After McKenna & Bell, 1997, Martín-Suárez & Mein, 1998, Ducroz, Volobouev & Granjon, 2001, Taylor, Denys & Mukerjee, 2003 (fig. 2b), Renaud et al., 2005, and “The Tree of Life”, with Recent species from Nowak, 1991
<==o Murinae Illiger, 1811 [taxonomic history] (Old World rats and mice; varsinaiset rotat & hiiret) |?-o †Huaxiamys Wu & L. Flynn, 1992; U.Mioc.-L.Plioc. As. [nomen nudum (McKenna & Bell, 1997)] | |-- †H. primitivus | `-- †H. downsi |?-o Deomyinae | |--o Acomys I. Geoffroy Saint-Hilaire, 1838 [taxonomic history] L.Plioc.-R. Af., Plioc. Eu., R. Mid., U.Pleist.-R. As. (spiny mice; okahiiret) [14] | | |-- A. cahirinus (okahiiri) | | |-- A. cilicicus | | |-- A. cinerasceus | | |-- A. ignitus | | |-- A. kempi | | |-- A. louisae | | |-- A. minous | | |-- A. mullah | | |-- A. nesiotes | | |-- A. percivali | | |-- A. russatus (kultaokahiiri) | | |-- A. spinosissimus | | |-- A. subspinosus | | `-- A. wilsoni | |-- Uranomys ruddi Dollman, 1909; R. Af. | `--o Lophuromys Peters, 1874 [Lasiomys Peters, 1866 (non Lasiomys Burmeister, 1854, a syn. of Sigmodon] [incl. Neanthomys Toschi, 1947; Kivumys Dieterlen, 1987] R. Af. [9] | |-- L. cinereus | |-- L. flavopunctatus | |-- L. luteogaster | |-- L. medicaudatus | |-- L. melanonyx | |-- L. nudicaudus | |-- L. rahmi | |-- L. sikapusi | `-- L. woosnami |?-o †Progonomys Schaup, 1938 | |?- †Progonomys debruijni Jacobs, 1978 | |?- †Progonomys yunnanensis Qiu & Storch, 1990 | |--+--+-- †Progonomys cathalai Schaup, 1938 | | | |-- †Progonomys woelferi Bachmayer & Wilson 1970 | | | |--o †Huerzelerimys Mein, Martín Suárez & Agustí, 1993; M.Mioc. Eu. | | | | |-- †H. turolensis | | | | `-- †H. vireti (Schaub, 1938) Mein et al., 1993 [Parapodemus vireti Schaub, 1938; Valerymys vireti (Schaub) Michaux, 1969] | | | `--o †Anthracomys Schaub, 1938; U.Mioc. NAf., U.Mioc.-L.Plioc. Eu. | | | `-- †A. lorenzi | | `--+?- †Parapodemus gaudryi (Dames, 1883) Schaub, 1938 [Mus gaudryi Dames, 1883] (nomen dubium, type lost) | | |?- Apodemus Kaup, 1826 [21] [relationship disputed by Martín-Suárez & Mein (1998)] | | |?-o †Rhagapodemus Kretzoi, 1959; U:Mioc.-L.Plioc. As., U.Mioc.-U.Plioc., Pleist. Eu., ?Plioc. Mid. | | | |?- †R. athensis de Bruijn & Van Der Meulen, 1975 | | | |?- †R. azzarolii Angelone & Kotsakis, 2001 | | | |?- †R. ballesioi Mein & Michaux, 1970 | | | |?- †R. debruijni | | | |?- †R. minor (Brandy, 1978) ==> †Rhagamys orthodon?? | | | |?- †R. vanderweerdi de Bruijn & van der Meulen, 1975 | | | `?-+-- †R. primaevus (Hugueney & Mein, 1965) | | | `--+-- †R. hautimagnensis Mein & Michaux, 1970 | | | `-- †R. frequens Kretzoi, 1959 | | `?- †Rhagamys orthodon (Hensel, 1856) Forsyth Major, 1905 [Mus orthodon Hensel, 1856] | `--+-- †Progonomys hispanicus | `--+--o †Castillomys Michaux, 1969 [incl. †Occitanomys Michaux, 1969; †Valerymys Michaux, 1969; †Hansdebruijnia Storch & Dahlmann, 1995] U.Mioc.-L.Plioc. As., U.Mioc.-L.Pleist. Eu., L.Plioc. Af. | | |-- †C gracilis | | `--+-- †C. crusafonti Michaux, 1969 | | `-- †C. rivas Martín-Suárez & Mein, 1991 [incl.? C. plinensis Ruiz Bustos, 1991; C. crusafonti meini Ruiz Bustos, 1991] | `--+-- †Centralomys de Giuli, 1989 | `--o †Occitanomys Michaux, 1969 [=? †Castillomys Michaux, 1969] | |?- †O. alcalai Adrover, Mein & Moissenet, 1988 | |?- †O. brailloni Michaux, 1969 | |?- †O. debruijni Sen, Jaeger, Dalfes, Mazin & Bocherens, 1989 | |?