Holotheria

After Luo, Cifelli & Kielan-Jaworowska, 2001, Luo, Kielan-Jaworowska & Cifelli, 2002 and, Kielan-Jaworowska, Cifelli & Luo, 2004

Alternative phylogeny after Wang, Clemens, Hu & Li, 1998

Alternative phylogeny after Rougier, Novacek, McKenna & Wible, 2001:

Alternative phylogeny after McKenna & Bell, 1997.

<==o Holotheria Wible et al., 1995 sensu Luo, Cifelli & Kielan-Jaworowska, 2001 [Theria sensu lato] ("korkeammat nisäkkäät")
   |?- †Chronoperates Fox, Youzwyshyn & Krause, 1992 [Chronoperatidae Fox, Youzwyshyn & Krause, 1992]
   `--o Trechnotheria McKenna, 1975 [Yangotheria Chow & Rich, 1982]
      |--o †Amphiodontidae Simpson, 1925 [Amphiodontoidea Prothero, 1981]
      |  |-- †Amphiodon superstes Simpson, 1925 [Mammalia incertae sedis sensu Averianov (2002)]
      |  |?- †Gobiotheriodon infinitus (Trofimov, 1980) Trofimov, 1997 [Gobiodon Trofimov, 1980]
      |  `--+?- †Manchurodon simplicidens Yabe & Shikama, 1938 ["Symmetrodonta" incertae sedis sensu Averianov (2002)]
      |     `?- †Nakunodon paikasiensis Yadagiri, 1985 [Holotheria incertae sedis sensu Averianov (2002)]
      `--+?- †Argentoconodon fariasorum Rougier, Garrido, Gaetano, Puerta, Corbitt & Novacek, 2007, M.Jur. SSAm.
         |?- †Hallautherium schalchi Clemens, 1980
         |?- †Liaotherium gracile Zhou, Cheng & Wang, 1991
         |?- †Tendagurodon janenschi Heinrich, 1998
         |?-o †Dyskritodon Sigogneau-Russell, 1995
         |  |-- †D. amazighi Sigogneau-Russell, 1995
         |  `-- †D. indicus Prasad & Manhas, 2002
         |?-o †Jeholodentidae Luo, P. Chen, Li & M. Chen, 2007
         |  |-- †Yanoconodon allini Luo, P. Chen, Li & M. Chen, 2007
         |  `-- †Jeholodens jenkinsi Ji, Luo & Ji, 1999
         |--+-- †Gobiconodontidae Chow & Rich, 1984 sensu Luo, Kielan-Jaworowska & Cifelli, 2002
         |  `--o †Klameliidae Martin & Averianov, 2006
         |     |-- †Ferganodon narynensis Martin & Averianov, 2006
         |     `-- †Klamelia zhaopengi Chow & Rich, 1984 [an amphilestid sensu Kielan-Jaworowska, Cifelli & Luo, 2004]
         `--+-- †Amphilestidae Osborn, 1888 sensu Luo, Kielan-Jaworowska & Cifelli, 2002
            `--+-- †Spalacotheroidea Prothero, 1981 [Spalacotheriida Prothero, 1981, incl. Quirogatheria Bonaparte, 1992]
               `--o Symmetrodonta Simpson, 1925
                  |?-o †Thereuodon Sigogneau-Russell, 1989 [Dryolestida, or Zatheria indet., milk tooth (Averianov, 2002)]
                  |  |-- †T. dahmani Sigogneau-Russell 1989
                  |  `-- †T. taraktes Sigogneau-Russell & Ensom, 1998
                  |?- †Atlasodon monbaroni Sigogneau-Russell, 1991 [Dryolestida, or Zatheria indet., milk tooth (Averianov, 2002)]
                  |?- †Eurylambda aequicrurius Simpson, 1925a
                  |?- †Peralestes longirostris Owen, 1871
                  `--+?- †Tribactonodon bonfieldi Sigogneau-Russell, Hooker, Ensom, 2001
                     `--+?- †Triconodontidae Marsh, 1887 [Eutriconodonta Kermack, Musset & Rigney, 1973 sensu Luo, Cifelli & Kielan-Jaworowska, 2001] [falls back to basal Mammalia sensu Luo, Kielan-Jaworowska & Cifelli, 2002]
                        `--+-- †Zhangheotherium quinquecuspidens Hu, Wang, Luo & Li, 1997 [transfered to Spalacotheroidea by Li & Luo, 2006]
                           `-- Cladotheria McKenna, 1975 sensu Luo, Cifelli & Kielan-Jaworowska, 2001

After Gao, Wilson, Luo, Maga, Meng & Wang, 2009 (modified from Fig. 3)

