After
<==o Phascolosomatidae Stephen & Edmonds, 1972 |--o Phascolosoma Leuckart, 1828 [Phascolosomum Diesing, 1851; Phymosomum de Quatrefages, 1865; Phymosoma Selenka, 1883; Physcosoma Selenka, 1897] | |--o (Fischerana) Stephen, 1964 | | |-- P. (F.) capitatum (Gerould, 1913) [Physcosoma capitatum Gerould, 1913; Physcosoma abyssorum Southern, 1913; Goldfingia (Fisherana) savalovi Murina, 1974] | | `-- P. (F.) lobostomum Fischer, 1895 [Golfingia (Apionsoma) lobostomum Cutler, 1979; Fischerana wasini Lanchester, 1905] | `--o (Phascolosoma) Leuckart, 1828 | |--o P. (P.) agassizii Keferstein, 1866 | | |-- P. (P.) a. agassizii Keferstein, 1866 [Phascolosoma lordi Baird, 1868; Phascolosoma japonicum Grübe, 1877; Physcosoma glaucum Satí, 1930; Physcosoma yezoense Ikeda, 1924; Physcosoma formosense Satí, 1939; Phascolosoma (Rueppellisoma) golikovi Murina, 1975] | | `-- P. (P.) a. kurilense (Satí, 1937) [Physcosoma kurilense Satí, 1937] | |-- P. (P.) albolineatum Baird, 1868 [Phymosoma microdonton Sluiter, 1886; Phascolosoma multiannulata Wesenberg-Lund, 1954; Phascolosoma andamanensis Johnson, 1971] | |-- P. (P.) annulatum Hutton, 1879 [Phascolosoma scolops mossambiciense Augener, 1903; Phascolosoma scolops tasmaniense Fischer, 1914] | |-- P. (P.) arcuatum (Gray, 1828) [Siphunculus arcuatus Gray, 1828; Phymosoma lurco Selenka & de Man, 1883; Phascolosoma hizophora Sluiter, 1891; Physcosoma ambonense Fischer, 1896; Phymosoma deani Ikeda, 1905; Physcosoma esculenta Chen & Yeh, 1958] | |--o P. (P.) glabrum (Sluiter, 1902) [Physcosoma glabrum Sluiter, 1902] | | |-- P. (P.) g. glabrum (Sluiter, 1902) [Physcosoma glabrum Sluiter, 1902; Phymosoma microdontodon Shipley, 1898; Physcosoma funafutiense Fischer, 1914] | | `-- P. (P.) g. multiannulatum Wesenberg-Lund, 1954 | |-- P. (P.) granulatum Leuckart, 1828 [Sipunculus flavus Risso, 1826; Sipunculus tigrinus Risso, 1826; Sipunculus genuensis de Blainville, 1827; Phascolosoma laeve Cuvier, 1830; Sipunculus verrucosus Cuvier, 1830; Sipunculus levis Cuvier, 1830; Sipunculus papillosum Thompson, 1840; Sipunculus multitorquatus de Quatrefages, 1865; Phascolosoma fasciatum Baird, 1868; Phascolosoma jeffreysii Baird, 1868; Phascolosoma loveni Koren & Danielssen, 1877; Physcosoma herowardii Hrubel, 1903; Physcosoma lanzarotae Harms, 1921] | |-- P. (P.) maculatum (Sluiter, 1886) [Phymosoma maculatum Sluiter, 1886] | |-- P. (P.) meteori (Hrubel, 1904) [Phymosoma meteori Hrubel, 1904] | |-- P. (P.) nigrescens Keferstein, 1865 [Phascolosoma puntarenae Grübe & Oersted, 1858; Sipunculus (Phymosomum) plicatus de Quatrefages, 1865; Phascolosoma varians Keferstein, 1865; Phascolosoma agassizii Keferstein, 1867; Phascolosoma plainispinosum Baird, 1868; Phymososma spengeli Sluiter, 1886; Phymososoma duplicigranulatum Sluiter, 1886; Phymosoma lacteum Sluiter, 1886; Phymosoma diaphanes Sluiter, 1886; Physcosma extortum Sluiter, 1902; Physcosoma evisceratum Lanchester, 1905; Physcosoma minutum ten Broeke, 1925; Physcosoma horsti ten Broeke, 1925] | |-- P. (P.) noduliferum Stimpson, 1855 [Sipunculus tuberculatus Gray, 1828; Sipunculus (Phymosomum) nodulosus de Quaterfages, 1865; Sipunculus (Phymosomum) javenensis de Quatrefages, 1865; Phascolosoma grayi Baird, 1868] | |-- P. (P.) pacificum Keferstein, 