-o †(Hansdebruijnia) Storch & Dahlmann, 1995 | | `-- †O. (H.) neutrum | |-- †Occitanomys sondaari | `--+-- †Occitanomys adroveri (Thaler, 1966) Michaux, 1969 [Parapodemus adroveri Thaler, 1966] | `--o †Stephanomys Schaub, 1938; U.Mioc.-L.Pleist. Eu., L.Plioc. NAf. | |?- †S. margaritae Adrover, 1986 | |?- †S. vandeweerdi Adrover, 1986 | |-- †S. ramblensis | `--+-- †S. dubari | `--+-- †S. medius Cordy, 1976 | `--+--+-- †S. donnezani (Depéret, 1890) | | `-- †S. thaleri Lopéz-Martínez, Micheux & Hutterer, 1998 | |-- †S. calveti Bachelet & Castillo, 1990 | `--+-- †S. minor Gmelig Meyling & Michaux, 1973 | `--+-- †S. priaetaensis Aguilar, Micheux, Delannoy & Guendon, 1993 | `--+-- †S. balcellsi Gmelig Meyling & Michaux, 1973 | `-- †S. progressus Cordy, 1976 [nomen nudum López-Martínez et al., 1998, synon. of †S. balcellsi] |?-o “Philippine relics” | |--o Phloeomys Waterhouse, 1839; R. SEAs. [2] | | |-- P. cumingi (cuminginrotta) | | `-- P. pallidus | |--o Crateromys Thomas, 1895; R. SEAs. [3] | | |-- C. australis | | |-- C. paulus | | `-- C. schadenbergi (pörrönhäntäpilvirotta) | |--o Batomys Thomas, 1895 [incl. Mindanaomys Sanborn, 1953]; R. SEAs. [3] | | |-- B. dentatus | | |-- B. granti | | `-- B. salomonseni (mindanaonrotta) | `--o Carpomys Thomas, 1895; R. SWAs. [2] | |-- C. melanurus | `-- C. phaeurus |?-o “Praomys -group” | |--o Praomys Thomas, 1915; U.Plioc.-R. Af. [9] | | |-- P. delectorum | | |-- P. hartwigi | | |-- P. jacksoni (afrikanmetsärotta) | | |-- P. minor | | |-- P. misonnei | | |-- P. morio | | |-- P. mutoni | | |-- P. rostratus | | `-- P. tullbergi | |--o Zelotomys Osgood, 1910 [incl. Ochromys Thomas, 1920] U.Plioc.-R. Af. [2] | | |-- Z. hildegardeae | | `-- Z. woosnami | |--o Mastomys Thomas, 1915; U.Plioc.-R. Af. U.Pleist. SWAs. (multimammate rats) [8] | | |-- M. angolensis | | |-- M. coucha | | |-- M. erythroleucus | | |-- M. hildebrandtii | | |-- M. natalensis (moninisärotta) | | |-- M. pernanus | | |-- M. shortridgei | | `-- M. verheyeni | |--o Myomys Thomas, 1915 [Myomys (Myomyscus) Shortridge, 1942] R. Af., R. SWAs. [7] | | |-- M. albipes | | |-- M. daltoni | | |-- M. derooi | | |-- M. fumatus (afrikanniittyrotta) | | |-- M. ruppi | | |-- M. verreauxii | | `-- M. yemeni | |--o Hylomyscus Thomas, 1926; R. Af. [7] | | |-- H. aeta | | |-- H. alleni (kiipeilijähiiri) | | |-- H. baeri | | |-- H. carillus | | |-- H. denniae | | |-- H. parvus | | `-- H. stella | |--o Stenocephalemys Frick, 1914 [Stenocephalomys lapsus calami?] R. EAf. [2] | | |-- S. albocaudata | | `-- S. griseicauda | |-- Colomys goslingi Thomas & Wroughton, 1907 [Nilopegamys Osgood, 1928] Pleist.-R. CAf | `-- Heimyscus fumosus Misonne, 1969; R. Af. |?-o “Australasian group” [Hydromyinae] | |--o “Philippine relics II” (shrew-rats) | | |--o Apomys Mearns, 1905; R. SEAs. [8] | | | |-- A. abrae | | | |-- A. datae | | | |-- A. hylocoetes | | | |-- A. insignis | | | |-- A. littoralis | | | |-- A. microdon | | | |-- A. musculus | | | `-- A. sacobianus | | `--+--o Chrotomys Thomas, 1895; R. SEAs. [<Luzon, Mindoro>] [3] | | | |-- C. gonzalesi | | | |-- C. mindorensis | | | `-- C. whiteheadi | | |--o Rhynchomys Thomas, 1895; R. SEAs. (shrew rat) [2] | | | |-- R. isarogensis | | | `-- R. soricoides | | |-- Archboldomys luzonensis Musser, 1982; R. SEAs. [<Luzon>] | | `-- Celaenomys silaceus Thomas, 1898; R. SEAs. [<Luzon>] (luzoninrotta) | |--o “New Guinea group” [=? Anisomyini] | | |--o Hyomys Thomas, 1904; Pleist. R. Aust. [<N.Gui.>] | | | |-- H. dammermani | | | `-- H. goliath (karkeahäntärotta) | | |-- Anisomys imitator Thomas, 1904; Pleist. R. Aust. [<N.Gui.