<==o Holotheria
   |--+-- †Aploconodon
   |  `--+--o †Amphiodontidae
   |     |  |-- †Amphiodon
   |     |  |-- †Gobiotheriodon
   |     |  `--+-- †Manchurodon
   |     |     `-- †Nakunodon
   |     `--+-- †Hakusanodon
   |        |-- †Juchilestes
   |        `-- †Comodon
   |--o †Gobiconodontidae
   |  |-- †Gobiconodon
   |  `-- †Rapenomamus
   `--+--+--o †Jeholodentidae
      |  |  |-- †Jeholodens
      |  |  `-- †Yanoconodon
      |  `--o †Triconodontidae
      |     |-- †Priacodon
      |     `--+--+-- †Triconodon
      |        |  `-- †Trioracodon
      |        `--+-- †Arundeloconodon
      |           `--+-- †Astroconodon
      |              `--+-- †Alticonodon
      |                 `-- †Corvicondodon
      `--o Trechnotheria
         |-- †Phascolotherium
         |-- †Amphilestes
         `--+-- †Spalacotheroidea
            `-- Cladotheria

After Luo, P. Chen, Li & M. Chen, 2007

<==+-- †Fruitafossor
   |--+-- †Amphilestes
   |  `--+--o †Gobiconodontidae
   |     |  |-- †Gobiconodon
   |     |  `-- †Rapenomamus
   |     `--+--o †Jeholodentidae
   |        |  |-- †Jeholodens
   |        |  `-- †Yanoconodon
   |        `--o †Triconodontidae
   |           |-- †Trioracodon
   |           `-- †Priacodon
   `--+-- †Allotheria
      `--o Trechnotheria
         |-- †Spalacotheroidea
         `-- Cladotheria

After Rougier, Novacek, McKenna & Wible, 2001

<==o Holotheria
   `--o Trechnotheria
      |?- †Liaotherium
      |--+-- †Gobiconodontidae
      |  `-- †Triconodontidae
      `--+--+-- †Aploconodon
         |  `--+--o †Amphiodontidae
         |     |  |-- †Amphiodon
         |     |  |-- †Gobiotheriodon
         |     |  `--+-- †Manchurodon
         |     |     `-- †Nakunodon
         |     `--+-- †Hakusanodon
         |        `-- †Comodon
         `--+-- †Jeholodens
            |-- †Amphilestes
            |-- †Phascolotherium
            `--o Symmetrodonta
               |-- †Eurylambda
               `--+--+-- †Zhangheotherium
                  |  `--+-- †Peralestes
                  |     `-- †Spalacotheroidea
                  `--o Mammalia sensu stricto
                     |?- †Tribactonodon
                     |--o Yinotheria
                     |  |-- †Shuontherium
                     |  `-- Australophenida
                     `-- Cladotheria

Note:

This tree is based on alternative phylogeny of Holotherians presented by Rougier, Novacek, McKenna & Wible, 2001. While doing this tree I made some presumptions not expressed in their original tree (Fig. 6).

I presume that they chose Amphiodon to present all other Amphiodontids as well.
I presume that Eurylambda presents Symmetrodonta
I presume that Amphilestes presents Cladotheria
I also turned their node 21 to get Cladotheria to the bottom of a tree.

Where indoubt, I included sistergroups from Kielan-Jaworowska, Cifelli & Luo, 2002.

After Wang, Clemens, Hu & Li, 1998

<==o Holotheria
   |?- †Chronoperates
   |-- †Kuehneotheria
   `--o Trechnotheria
      |--o †Amphiodontidae
      |  |-- †Amphiodon
      |  |-- †Gobiotheriodon
      |  `--+-- †Manchurodon
      |     `-- †Nakunodon
      `--+-- †Spalacotheroidea
         `--o Symmetrodonta
            |?- †Asfaltomylos
            |?- †Atlasodon
            |?-o †Ausktribosphenidae
            |  |?- †Bishops
            |  `-- †Ausktribosphenos
            |?- †Casamiquelia
            |?- †Thereuodon
            |?- †Eurylambda
            |-- †Shuontherium
            `-- Cladotheria

After McKenna & Bell, 1997

<==o Holotheria
   |?- †Chronoperates
   |-- †Kuehneotheria
   `--o Trechnotheria
      |--o Symmetrodonta
      |  |?- †Casamiquelia
      |  |?- †Thereuodon
      |  |?- †Atlasodon
      |  |?- †Eurylambda
      |  |?- †Peralestes
      |  |?- †Shuontherium
      |  |--o †Amphiodontoidea
      |  |  `--o †Amphiodontidae
      |  |     |-- †Amphiodon
      |  |     |-- †Gobiotheriodon
      |  |     `--+-- †Manchurodon
      |  |        `-- †Nakunodon
      |  `-- †Spalacotheroidea
      `-- Cladotheria

