After Rieppel, 1994, Evans & Barbadillo, 1997, Evans & Chure, 1998, Gao & Norell, 2000, and Alifanov, 2000
Alternative phylogeny of Scincomorpha after Kearney, 2003
<==o Scincomorpha Camp, 1923 (skink-like lizards; skinkkimäiset liskot) |?- †Scincomorpha genera et species indet. [Ceder Mountain formation, Utah, USA.] [Nydam, 2002] |?- †Scincomorpha genera et species indet. [Ukhaa Tolgod, Djadokhta formation, Mongolia] [Gao & Norell, 2000] |?- †Calanguban alamoi Simőes, Caldwell & Kellner, 2014; E.Cret. SAm. |?- †Bothriagenys mysterion Nydam, 2002 |?- †Globaura venusta Borsuk-Bialynicka, 1988 |-- †Tepexisaurus tepexii Reynoso & Callison, 2000 `--+--+?- †Changetisaurus estesi Federov & Nessov, 1992 [Chengetisaurus lapsus calami] [Dorsetisauridae: Changetisaurinae Federov & Nessov, 1992] | `--o †Paramacellodidae Estes, 1983 [a paraphyletic form-family?] | |?- †Atokasaurus metarsiodon Nydam & Cifelli, 2002b, L. Cret. NA. [cf. “Paramacellodidae”] | |?- †Pseudosaurillus becklesi Hoffstetter, 1967 | |?-o †Saurillus Owen, 1855 | | |-- †S. henkeli Hoffstetter, 1967 | | |-- †S. obtusus Owen, 1855 | | `-- †S. robustidens Hoffstetter, 1967 | |?- †Saurillodon proraformis Estes, 1983 | |--o †Paramacellodus Hoffstetter, 1967; U. Jur. L. Cret. NA. | | |?- cf. P. sp. [Ceder Mountain formation, Utah, USA.] [Nydam, 2002] | | |?- cf. †P. keebleri Nydam & Cifelli, 2002b, L. Cret. NA. | | |-- †P. oweni Hoffstetter, 1967 U. Jur. NA. | | `-- †P. keebleri Nydam & Cifelli, 2002b | |--o †Becklesius Estes, 1983 | | |-- †B. hoffstetteri (Seiffert, 1973) [Macellodus brodei Hoffsteetr, 1967 (non Macellodus brodei Owen, 1854); Becklesisaurus hoffstetteri Seiffert, 1973] | | |-- †B. cataphractus Richter, 1994 | | |-- †B. sp. [Richter, 1994] | | `-- †B. nopcsai Folie & Codrea, 2005 | |-- †Sharovisaurus karatauensis Hecht & Hecht, 1984; U. Jur. CAs. | `-- †Mimobecklesiosaurus gansuensis Li, 1985; U. Jur. EAs. |--o Scincoidea Oppel, 1811 (skink-related lizards; skinkinsukuiset liskot) | |?- †Dimekodontosaurus madseni Nydam, 2002 | |?- †Hymenosaurus clarki Gao & Norell, 2000 | |?- †Scincoideus haininensis Folie, Sigé & Smith, 2005, Pal. Eu. | |?- †Parmeosaurus scutatus Gao & Norell, 2000 | |?-o †Slavoiidae Alifanov, 1993 | | |-- †Eoxanta lacertifrons Borsuk-Bialynicka, 1988 | | `-- †Slavoia darevskii Sulimski, 1984 | |-- Scincidae [incl. Feyliniidae] (skinks; skinkit) [124/1206] | `--o Cordyloidea [Cordyliformes Fitzinger, 1826] | |-- Cordylidae Mertens, 1937 (non Gray, 1838?, non Fitzinger, 1826?) (Girdled Lizards, vyöliskot) | `-- Gerrhosauridae (kilpiliskot) `--o Lacertoidea Camp, 1923 (sisiliskonsukuiset liskot) |--o Xantusiomorpha | |?-o †Hodzhakuliidae Alifanov, 1993 sensu Alifanov, 2016 [previously treated as Anguimorpha incertae sedis] | | |-- †Bagaluus primigenius Alifanov, 2016 | | `--+-- †Hodzhakulia magna Nessov, 1985 | | `--+-- †“Platynotoides” altidentatus Alifanov, 2016 [Platynotoides Alifanov, 2016 (preocc.), non Platynotoides Kaszab, 1975 (a beetle)] | | `--+-- †Carnoscincus eublepharus Alifanov, 2016 | | `--o †Pachygenys Gao & Cheng, 1999; E.Cret. EAs. | | |-- †P. thlastesa Gao & Cheng, 1999 | | `-- †P. adachii Ikeda, Ota & Saegusa, 2015 | `-- Xantusiidae (yöliskot) `--o Lacertiformes Estes, de Queiroz & Gauthier, 1988 (sisiliskomaiset liskot) |--+?-o †Eolacertidae Cernanský & Smith, in press (2017) | | |-- †Eolacerta robusta Nöth, 1940 [however, see Anguimorpha for alternative position] | | `-- †Stefanikia siderea Cernanský & Smith, in press (2017) | `-- Lacertidae Bonaparte, 1831 (sisiliskot) [26/273] `--+--o †Meyasaurus Vidal, 1915 [Ilerdaesaurus] | |-- †M. faurai Vidal, 1915 (emend. von Huene, 1952) | |-- †M. diazromeri Evans & Barbadillo, 1997 | |-- †M. crusafonti (Hoffstetter, 1966) | `-- †M. unensis (Richter, 1994) `--+?