>] | | |--+--o Chiruromys Thomas, 1888; R. Aust. [<N.Gui.>] | | | | |-- C. forbesi (isopuuhiiri) | | | | |-- C. lamia | | | | `-- C. vates | | | `--o Pogonomys Milne-Edwards, 1877; R. Aust. Pac. | | | |-- P. championi | | | |-- P. loriae | | | |-- P. macrourus | | | `-- P. sylvestris | | |--o Coccymys Menzies, 1990; R. Aust. [N.Gui.] | | | |-- C. albidens | | | `-- C. ruemmleri | | |--o Mallomys Thomas, 1898 [Dentrosminthus de Vis, 1907] Pleist. R. Aust. [<N.Gui.>] | | | |-- M. aroaensis | | | |-- M. gunung | | | |-- M. istapantap | | | `-- M. rothschildi (pehmeähäntärotta) | | `--o Macruromys Stein, 1933; R. Aust. [<N.Gui.>] | | |-- M. elegans (lännenpienihammasrotta) | | `-- M. major | `--o Hydromyini [incl. Conilurini] | |--o Conilurini | | |--o Conilurus Ogilby, 1838 [Hapalotis Lichtenstein, 1829 (non Hapalotis Hüpner, 1816, a genus of butterflies)] L.Pleist.-R. Aust. (brush-tailed tree rat; white-footed tree rat; rabbit rat) | | | |-- C. albipes (white-footed tree rat) | | | `-- C. penicillatus (brush-tailed tree rat) | | |--o Leporillus Thomas, 1906; L.Pleist.-R. Aust. (stick-nest rats; tillikins) | | | |-- L. apicalis | | | `-- L. conditor (tikkupesärotta) | | `--o Mesembriomys Palmer, 1906 [Ammomys Thomas, 1906 (non Ammomys Bonaparte, 1831, a subgenus of Microtus?)] ?Pleist. R. Aust. (golden-backed tree rats; black-footed tree rats) | | |-- M. gouldii | | `-- M. macrurus | |--+-- Mastacomys Thomas, 1882 [treated as a synon. of Pseudomys by McKenna & Bell, 1997] | | |-- Pseudomys Gray, 1832 [taxonomic history] Plioc. L.Pleist.-R. Aust. (broad-toothed rat; Australian false mice; Australian field mice) [25] | | `--o Notomys Lesson, 1842 [incl. Podanomalus Waite, 1898; Thylacomys Waite, 1898; Ascopharynx Waite, 1900] Pleist.-R. Aust. (hopping mice) [9] | | |-- N. alexis | | |-- N. amplus | | |-- N. aquilo | | |-- N. cervinus | | |-- N. fuscus | | |-- N. longicaudatus | | |-- N. macrotis | | |-- N. mitchellii | | `-- N. mordax | |--o Zyzomys Thomas, 1909 [incl. Laomys Thomas, 1909] ?Pleist.-R. Aust. (rock rats) | | |-- Z. argurus | | |-- Z. maini | | |-- Z. palatilis | | |-- Z. pedunculatus | | `-- Z. woodwardi | |--o Leggadina Thomas, 1910 [Pseudomys (Leggadina) Thomas, 1910] Pleist.-R. Aust. | | |-- L. forresti | | `-- L. lakedownensis | |--o Uromys Peters, 1867 [Gymnomys Gray, 1867] [incl. Cyromys Thomas, 1910; Melanomys Winter, 1983 (non Melanomys Thomas, 1902, a synon. of Oryzomys, a sigmodontine murid)] Pleist. R. Aust., R. SEAs. Pac. | | |-- U. anak | | |-- U. caudimaculatus | | |-- U. hadrourus | | |-- U. imperator | | |-- U. neobritanicus | | |-- U. porculus | | |-- U. rex | | |-- (Cyromys) Thomas, 1910 | | `-- (Uromys) Peters, 1867 | |--+--o Solomys Thomas, 1922 [incl. Unicomys Troughton, 1935] R. Pac. [<Solomon Islands>] | | | |-- S. ponceleti | | | |-- S. salamonis | | | |-- S. salebrosus | | | |-- S. sapientis | | | `-- S. spriggsarum | | `--o Melomys Thomas, 1922 [incl. Paramelomys Rümmler, 1936; Mammelomys Menzies, 1996; Protochromys Menzies, 1996] Pleist.