1866 [Sipunculus (Phymosomum) javenensis de Quatrefages, 1865; Phasacolosoma asperum Grübe, 1868] | |-- P. (P.) perlucens Baird, 1868 [Sipunculus (Phascolosomum) vermiculus de Quatrefages, 1865; Phymosoma dentigerum Selenka & de Man, 1883; Physcosoma thomense Augener, 1903; Aspidosiphon insularis Lanchester, 1905; Physcosoma microdentigerum ten Broeke, 1925; Phascolosoma spinosum Johnson, 1971] | |-- P. (P.) saprophagicum Gibbs, 1987 | |-- P. (P.) scolops (Selenka & de Man, 1883) [Phymosoma scolops Selenka & de Man, 1883; Phymosoma scolops mossambiciense Selenka & de Man, 1883; Phascolosoma carneum Leuckart & Rüppell, 1828; Sipunculus (Phymosomum) guttatus de Quatrefahges, 1865; Phymosoma psaron Sluiter, 1886; Phymosoma spongicola Sluiter, 1902; Phymosoma nahaense Ikeda, 1904; Phymosoma socium Lanchester, 1905; Phascolosoma rottnesti Edmonds, 1956; Phascolosoma dunwichi Edmonds, 1956; Phascolosoma riukiuensis Murina, 1975] | |-- P. (P.) stephensoni (Stephen, 1942) [Physcosoma stephensoni Stephen, 1942; Phascolosoma laeve Keferstein, 1862; Sipunculus (Phymosomum) spinicauda de Quatrefages, 1865; Physcosoma lanzarotae Harms, 1921; Phascolosoma heronis Edmonds, 1956; Phascolosoma granulatum Murina, 1981] | `-- P. (P.) turnerae Rice, 1985 [Phascolosoma kapalum Edmonds, 1985] |--o Antillesoma Stephen & Edmonds, 1972 [Phascolosoma (Antillesoma) Stephen & Edmonds, 1972] | `-- A. antillarum (Grübe & Oersted, 1858) [Phascolosoma antillarum Grübe & Oersted, 1858; Phascolosoma fuscum Keferstein, 1862; Sipunculus (Aedematosomum) glans de Quatrefages, 1865; Phascolosoma nigriceps Baird, 1868; Phymosoma asser Selenka & de Man, 1883; Phymosoma pelam Selenka & de Man, 1883; Physcosoma weldoni Shipley, 1892; Phymosoma onomichianum Ikeda, 1904; Physcosoma gaudens Lanchester, 1905; Physcosoma similkis Chen & Yeh, 1958; Golfingia mokyevskii Murina, 1964; Phascolosoma schmidti Murina, 1975] `--o Apionsoma Sluiter, 1902 [Golfingia (Mitosiphon) Fischer, 1950; Golfingia (Phascolana) Wesenberg-Lund, 1959] |--o (Apionsoma) Sluiter, 1902 | |?- A. (A.) immunitum (Sluiter, 1902) [Phascolsosoma immunitum Sluiter 1902][sp. inquirenda] | |?- A. (A.) papilliferum (Keferstein, 1865) [Phascolsosoma papilliferum Keferstein, 1865][sp. inquirenda] | |?- A. (A.) reconditum (Sluiter, 1900) [Phascolsosoma reconditum Sluiter 1900][sp. inquirenda] | |-- A. (A.) misakianum (Ikeda, 1904) [Phascolosoma misakianum Ikeda, 1904; Phascolosoma hespera Chamberlin, 1920; Golfingia (Phascolana) longirostris Wesenberg-Lund, 1959; Golfingia (Phasacolana) tenuissima Wesenberg-Lund, 1959] | |---o A. (A.) murinae (Cutler, 1969) [Golfingia (Mitosiphon) murinae Cutler, 1969] | | |-- A. (A.) m. murinae (Cutler, 1969) [Golfingia murinae Cutler, 1969; Golfingia hespera Murina, 1964] | | `-- A. (A.) m. bilobatae (Cutler, 1969) [Golfingia (Mitosiphon) murinae bilobatae Cutler, 1969] | `-- A. (A.) trichoscephalus Luiter, 1902 [Phascolosoma pusillum Sluiter, 1912] `--o (Edmondsius) Gibbs & Cutler, 1987 [Phascolosoma (Satínus) Stephen & Edmonds, 1972] `-- A. (E.) pectinatum (Keferstein, 1867) [Phascolosoma pectinatum Keferstein, 1867; Phascolosoma abnorme Sluiter, 1886]
Reference(s):
Special thanks to:
Jos H. M. Dols, for gathering additional information and species of these animals.