- †Dalinghosaurus longidigitus Ji, 1998 `--+--o Teiioidea Estes, de Queiroz & Gauthier, 1988 (Teju-like lisards; teijunsukuiset liskot) | |?- †Haptosphenus placodon Estes, 1964 | `--+?- †Adamisaurus magnidentatus Sulimski, 1972 [Adamisauridae Sulimski, 1978] | |-- Teiidae Gray, 1827 (tejus; teijut) [Recent: 9 genera / 116 species] | `-- Gymnophthalmidae `--o †Boreoteiioidea Nydam, Eaton & Sankey, 2007 |?- †Tijubina pontei Bonfim-Júnior & Marques, 1997 |?- †Olindalacerta brasiliensis Evans & Yabumoto, 1998 |-- †Kuwajimalla kagaensis Evans & Manabe, 2008 |-- †Tianyusaurus zhengi Lü et al., 2008 |-- †Mongolochamopidae Alifanov, 1993 |-- †Macrocephalosauridae Sulimski, 1975 `-- †Polyglyphanodontidae Gilmore, 1943b
After Alifanov, 2016<==o Scincomorpha |--o Leptoglossa | |--+-- †Globauridae | | `--+-- †Ardeosauridae | | `-- †Carusiidae | `--+--o ?Cordyloidea | | |-- †Paramacellodidae | | `-- Cordylidae | `--o ?Scincoidea | |-- Acontiidae | `-- Scincidae `--o Xantusiomorpha |-- Xantusiidae `--+-- Dibamidae `--+--o †Eoxantidae | `-- †Eoxanta `--+--o †Slavoiidae | `-- †Slavoia `--o †Hodzhakuliidae |-- †Bagaluus `--+-- †Hodzhakulia `--+-- †“Platynotoides” `--+-- †Carnoscincus `-- †Pachygenys Reference(s):
|
After Kearney, 2003<==o Scincomorpha |--o Cordyliformes | `--o Cordyloidea | |-- Cordylidae | `-- Gerrhosauridae `--+--o Scincoidea | |?- †Dimekodontosaurus | |?- †Hymenosaurus | |?- †Parmeosaurus | `-- Scincidae `--+--o †Eichstaettisauridae | |-- †Eichstaettisaurus | `-- †Globaura `--+-- †Eoxanta `--+-- Xantusiidae |--o Lacertiformes | |-- Lacertidae Bonaparte, 1831 | `--+-- †Meyasaurus | `-- Teiidae `--+--o †Slavoiidae | `-- †Slavoia |--o †Adamisauridae | `-- †Adamisaurus `--+-- †Polyglyphanodontidae `--+-- †Sineoamphisbaena `-- †Macrocephalosauridae Reference(s):
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Reference(s):
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in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000, xxxix-696 - Alifanov, V. R., 2016: Lizards of the family Hodzhakuliidae (Scincomorpha) from the lower Cretaceous of Mongolia.
–Paleontological Journal: Vol. 50, #5, pp. 504-513 [doi: 10.1134/s0031030116050038] - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
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–W. H. Freeman and company, New York, 1988, 698. - Cernanský, A. & Smith, K. T., 2017: Eolacertidae: a new extinct clade of lizards from the Palaeogene; with comments on the origin of the dominant European reptile group – Lacertidae.
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–Bulletin of the American Museum of Natural History: #249, pp. 1-118 - Ikeda, T., Ota, H. & Saegusa, H., 2015: A new fossil lizard from the Lower Cretaceous Sasayama Group of Hyogo Prefecture, western Honshu, Japan.
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–Journal of Vertebrate Paleontology: Vol. 22, #3, pp. 645-660 - Nydam, R. L. & Cifelli, R. L., 2002: Lizards from the Lower Cretaceous (Aptian-Albian) Antlers and Cloverly Formations.
–Journal of Vertebrate Paleontology: Vol. 22, #2, pp. 286-298 - Reynoso, V.-H. & Callison, G., 2000: A new scincomorph lizard from the Early Cretaceous of Puebla, Mexico.
–Zoological Journal of the Linnean Society: vol. 130, #2, pp. 183-212. - Rieppel, O. 1994: The Lepidosauromorpha: an overview with special emphasis on the Squamata. 23-37.
in Fraser, N. C. & Sues, H-D. 1994: In the Shadow of the Dinosaurs.
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–iNet: http://www.reptile-database.org/