-R. Aust., R. SEAs. Pac. N.Gui. | | |-- M. aerosus | | |-- M. bougainville | | |-- M. burtoni | | |-- M. capensis | | |-- M. cervinipes | | |-- M. fellowsi | | |-- M. fraterculus | | |-- M. gracilis | | |-- M. lanosus | | |-- M. leucogaster | | |-- M. levipes | | |-- M. lorentzii | | |-- M. mollis | | |-- M. moncktoni | | |-- M. obiensis | | |-- M. platyops | | |-- M. rattoides | | |-- M. rubex (ylänköbanaanirotta) | | |-- †M. rubicola (Bramble Cay Melomys) [last seen on location 2009, not in survey on 2014, declared extinct 2019] | | |-- M. rufescens | | `-- M. spechti | `--o Hydromyini sensu stricto? | |--+-- Mayermys ellermani Laurie & Hill, 1954;; R. Aust. [N.Gui.] (yksihammaspäästäishiiri) | | |-- Neohydromys fuscus Laurie, 1952; R. Aust. [N.Gui.] (lyhythäntäpäästäishiiri) | | |--o Pseudohydromys Rümmler, 1934; R. Aust. [N.Gui.] | | | |-- P. murinus | | | `-- P. occidentalis | | |--o Microhydromys Tate & Archbold, 1941; R. Aust. [N.Gui.] | | | |-- M. musseri | | | `-- M. richardsoni (uurteishammaspäästäishiiri) | | `--o Paraleptomys Tate & Archbold, 1941; R. Aust. [N.Gui.] | | |-- P. rufilatus | | `-- P. wilhelmina | |--+-- Xeromys myoides Thomas, 1889; R. Aust. (valesuorotta) | | |-- Crossomys moncktoni Thomas, 1907; R. Aust. [N.Gui.] (korvatonvesirotta) | | `--o Leptomys Thomas, 1897; R. Aust. [N.Gui.] | | |-- L. elegans (pitkäjalkavesirotta) | | |-- L. ernstmayri | | `-- L. signatus | `--+-- Parahydromys asper Poche, 1906 [Drosomys Thomas, 1906; Limnomys Thomas, 1906 (non Limnomys Mearns, 1905, another murine)] (karkeakarvavesirotta) | `--o Hydromys É. Geoffroy Saint-Hilaire, 1804 [incl. Baiyankomys Hinton, 1943] Pleist-R. Aust. R. SEAs., R. Pac. Oc. [New Britan] | |-- H. chrysogaster (australianvesirotta) | |-- H. habbema (vuorivesirotta) | |-- H. hussoni | |-- H. neobrittanicus | `-- H. shawmayeri |?-o “Arvicanthines” | |--o Aethomys Thomas, 1915 [incl. Micaelamys Ellerman, 1941] [Epimys (Anthomys) Thomas, 1915; Rattus (Micaelamys) Ellerman, 1941] L.Plioc.-R. Af. [10] | | |-- A. bocagei | | |-- A. chrysophilus | | |-- A. granti | | |-- A. hindei (hindinpensasrotta) | | |-- A. kaiseri | | |-- A. namaquensis | | |-- A. nyikae | | |-- A. silindensis (silindanpensasrotta(!!"selindan")) | | |-- A. stannarius | | `-- A. thomasi | |--o Lemniscomys Trouessart, 1881; U.Plioc., ?L.Pleist., M.Pleist.-R. Af. [10] | | |-- L. barbarus (single-striped mouse; raitahiiri) | | |-- L. bellieri | | |-- L. griselda | | |-- L. hoogstraali | | |-- L. linulus | | |-- L. macculus | | |-- L. mittendorfi | | |-- L. rosalia | | |-- L. roseveari | | `-- L. striatus (täpläraitahiiri) | |--o Grammomys Thomas, 1915; U.Plioc.-R. Af (woodland mice) [11] | | |-- G. aridulus | | |-- G. buntingi | | |-- G. caniceps | | |-- G. cometes | | |-- G. dolichurus (puurotta) | | |-- G. dryas | | |-- G. gigas | | |-- G. ibeanus | | |-- G. macmillani | | |-- G. minnae | | `-- G. rutilans (peterinpuurotta) | |--o Oenomys Thomas, 1904; U.Plioc. R. Af. [2] | | |-- O. hypoxanthus | | `-- O. ornatus | |--o Thallomys | | |-- T. loringi | | |-- T. nigricauda | | |-- T. paedulcus | | `-- T. shortridgei | |--+-- Stochomys longicaudatus Thomas, 1926; R. Af. | | `--o Hybomys Thomas, 1910 [incl. Typomys Thomas, 1911] [6] | | |-- H. basilii | | |-- H. eisentrauti | | |-- H. lunaris | | |-- H. planifrons | | |-- H. trivirgatus (kolmijuovahiiri) | | `-- H. univittatus (peterinjuovahiiri) | |--+-- Desmomys harringtoni Thomas, 1910; R. EAf. | | `-- Rhabdomys pumilio Thomas, 1916; U.Plioc., ?Pleist., R. Af. (striped mouse; four-striped grass mouse) | `--+-- Mylomys dybowskii Thomas, 1906; R. Af. | |--o Arvicanthis Lesson, 1842 [taxonomic history] L.Plioc., U.Pleist.-R. SWAs., U.Plioc.-M.Pleist., ?U.Pleist., R., EAf. [5] | | |-- A. abyssinicus | | |-- A. blicki | | |-- A. nairobae | | |-- A. neumanni | | `-- A. niloticus (niilinrotta) | `--+--o Pelomys Peters, 1852 [Komemys de Beaux, 1924] L.Plioc. As.; U.Plioc.