After Sørensen, 2002 [fig. 2.]
<==o Gnathostomulida (kampaleukamadot) |--o Filospermoidea | |-- Haplognathia [Haplognathiidae] | `--o Pterognathiidae | |-- Cosmognathia | `-- Pterognathia `--o Bursovaginoidea |-- Paudicentula [Paudicentulidae] `--+--o Onychognathiidae | |-- Nanognathia | `--+-- Vampyrognathia | `--+-- Onychognathia | `--+-- Rastrognathia [Rastrognathiidae] | `-- Valvognathia `--+--+-- Gnathostomaria [Gnathostomariidae] | `--+-- Goannagnathia | `--+--o Mesognathariidae | | |-- Tenuignathia | | `--+-- Lapidognathia | | `-- Mesognatharia | `--+-- Clausognathia [Clausognathiidae] | `--o Agnathiellidae | |-- Paragnathiella | `-- Agnathiella `--+-- Problognathia [Problognathiidae] `--+--o Gnathostomulidae | |-- Corculognathia | `--+-- Ratugnathia | `--+-- Semaeognathia | `-- Gnathostomula [incl. Remanella paradoxa Meixner, nomen nudum] `--o Austrognathiidae [Conophoralia] |-- Triplignathia `--+-- Austrognatharia `-- Austrognathia
Reference(s):
- Parker, S. P. (ed.), 1982: Synopsis and classification of living organisms. Vols. 1 & 2
–McGrew-Hill Book Company - Sørensen, M. V., 2002: Phylogeny and jaw evolution in Gnathostomulida, with a cladistic analysis of the genera.
–Zoologica Scripta: Vol. 31, #5, pp. 461-480