Reference(s):

Averianov, A. O., 2002: Early Cretaceous "symmetrodont" mammal Gobiotheriodon from Mongolia and the classification of "Symmetrodonta".
–Acta Palaeontologica Polonica: Vol. 47, #4, pp. 705-716
Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698
Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698
Cifelli, R. L., Wible, J. R. & Jenkins, F. A. jr., 1998: Triconodont mammals from the Cloverly Formation (Lower Cretaceous), Montana and Wyoming.
–Journal of Vertebrate Paleontology: Vol. 18, #1, pp. 237-241
Dykes, T., 2002: Personal correspondence.
Dykes, T., 2001: Spalacotheriidae, Tinodontidae and other derived symmetrodonts.
–iNet: Mesozoic mammals – an internet directory: http://home.arcor.de/ktdykes/symmetro.htm
Heinrich, W.-D., 1998: Late Jurassic Mammals from Tendaguru, Tanzania, East Africa.
–Journal of Mammalian Evolution: Vol. 5, #4, pp. 269-290 [doi: 10.1023/A:1020548010203]
Gao, C., Wilson, G. P., Luo, Z.-X., Maga, A. M., Meng, Q. & Wang, X., 2009: A new mammal skull from the Lower Cretaceous of China with implications for the evolution of obtuse-angled molars and ‘amphilestid’ eutriconodonts.
–Proceedings of the Royal Society of London: Biological series: Vol. 277, #1, pp. 237-246 [doi: 10.1098/rspb.2009.1014]
Kielan-Jaworowska, Z., Cifelli, R. L. & Luo, Z.-X., 2002: Dentition and relationships of the Jurassic mammal Shuotherium.
–Acta Palaeontologica Polonica: Vol. 47, #3, pp. 479-486
Kielan-Jaworowska, Z., Cifelli, R. L. & Luo, Z.-X., 2004: Mammals from the Age of Dinosaurs – origins, evolution, and structure.
–Columbia University Press, New York, 2004, xv-630
Luo, Z. 1994: Sister-group relationships of mammals and transformations of diagnostic mammalian characters. 98-128.
in Fraser, N. C. & Sues, H-D. (eds.) 1994: In the Shadow of the Dinosaurs.
–Cambridge University Press, New York, 1994, x-435
Luo, Z.-X., Chen, P.-j., Li, G. & Chen, M., 2007: A new eutriconodont mammal and evolutionary development in early mammals.
–Nature: Vol. 446, pp. 288-293 [doi: 10.1038/nature05627]
Luo, Z.-X., Cifelli, R. L. & Kielan-Jaworowska, Z., 2001: Dual origin of tribosphenic mammals.
–Nature: Vol. 409, 4 January, pp. 53-57
Luo, Z.-X., Kielan-Jaworowska, Z. & Cifelli, R. L., 2002: In quest for a phylogeny of Mesozoic mammals.
–Acta Palaeontologica Polonica: Vol. 47, #1, pp. 1-78
Martin, T. & Averianov, A. O., 2006: A previously unrecognized group of Middle Jurassic triconodontan mamals from Central Asia.
–Naturwissenschaften: Vol. 94, #1, pp. 43-48 [doi: 10.1007/s00114-006-0135-5]
McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631
Rougier, G. W., Garrido, A., Gaetano, L., Puerta, P. F., Corbitt, C. & Novacek, M. J., 2007: First Jurassic triconodont from South America.
–American Museum Novitates: #3580, pp. 1-17
Rougier, G. W., Novacek, M. J., McKenna, M. C. & Wible, J. R., 2001: Gobiconodonts from the Early Cretaceous of Oshih (Ashile), Mongolia.
–American Museum Novitates: #3348, pp. 1-30
Rougier, G. W., Spurlin, B. K. & Kik, P. K., 2003: A New Specimen of Eurylambda aequicrurius and Considerations on “Symmetrodont” Dentition and Relationships.
–American Museum Novitates: #3398, pp. 1-15 [doi: 10.1206/0003-0082(2003)398<0001:ANSOEA>2.0.CO;2]
Wang, Y., Clemens, W. M., Hu, Y. & Li, C., 1998: A probable pseudo-tribosphenic upper molar from the Late Jurassic of China and the early radiation of the Holotheria.
–Journal of Vertebrate Paleontology: vol. 18, #4, pp. 777-787

Mikko's Phylogeny Archive

Holotheria – holotheres