-R. Af. (groove-toothed rats) [5] | | |-- P. campanae | | |-- P. fallax | | |-- P. hopkinsi | | |-- P. isseli (afrikanpurorotta) | | `-- P. minor (pikkupurorotta) | `?-o Otomyini (laminate-toothed rats; korvarotat) | |--o †Euryotomys T.N. Pocock, 1976; L.Plioc. EAf. | | |-- †E. pelymoides | | `-- †E. bolti | `--o Parotomys Thomas, 1912 [incl. Liotomys Thomas, 1918] M.Pleist.-R. SAf. (whistling rats) [2] | |-- Parotomys littledalei | `--+-- Otomys unisulcatus | `--+-- Parotomys brantsii | `--o Otomys F.Cuvier, 1824 sensu stricto; U.Plioc.-R. Af. [taxonomic history] [12] | |?- †O. sp. cf. O. sloggetti | |?- †O. sp. cf. O. petteri | |-- O. sloggetti | `--+--+-- †O. sp. cf. O. gracilis | | `-- O. saundersiae | `--+-- O. denti | `--+-- O. irroratus | `--+-- O. tropicalis tropicalis | `--+--+-- †O. petteri | `--+-- O. angoniensis (angoninkorvarotta) | | `--+-- O. maximus (isokorvarotta) | | `-- O. tropicalis burtoni | `--+-- O. anchietae anchietae | |?- O. anchietae lacustris | `--+-- O. anchietae barbouri | `--+-- O. occidentalis | `--+-- O. laminatus | `--o O. typus | |-- O. t. jacksoni | `-- O. t. typus `?-o “Rattus -group” |?- Halmaheramys bokimekot Fabre, Pagès, Musser, Fitriana, Semiadi & Helgen, 2013 |--o Maxomys Sody, 1936; R. SEAs. [Indonesia & Philippines] | |-- M. alticola (vuoriokarotat) | |-- M. baeodon | |-- M. bartelsii | |-- M. dollmani | |-- M. hellwaldii | |-- M. hylomyoides | |-- M. inas | |-- M. inflatus | |-- M. moi | |-- M. musschenbroekii | |-- M. ochraceiventer | |-- M. pagensis | |-- M. panglima | |-- M. rajah | |-- M. surifer | |-- M. wattsi | `-- M. whiteheadi `--+--o “Dacnomys Division” | |?- Chiromyscus chiropus Thomas, 1925; R. SEAs. | |?- Saxatilomys paulinae Musser, Smith, Robinson & Lunde, 2005; R. SEAs. | |--o Niviventer Marshall, 1976; U.Plioc.-R. S.-SEAs. | | |-- N. andersoni | | |-- N. brahma | | |-- N. confucianus | | |-- N. coxingi | | |-- N. cremoriventer | | |-- N. culturatus | | |-- N. eha | | |-- N. excelsior | | |-- N. fulvescens | | |-- N. hinpoon | | |-- N. langbianis | | |-- N. lepturus | | |-- N. niviventer | | |-- N. rapit | | `-- N. tenaster | `--+-- Dacnomys millardi Thomas, 1916; R. S.-SEAs. | `--o Leopoldamys Ellerman, 1947; U.Plioc.-R. SEAs. | |-- L. edwardsi | |-- L. neilli | |-- L. sabanus | `-- L. siporanus `--+--o Berylmys Ellerman, 1947 [Rattus (Berylmys) Ellerman, 1947] L.-M.Pleist. R. As., R. SEAs. | |-- B. berdmorei (pienivalkohammasrotta) | |-- B. bowersi | |-- B. mackenziei | `-- B. manipulus |--o Tarsomys Mearns, 1905; R. SEAs. [Mindanao] | |-- T. apoensis | `-- T. echinatus |-- Limnomys sibuanus Mearns, 1905; R. SEAs. [Mindanao] |--o Bandicota Gray, 1873 [incl. Gunomys Thomas, 1907] R. As. (bandicoot rats) | |-- B. bengalensis (intianrotta) | |-- B. indica (intianisorotta) | `-- B. savilei |--o Bullimus Mearns, 1905; R. SEAs. [Philippines] | |-- B. bagobus | `-- B. luzonicus |--o Sundamys Musser & Newcomp, 1983; R. SEAs. | |-- S. infraluteus | |-- S. maxi | `-- S. Mülleri `-- Rattus Fischer de Waldhein, 1803 [taxonomic history] L.Pleist.-R., As., Aust.; M.Pleist.-R. SEAs..; R. Ind. Oc., Pac. Oc. (rats; rotat)
Murinae without phylogenetic association:
†Ratchaburimys Chaimanee, Suteethorn, Triamwichanon & Jaeger, 1996 Millardia Thomas, 1911 [incl. Grypomys Thomas, 1911; Guyia Thomas, 1917; Millardomys Sody, 1941] ?L.Pleist.-R. As. M. gleadowi M. kathleenae M. kondana M. meltada Cremnomys Wroughton, 191 [incl. Madromys Sody, 1941] R. SAs. C. blanfordi C. cutchicus C. elvira Diomys crumpi Thomas, 1917; R. SAs. Micromys Dehne, 1841; U.Mioc.-L.Plioc., Pleist.-R. As., U.Mioc.-R. Eu. †M. bendai †M. chalceus †M. cingulatus Storch & Dahlmann, 1995 ?†M. coronensis (Schaub, 1938) [†Parapodemus coronensis Schaub, 1938] †M. kozaniensis †M. paricioi †M. steffensi †M. tedfordi Wu & Flynn, 1992 †M. praeminutus M. minutus (Pallas, 1771) [incl. M. danubialis; M. agilis Dehne, 1841] (harvest mouse; vaivaishiiri) M. m. minutus M. m. soricinus M. m. subobscurus M. m. pratensis M. m. ussuricus Vandeleuria Gray, 1842; L.Pleist.-R. As. (Long-Tailed Tree Mice; palmuhiiret) V. nilagirica [ssp. => V. oleracea] V. nolthenii Phillips, 1929 (Ceylon Highland Long-Tailed Tree Mouse, Nolthenius's Long-tailed Climbing Mouse; ceyloninpalmuhiiri) V. oleracea (Bennett, 1832) (Indian Long-Tailed Tree Mouse; palmuhiiri) Vernaya fulva Anthony, 1941 [Octopdomys Sody, 1941] U.Plioc.-M.Pleist. R. As. Chiropodomys Peters, 1868 [incl. Insulaemus Taylor, 1934] U.Plioc., R. As. †C. penicillatus C. calamianensis C. gliroides (pensselihäntähiiri) C. karlkoopmani C. major C. muroides C. pusillus Hapalomys Blyth, 1859; U.Plioc., U.Pleist.-R. As. [2] H. delacouri H. longicaudatus Tokudaia Kuroda,1943 [Acanthomys Tokuda, 1941 (non Acanthomys Lesson, 1842, a synon. of Acomys; non Acanthomys Gray, 1867, a synon. of Rattus); Tokudamys Johnson, 1946] U.Pleist.-R. As. T. muenninki T. osimensis (japaninokahiiri) †Microtia elva Freudenthal, 1976; U.Mioc. Eu. †Paraethomys F.Petter, 1968 †P. abaigari Adrover, Mein & Moissenet, 1988 †P. athmeniae †P. belmezi Castillo, 1990 †P. meini (Micheux, 1969) †P. jaegeri Montenat & de Bruijn, 1976 †P. chikeri Jaeger, 1975 †P. miocaenicus †P. pusillus †P. anomalus †P. rbiae †P. darelbeidae †P. filfilae †Orientalomys de Bruijn & van der Meulen, 1975 [incl. †Euxinomys Sen, 1975] U.Mioc.-U.Plioc. As., L.-U.Plioc. Eu. †O. nihewanicus Zheng, 1981 †O. similis (Argyropulo & Pidoplichka, 1939) de Bruijn & van der Meulen, 1975 [Parapodemus similis Argyropulo & Pidoplichka, 1939] †O. galaticus †O. sibiricus †Chardinomys Jacobs & Li, 1982 [incl. Teilhardomys Cordy, 1977 (nomen nudum)] L.Plioc.-L.Pleist. As. †O. yusheensis †O. louisi †O. schaubi †Castromys Martín-Suárez & Freudenthal, 1994; U.Mioc. Eu. †C. inflatus †Beremendimys Kretzoi, 1956; Mus [taxonomic history] (mice; hiiret) [38] Muriculus imberbis Thomas, 1903; R. EAf. †Saidomys James & Slaughter, 1974; U.Mioc., U.Plioc. Af., L.Plioc. Eu. †S. afarensis †S. natrunensis James & Slaughter, 1974 †Yunomys wui Qiu & Storch, 1990 †Parapelomys Jacops, 1978; U.Mioc.-U.Plioc. As. Hadromys humei Thomas, 1911; L.Pleist.-R. As. Golunda ellioti Gray, 1837; ?U.Plioc., EAf; U.Plioc., Pleist., R., SAs. †Dilatomys Sen, 1983; L.Plioc. CAs. Dasymys Peters, 1875; U.Plioc.-L.Pleist. R. Af. [5] D. foxi D. incomtus (afrikansuorotta) D. montanus D. nudipes D. rufulus Lamottemys okuensis F.Petter, 1986; R. WAf. Dephomys Thomas, 1926; R. Af. [2] D. defua D. eburnea †Canariomys tamarani Crusafont-Pairó & F. Patter, 1964; Pleist.-R. At. [Canary islands, till 2000 B.P.] †Malpaisomys insularis Hutterer, López-Martínez & Michaux, 1988; Pleist.