Mikko's Phylogeny Archive

Lacertidae – lacertid lizards

After Fu, 2000, with recent species from Uetz, Etzold & Chenna, 1995-2000

Genera only:

<==o Lacertidae Bonaparte, 1831 (lacertids; sisiliskot)
   |?-o †Amblyolacerta Olig. Mio. CEu.
   |  |-- †A. sp. Olig. CEu.
   |  `-- †A. dolnicensis Rocek, 1984; Mio. CEu.
   |?-o †Dormaalisaurus Augé & Smith, 2002; L.Eoc. WEu.
   |  |-- †D. girardoti Augé & Smith, 2002
   |  `-- †D. rossmanni Augé & Smith, 2002
   |?- †Dracaenosaurus croizeti Hoffstetter, 1944; Olig. SEu.
   |?- †Miolacerta Rocek [=? Takydromus], L. Mio. CEu.
   |?- †Plesiolacerta lydekkeri Hoffstetter, 1942; Eoc. Olig. Eu.
   |?- †Pseudeumeces cadurcensis (Filhol, 1877) Hoffstetter, 1944; Olig. SEu.
   |?- †Ligerosaurus pouiti (Augé, 1993) Augé, Bailon & Malfay, 2003 [Pseudeumeces pouiti Augé, 1993]; Mio. SEu.
   |--o Gallotiinae
   |  |-- Gallotia
   |  `-- Psammodromus (hietajuoksijat)
   `--o Lacertinae
      |-- Takydromus Daudin, 1802 [Tachysaurus Gray, 1845; Platyplacopus Boulenger, 1917; Apeltonotus Boulenger, 1917] (itä-aasiansisiliskot; East Asian grass lizards) [17]
      `--+--+-- Algyroides
         |  |-- Podarcis
         |  |-- Timon [sg. => Lacerta (Timon)?]
         |  |?- Darevskia [sg. => Lacerta (Darevskia)?]
         |  `==o Lacerta [paraphyletic]
         |     |-- (Apathya)
         |     |-- (Archaeolacerta)
         |     |-- (Omanosaura)
         |     |-- (Teira)
         |     `-- (Zootoca)
         `--+--+-- Mesalina
            |  `-- Acanthodactylus (ripsusisiliskot)
            `--+--+-- Nucras
               |  `--+-- Latastia
               |     `-- Heliobolus
               `--+--+-- Adolfus
                  |  |--o Eremias
                  |  |  |-- (Ommateremias)
                  |  |  |-- (Pareremias)
                  |  |  |-- (Rhabderemias)
                  |  |  `-- (Scapteira)
                  |  `-- Ophisops
                  `--+-- Meroles
                     `--+-- Pedioplanis
                        `-- Tropidosaura

Lacertids not treated by Fu, 2000:

    Australolacerta
    Eremiascincus
    Gastropholis
    Holaspis
    Ichnotropis
    Philochortus
    Poromera
    Pseuderemias

With Species:

<==o Lacertidae Bonaparte, 1831 (lacertids; sisiliskot)
   |?-o †Amblyolacerta Olig. Mio. CEu.
   |  |-- †A. sp. Olig. CEu.
   |  `-- †A. dolnicensis Rocek, 1984; Mio. CEu.
   |?-o †Dormaalisaurus Augé & Smith, 2002; L.Eoc. WEu.
   |  |-- †D. girardoti Augé & Smith, 2002
   |  `-- †D. rossmanni Augé & Smith, 2002
   |?- †Dracaenosaurus croizeti Hoffstetter, 1944; Olig. SEu.
   |?- †Miolacerta Rocek [=? Takydromus], L. Mio. CEu.
   |?- †Plesiolacerta lydekkeri Hoffstetter, 1942; Eoc. Olig. Eu.
   |?- †Pseudeumeces cadurcensis (Filhol, 1877) Hoffstetter, 1944; Olig. SEu.
   |?- †Ligerosaurus pouiti (Augé, 1993) Augé, Bailon & Malfay, 2003 [Pseudeumeces pouiti Augé, 1993]; Mio. SEu.
   |--o Gallotiinae
   |  |--o Gallotia
   |  |  |-- G. atlantica 
   |  |  |-- G. caesaris
   |  |  |-- G. galloti
   |  |  |-- G. goliath
   |  |  |-- G. gomerana Hutterer, 1985 [G. goliath gomerana Hutterer, 1985]
   |  |  |-- G. intermedia
   |  |  |-- G. simonyi
   |  |  `-- G. stehlini
   |  `--o Psammodromus (hietajuoksijat)
   |     |-- P. algirus (algerianhietajuoksija)
   |     |-- P. blanci
   |     |-- P. hispanicus (espanjanhietajuoksija)
   |     `-- P. microdactylus
   `--o Lacertinae
      |-- Takydromus Daudin, 1802 [Tachysaurus Gray, 1845; Platyplacopus Boulenger, 1917; Apeltonotus Boulenger, 1917] (itä-aasiansisiliskot; East Asian grass lizards)
      `--+--+--o Algyroides
         |  |  |-- A. fitzingeri
         |  |  |-- A. marchi
         |  |  |-- A. moreoticus
         |  |  `-- A. nigropunctatus
         |  |--o Podarcis
         |  |  |-- P. bocagei
         |  |  |-- P. dugesii
         |  |  |-- P. erhardii
         |  |  |-- P. filfolensis
         |  |  |-- P. gaigeae
         |  |  |-- P. graeca
         |  |  |-- P. hispanica
         |  |  |-- P. lilfordi
         |  |  |-- P. melisellensis
         |  |  |-- P. milensis
         |  |  |-- P. muralis
         |  |  |-- P. peloponnesiaca
         |  |  |-- P. pityusensis
         |  |  |-- P. raffonei
         |  |  |-- P. sicula
         |  |  |-- P. taurica
         |  |  |-- P. tiliguerta
         |  |  `-- P. wagleriana
         |  |-- Timon lepidus [sg. => Lacerta (Timon)?]
         |  |?- Darevskia [sg. => Lacerta (Darevskia)?]
         |  `==o Lacerta [paraphyletic]
         |     |-- (Apathya)
         |     |-- (Archaeolacerta)
         |     |-- (Omanosaura)
         |     |-- (Teira)
         |     `-- (Zootoca)
         `--+--+-- Acanthodactylus (ripsusisiliskot)
            |  `--o Mesalina
            |     |-- M. adramitana
            |     |-- M. ayunensis
            |     |-- M. balfouri
            |     |-- M. brevirostris
            |     |-- M. ercolinii
            |     |-- M. guttulata
            |     |-- M. martini
            |     |-- M. olivieri
            |     |-- M. pasteuri
            |     |-- M. rubropunctata
            |     |-- M. simoni
            |     `-- M. watsonana
            `--+--+--o Nucras
               |  |  |-- N. boulengeri
               |  |  |-- N. caesicaudata
               |  |  |-- N. intertexta
               |  |  |-- N. lalandii
               |  |  |-- N. scalaris
               |  |  |-- N. taeniolata
               |  |  `-- N. tessellata
               |  `--+--o Latastia
               |     |  |-- L. boscai
               |     |  |-- L. carinata
               |     |  |-- L. cherchii
               |     |  |-- L. doriai
               |     |  |-- L. johnstonii
               |     |  |-- L. lanzai
               |     |  |-- L. longicaudata
               |     |  |-- L. ornata
               |     |  |-- L. siebenrocki
               |     |  `-- L. taylori
               |     `--o Heliobolus
               |        |-- H. lugubris
               |        |-- H. neumanni
               |        |-- H. nitida
               |        `-- H. spekii
               `--+--+--o Adolfus
                  |  |  |-- A. africanus
                  |  |  |-- A. alleni
                  |  |  |-- A. jacksoni
                  |  |  `-- A. vauereselli
                  |  |--o Eremias
                  |  |  |-- (Ommateremias)
                  |  |  |-- (Pareremias)
                  |  |  |-- (Rhabderemias)
                  |  |  `-- (Scapteira)
                  |  `--o Ophisops
                  |     |-- O. beddomei
                  |     |-- O. elbaensis
                  |     |-- O. elegans
                  |     |-- O. jerdonii
                  |     |-- O. leschenaultii
                  |     |-- O. microlepis
                  |     |-- O. minor
                  |     `-- O. occidentalis
                  `--+--o Meroles
                     |  |-- M. anchietae
                     |  |-- M. ctenodactylus
                     |  |-- M. cuneirostris
                     |  |-- M. knoxii
                     |  |-- M. micropholidotus
                     |  |-- M. reticulatus
                     |  `-- M. suborbitalis
                     `--+--o Pedioplanis
                        |  |-- P. benguelensis
                        |  |-- P. breviceps
                        |  |-- P. burchelli
                        |  |-- P. gaerdesi
                        |  |-- P. husabensis
                        |  |-- P. laticeps
                        |  |-- P. lineoocellata
                        |  |-- P. namaquensis
                        |  |-- P. rubens
                        |  `-- P. undata
                        `--o Tropidosaura
                           |-- T. cottrelli
                           |-- T. essexi
                           |-- T. gularis
                           `-- T. montana

