †Minmi `--+-- †Shamosaurus |-- †Gobisaurus `--+-- †Tsagantegia |--+-- †Gastonia | `-- †Tatankacephalus `--+== †Pinacosaurus |-- †Tianzhenosaurus |-- †Talarurus |-- †Tarchia |-- †Saichania `--+-- †Ankylosaurus `-- †Euoplocephalus |
After Hill, Witmer & Norell, 2003<==o Ankylosauria |-- Nodosauridae `--o †Ankylosauridae |-- †Gargoyleosaurus `--+-- †Gastonia `--+-- †Gobisaurus `--+-- †Minmi `--+-- †Pinacosaurus `--+-- †Talarurus `--+--o Shamosaurinae | |-- †Shamosaurus | `-- †Tsagantegia `--o †Ankylosaurinae |--+-- †Tarchia | `-- †Saichania |-- †Tianzhenosaurus |-- †Nodocephalosaurus |-- †Shanxia |-- †Ankylosaurus `-- †Euoplocephalus |
After Lü, Ji, Gao & Li, 2007<==o Ankylosauria |-- †Nodosauridae `--+-- †Polacanthidae `--o †Ankylosauridae |--o Shamosaurinae | |?- †Aletopelta | |?- †Cederpelta | `--+-- †Gobisaurus | `-- †Shamosaurus `--o †Ankylosaurinae |-- †Ankylosaurus `--+-- †Crichtonsaurus `--+--+-- †Tarchia | `--+-- †Saichania | `-- †Tsagantegia `--+-- †Talarurus `--+-- †Euoplocephalus `--+-- †Nodocephalosaurus |-- †Tianzhenosaurus `-- †Pinacosaurus |
After Carpenter, 2001:<==o †Ankylosauria |-- †Minmi `--+--o †Nodosauridae | |?- †Propanoplosaurus | |--+-- †Panoplosaurus | | `--+-- †Stegopelta | | `--+-- †Niobrarasaurus | | `--+-- †Texasetes | | `--+-- †Amanantarx | | `-- †Edmontonia | `--+-- †Anoplosaurus | `--+--+-- †Nodosaurus | | `-- †Struthiosaurus | `--+-- †Sauropelta | `--+-- †Pawpawsaurus | `-- †Silvisaurus `--o †Ankylosauroidea |--o †Polacanthidae | |--+-- †Gargoyleosaurus | | `--+-- †Hylaeosaurus | | `-- †Mymoorapelta | `--+-- †Gastonia | `--+-- †Hoplitosaurus | `-- †Polacanthus `--o †Ankylosauridae |--o Shamosaurinae | |?- †Aletopelta | |?- †Cederpelta | `--+-- †Gobisaurus | `-- †Shamosaurus `--o †Ankylosaurinae |-- †Ankylosaurus `--+?- †Crichtonsaurus `--+--+-- †Tarchia | `--+-- †Saichania | `-- †Tsagantegia `--+-- †Talarurus `--+-- †Euoplocephalus `--+-- †Nodocephalosaurus `-- †Pinacosaurus |
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