-R. At. [Canary islands] Nilopegamys Osgood, 1908; R. EAf. (Ethiopian water mouse) [treated as a synonym of Colomys elsewhere in McKenna & Bell, 1997] Malacomys Milne-Edwards, 1877; R. Af. [5] M. cansdalei M. edwardsi M. longipes (pitkäjalkarotta) M. lukolelae M. verschureni †Kritimys kiridus (Bate) Kuss & Misonne, 1968; M.Pleist. Mid. [Crete] Nesokia Gray, 1842 [Spalacomys Peters, 1861; Nesocia Blanford, 1891] [incl. Erythronesokia Khjuria, 1981] Plioc., ?L.Pleist. R. As., U.Pleist.-R. NEAf. (pest rats) [2] N. bunnii N. indica (lyhythäntärotta) Diplothrix legatus Thomas, 1916; U.Pleist.-R. EAs. [Ryukyu islands] Palawanomys furvus Musser & Newcomp, 1983; R. SWAs. [Palawan] Kadarsanomys sodyi Musser, 1981; SEAs. [Java] Tryphomys adustus Miller, 1910; R. SEAs. [Luzon] Abditomys latidens Musser, 1982; R. SEAs. [Luzon] Taeromys Sody, 1941 [incl. Arcuomys Sody, 1941] R. SEAs. [Sulawesi] [6] T. arcuatus T. callitrichus T. celebensis T. hamatus T. punicans T. taerae Paruromys Ellerman, in Laurie & Hill, 1954; R. SEAs. [Sulawesi] [2] P. dominator P. ursinus †Hooijeromys Musser, 1981; U.Pleist. SEAs. [Flores] Papagomys Sody, 1941; R. SEAs. [Flores] P. armandvillei P. theodorverhoeveni Komodomys rintjanus Musser & Boeadi, 1980; R. SEAs. [Rintja, Padar & Flores Islands] Bunomys Thomas, 1910 [incl. Frateromys Sody, 1941] R. SEAs. [Sulawesi] [7] B. andrewsi B. chrysocomus B. coelestis B. fratrorum B. heinrichi B. penitus B. prolatus Paulamys Musser, in Musser, van de Weerd & Strasser, 1986 [†Florensomys Musser, 1981, non Florensomys Fries, Hibbard & Dunkle, 1955, a geomyoid] R. SEAs. [Flores] †P. guanajuatoensis P. naso Stenomys Thomas, 1910 [incl. Nesoromys Thomas, 1920] R. SEAs. [Seram & New Guinea] [5] S. ceramicus S. niobe S. richardsoni S. vandeuseni S. verecundus (hoikkarotta) †Wushanomys Zheng, 1993; U.Plioc.-L.Pleist. EAs. †Qianomys Zheng, 1993; M.Pleist. EAs. Srilankamys ohiensis Musser, 1981; R. SAs. [Ceylon] Lenothrix canus Miller, 1903 R. SEAs. (gray tree rat) Pithecheir F.Cuvier, 1838 [Pithecochirus Gloger, 1841; Pithechirus Agassiz, 1842; Pitechirus Kaup, 1844 (lapsus calami); Pitcheir Schinz, 1845; Pithechir Jentink, 1892] [incl. Pithecheirops Emmons, 1993] R. SEAs. [2] P. melanurus P. parvus Eropeplus canus Miller & Hollister, 1921; R. SWAs. [Sulawesi] (Sulawesi soft-furred rat) Lenomys meyeri Thomas, 1898; R. SWAs. [Sulawesi] Margaretamys Musser, 1981; SWAs. [Sulawesi] [3] M. beccarii M. elegans M. parvus Haeromys Thomas, 1911; R. SWAs. [Borneo, Sulawesi, Palawan] [3] H. margarettae (raneehiiri) H. minahassae H. pusillus (pikkuraneehiiri) Anonymomys mindorensis Musser, 1981; R. SEAs. [Mindanao] Crunomys Thomas, 1897; R. SEAs. [4] C. celebensis C. fallax C. melanius C. rabori Lorentzimys nouhuysi Jentick, 1911; R. Aust. [N.Gui.] (uudenguineanhyppyhiiri) Pogonomelomys Rümmler, 1936 [incl. Abelomelomys Menzies, 1990] R. SEAs. Aust. [N.Gui.] P. bruijni P. mayeri P. sevia Xenuromys barbatus Tate & Archbold, 1941; R. Aust. [N.Gui.] (matkijapuurotta) †Spelaeomys florensis Hooijer, 1957; R. SEAs. [<Flores>] †Coryphomys buehleri Schaub, 1937; R. SEAs. [<Timor>] “Sulawesi shrew-rats” Echiothrix Gray, 1867 [Craurothrix Thomas, 1896] R. SEAs. [Sulawesi] E. centrosa Miller & Hollister, 1921 (Central Sulawesi spiny-rat) E. leucura Gray, 1867 (Sulawesi spiny-rat; sulawesinokarotta) Melasmothrix naso Miller & Hollister, 1921 (Sulawesian shrew-rat ) R. SWAs. [Sulawesi] Tateomys Musser, 1969; R. SWAs. [Sulawesi] T. macrocercus Musser, 1982 (Long-tailed shrew-rat) T. rhinogradoides Musser, 1969 (Tate's shrew-rat) Paucidentomys vermidax Esselstyn, Achmadi & Rowe, 2012 (longnose shrew-rat) R. SEAs. [Sulawesi] Sommeromys macrorhinos Musser & Durden, 2002 (Sommer’s Sulawesi Rat) R. SEAs. [Sulawesi]
