After McKenna & Bell, 1997, with Recent species from Nowak, 1991
<==o Echimyinae Gray, 1825 [taxonomic history] |-- †Callistomys Emmons & Vucetich, 1998 |-- Echimys (Echimys) semivillosus (I. Geoffroy Saint-Hilaire, 1838) [Makalata semivillosa] (Speckeled Tree Rat) `--+-- Echimys (Echimys) chrysurus (Zimmermann, 1780) (arboreal white-faced spiny rat, Whitefaced Tree Rat, Sauria[']) `--+--o Isothrix Wagner, 1845 [Lasiuromys Deville, 1852] Pleit.-R. SA. | |-- I. bistriata Wagner, 1845 (toro) | `-- I. pagurus Wagner, 1845 (Plain Brush-tailed Rat) |-- Makalata didelphoides (Desmarest, 1817) [Echimys didelphoides Desmarest, 1817; Mesomys didelphoides (Desmarest, 1817); Makalata armata (I. Geoffroy, 1838); Makalata castanea (Allen & Chapman, 1893); Makalata guianae (Thomas, 1888); Makalata hispida (Lichtenstein, 1830); Echimys longirostris Anthony, 1921] (Brazilian Spiny Tree Rat; Red-nosed Tree Rat) `--+-- Echimys (Echimys) grandis (Wagner, 1845) [Makalata grandis] (Giant Tree Rat, Toro['] Pre[^]to) `--+--o Echimys (Phyllomys) Lund, 1839 | |-- E. (P.) dasythrix Hensel, 1872 (Drab Atlantic Tree Rat) | |?- †E. (P.) fossilis Lund [nomen dubium] | |-- E. (P.) kerri (Moojen, 1950) | |-- E. (P.) lundi Leite, 2003 | |-- E. (P.) mantiqueirensis Leite, 2003 | |-- E. (P.) medius (Thomas, 1909) | |-- E. (P.) nigrispinus (Wagner, 1842) (Blackspined Atlantic Tree Rat) | |-- E. (P.) pattoni Emmons et al., 2002 | |-- E. (P.) thomasi (Ihering, 1897) (Giant Atlantic Tree Rat) | |-- E. (P.) unicolor (Wagner, 1842) | |-- E. (P.) brasiliensis Lund, 1840 | |-- E. (P.) lamarum (Thomas, 1916) (Pallid Atlantic Tree Rat) | `-- E. (P.) blainvillei (Jourdan, 1837) (Golden Atlantic Tree Rat) `--o Diplomys Thomas, 1916; R. CA.SA. |-- D. caniceps (Günther, 1876) (pehmeäpiikkirotta) |-- D. labilis (Bangs, 1901) (Central American Rufous Tree Rat, Gliding Spiny Rat) `-- D. rufodorsalis (J.A. Allen, 1899) (Red-crested Tree Rat)
Untreated Echimyinae:
Echimys G. Cuvier, 1809 [Loncheres Illiger, 1811; Echinomys Wagner, 1840] [incl. Nelomys F. Cuvier, 1837; Phyllomys Lund, 1839] Pleist.-R. SA. E. vieirai Iack-Ximenes, Vivo & Percequillo, 2005 (Echimys) E. (E.) braziliensis (Waterhouse, 1848) (Rusty-sided Atlantic Tree Rat) E. (E.) pictus (Pictet, 1841 (Painted Tree Rat) E. (E.) rhipidurus Thomas, 1928 [Makalata rhipidura (Thomas, 1928)] (Peruvian Tree Rat) E. (E.) saturnus Thomas, 1928 (Dark Tree Rat) Makalata Husson, 1978; R. Caribb.?, R. SA. M. macrura (Wagner, 1842) [Echimys macrurus Wagner, 1842] (Long-tailed armored tree-rat) M. carrikeri M. castanea M. flavida M. guianae M. longirostris M. obscura (Wagner, 1840) [Mesomys obscurus (Wagner, 1840)] (Dusky spiny tree-rat) M. occasius M. punctatus
Reference(s):
- Carvalho, G. A. S. & Salles, L. O., 2004: Relationships among extant and fossil echimyids (Rodentia: Hystricognathi).
–Zoological Journal of the Linnean Society: Vol. 142, #4, pp. 445-477 [doi: 10.1111/j.1096-3642.2004.00150.x] - Iack-Ximenes, G. E., de Vivo, M. & Percequillo, A. R., 2005: A new species of Echimys Cuvier, 1809 (Rodentia, Echimyidae) from Brazil.
–Papéis Avulsos de Zoologia (São Paulo): Vol. 45, pp. 51-60 [doi: 10.1590/S0031-10492005000500001] - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629