Lacertids not treated by Fu, 2000:

    Australolacerta
        A. australis
        A. rupicola
    Eremiascincus
        E. fasciolatus
        E. richardsonii
    Gastropholis
        G. echinata
        G. prasina
        G. tropidopholis
        G. vittata
    Holaspis guentheri
    Ichnotropis
        I. bivittata
        I. capensis
        I. chapini
        I. grandiceps
        I. microlepidota
        I. squamulosa
        I. tanganicana
    Philochortus
        P. hardeggeri
        P. intermedius
        P. lhotei
        P. neumanni
        P. phillipsi
        P. spinalis
        P. zolii
    Poromera fordii
    Pseuderemias
        P. brenneri
        P. erythrosticta
        P. mucronata
        P. savagei
        P. septemstriata
        P. smithii
        P. striatus

<==o Lacertidae
   |--o Gallotiinae
   |  |-- Gallotia
   |  `-- Psammodromus
   `--o Lacertinae [23/260]
      |-- Acanthodactylus (ripsusisiliskot) [36]
      |-- Adolfus
      |-- Algyroides
      |-- Australolacerta
      |-- Eremias [27]
      |-- Eremiascincus
      |-- Gastropholis
      |-- Holaspis
      |-- Heliobolus
      |-- Ichnotropis
      |-- Lacerta [62]
      |-- Latastia
      |-- Meroles
      |-- Mesalina
      |-- Nucras
      |-- Ophisops
      |-- Pedioplanis
      |-- Philochortus
      |-- Podarcis
      |-- Poromera
      |-- Pseuderemias
      |-- Takydromus Daudin, 1802 [Tachysaurus Gray, 1845; Platyplacopus Boulenger, 1917; Apeltonotus Boulenger, 1917] (itä-aasiansisiliskot; East Asian grass lizards)
      |-- Timon [sg. => Lacerta?]
      `-- Tropidosaura

Reference(s):

• Arnold, E. N., 1997: Interrelationships and evolution of the east Asian grass lizards, Takydromus (Squamata: Lacertidae).
  –Zoological Journal of the Linnean Society: Vol. 119, pp. 267-296

• Augé, M., 2003: La faune de Lacertilia (Reptilia, Squamata) de l'Éocène inférieur de Prémontré (Bassin de Paris, France)
  [The lizard fauna (Reptilia, Squamata) from the lower Eocene locality of Prémontré (Paris Basin, France)].
  –Geodiversitas: Vol. 25, #3, pp. 539-574
• Augé, M., Bailon, S. & Malfay, J.-P., 2003: Un nouveau genre de Lacertidae (Reptilia, Lacertilia) dans les faluns miocènes de l'Anjou-Touraine (Maine-et-Loire, France).
  [A new genus of Lacertidae (Reptilia, Lacertilia) from the Miocene deposits of Anjou-Touraine (Maine-et-Loire, France)].
  –Geodiversitas: Vol. 25, #2, pp. 289-295
• Borsuk-Bialynicka, M., Lubka, M., & Böhme, W. 1999: A lizard from Baltic amber (Eocene) and the ancestry of the crown group lacertids.
  – Acta Palaeontologica Polonica: Vol. 44, #4, pp. 349-382.
  [from iNet Abstract: http://www.paleo.pan.pl/acta/acta44-4.htm#Borsuk]
• Carroll, R. L., 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698
• Carroll, R. L., 1988: Appendix. 594-648.
  in Carrol, R. L. 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698
• Fu, J., 2000: Toward the phylogeny of the family Lacertidae - Why 4708 base pairs of mtDNA sequences cannot draw the picture.
  –Biological Journal of the Linnean Society: Vol. 71, #2, pp. 203-217
Murphy, R. W., Fu, J., MacCulloch, R. D., Darevsky, I. S. & Kupriyanova, L. A., 2000: A fine line between sex and unisexuality: the phylogenetic constraints on parthenogenesis in lacertid lizards.
–Zoological Journal of the Linnean Society: Vol. 130, pp. 527-549
• Rieppel, O. 1994: The Lepidosauromorpha: an overview with special emphasis on the Squamata. 23-37.
  in Fraser, N. C. & Sues, H-D. 1994: In the Shadow of the Dinosaurs.
  –Cambridge University Press, New York, 1994
• Uetz, P., Etzold, T. & Chenna, R., 1995-2000: The Reptile Database.
  –iNet: http://www.reptile-database.org/