Murinae incertae sedis:
†“Mus” hipparionum Schlosser, 1924 [Parapodemus hipparionum (Schlosser) Schaub, 1938; Karnimata hipparionum (Schlosser, 1924) Jacobs, 1978] †“Parapodemus” coronensis Schaub, 1938 [chimera?] †“Mastomys” colberti Lewis, 1939 [Parapodemus colberti (Lewis) Misonne, 1969; Rattus colberti (Lewis) Black, 1972; Karnimata colberti (Lewis) Jacobs, 1978] Thamnomys Thomas, 1907; U.Pleist.-R. Af. T. kempi Dollman, 1911 (Kemp's Thicket Rat) T. major Hatt, 1934 (Hatt's Thicket Rat) T. venustus Thomas, 1907 (Charming Thicket Rat)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Ducroz, J. F., Volobouev, V. & Granjon, L., 2001: An assessment of the systematics of arvicanthine rodents using mitochondrial DNA sequences: evolutionary and biogeographical implications.
–Journal of Mammalian Evolution: Vol. 8, #3, pp. 173-206 - Esselstyn, J. A., Achmadi, A. S. & Rowe, K. C., 2012: Evolutionary novelty in a rat with no molars.
–Biology Letters: in press [doi: 10.1098/rsbl.2012.0574] - Jacobs, L. L., 1977: A new genus of murid rodent from the Miocene of Pakistan and comments on the origin of the Muridae.
–PaleoBios: Vol. 25 [iNet Abstract at: http://www.ucmp.berkeley.edu/museum/pbios/abstracts_17to28.html] - López Antoñanzas, R., 2009: First Potwarmus from the Miocene of Saudi Arabia and the early phylogeny of murines (Rodentia: Muroidea).
–Zoological Journal of the Linnean Society: Vol. 156, #3, pp. 664-679 [doi: 10.1111/j.1096-3642.2008.00494.x] - Martín-Suárez, E. & Mein, P., 1998: Revision of the genera Parapodemus, Apodemus, Rhagamys and Rhagapodemus (Rodentia, Mammalia).
–Geobios: Vol. 31, #1, pp. 87-97 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Minwer-Barakat, R., García-Alix, A., Martín-Suárez, E. & Freudenthal, M., 2005: Muridae (Rodentia) from the Pliocene of Tollo de Chiclana (Grenada, southeastern Spain).
–Journal of Vertebrate Paleontology: Vol. 25, #2, pp. 426-441 - Musser, G. G., Smith, A. L., Robinson, M. F. & Lunde, D. P., 2005: Description of a new genus and species of rodent (Murinae, Muridae, Rodentia) from the Khammouan Limestone National Biodiversity Conservation Area in Lao PDR.
–American Museum Novitates: #3497, pp. 1-31 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629 - Qiu, Z. & Storch, G., 1990: New murids (Mammalia: Rodentia) from the Lufeng hominoid locality, Late Miocene of China.
–Journal of Vertebrate Paleontology: Vol. 10, #4, pp. 467-472 - Renaud, S., Benammi, M. & Jaeger, J.-J., 1999: Morphological evolution of the Murine rodent Paraethomys in response to climatic variations (Mio-Pleistocene of North Africa).
–Paleobiology: Vol. 25, #3, pp. 369-382
[stable url: http://links.jstor.org/sici?sici=0094-8373%28199922%2925%3A3%3C369%3AMEOTMR%3E2.0.CO%3B2-7] - Renaud, S., Michaux, J., Schmidt, D. N., Aguilar, J.-P., Mein, P. & Auffray, J.-C., 2005: Morphological evolution, ecological diversification and climate change in rodents.
†Proceedings of the Royal Society: Biological Sciences: Vol. 272, pp. 609-617 - Taylor, P. J., Denys, C. & Mukerjee, M., 2003: Phylogeny of the African murid tribe Otomyini (Rodentia), based on morphological and allozyme evidence.
–Zoologica Scripta: Vol. 33, pp. 389-402 - Zheng, S. & Cai, B., 1991: Fossil micromammals from the Donggou section of Dongyaozitou, Yuxian County, Hebei Province.
–Contributions to INQUA XIII: pp. 100-131