After Uetz, Etzold & Chenna, 1995-2000 [“The Reptile Database” (former EMBL Reptile Database)]
<==o Scincidae Gray, 1825 [incl. Acontidae, Mabuyidae, Egerniidae, Eugongylidae, Lygosomidae, and Sphenomorphidae] (scincs; skinkit) |?- †Axonoscincus sabatieri Augé, 2005; L.Eoc. WEu. |?- †Contogenys sloani |?- †Estescincosaurus cooki (Estes, 1964) [Sauriscus Estes, 1964 non Sauriscus Lawrence, 1949, a tick (Arachnida: Trombiculidae)] L.Pal. NA. |?- †Palaeoscincosaurus middletoni Sullivan & Lucas, 1996 |--o Ablepharus | |-- A. bivittatus Menetries, 1832 | |-- A. chernovi Darevsky, 1953 | |-- A. darvazi Jeriomtshenko & Panfilov, 1990 | |-- A. deserti Strauch, 1868 | |-- A. grayanus Stoliczka, 1872 | |-- A. kitaibelii Bibron & Bory, 1833 | `-- A. pannonicus Fitzinger, 1823 |-- Acontophiops lineatus Sternfeld, 1919 |--o Afroblepharus | |-- A. duruarus Monard, 1949 | |-- A. seydeli de Witte, 1933 | |-- A. tancredi Boulenger, 1909 | `-- A. wilsoni Werner, 1919 |--o Asymblepharus | |-- A. alaicus Elpatjevsky, 1901 | |-- A. mahabharatus Eremchenko, Shah & Panfilov, 1998 | |-- A. nepalensis Eremchenko & Helfenberger, 1998 | `-- A. tragbulense Alcock, 1898 |-- Bartleia jigurru Covacevich, 1984 |--o Caledoniscus | |-- C. aquilonius Sadlier, Bauer & Colgan, 1999 | |-- C. atropunctatus Roux, 1913 | |-- C. auratus Sadlier, Bauer & Colgan, 1999 | |-- C. austrocaledonicus Bavay, 1869 | |-- C. chazeaui Sadlier, Bauer & Colgan, 1999 | |-- C. cryptos Sadlier, Bauer & Colgan, 1999 | |-- C. festivus Roux, 1913 | |-- C. haplorhinus Günther, 1872 | |-- C. orestes Sadlier, 1987 | |-- C. renevieri Sadlier, Bauer & Colgan, 1999 | `-- C. terma Sadlier, Bauer & Colgan, 1999 |-- Cautula zia Ingram & Ehmann, 1981 |-- Chalcidoseps thwaitesi Günther, 1872 |-- Coggeria naufragus Couper et al., 1996 |--o Cophoscincopus | |-- C. durus Cope, 1862 | |-- C. greeri Böhme, Schmitz & Ziegler, 2000 | `-- C. simulans Vaillant, 1884 |-- Cryptoscincus minimus Mocquard, 1906 |--o Cyclodina | |-- C. aenea Girard, 1857 | |-- C. alani Robb, 1970 | |-- C. lichenigera O'Shaughnessy, 1874 | |-- C. macgregori Robb, 1975 | |-- C. oliveri McCann, 1955 | |-- C. ornatus Gray, 1843 | `-- C. whitakeri Hardy, 1977 |-- Davewakeum miriamae Heyer, 1972 |-- Eroticoscincus graciloides Lönnberg & Andersson, 1913 |--o Eumecia | |-- E. anchietae Bocage, 1870 | `-- E. johnstoni Boulenger, 1897 |--o Euprepes | |-- E. chaperi Vaillant, 1884 | `-- E. vittatus Peters, 1862 |-- Evesia bellii Dúmeril & Bibron, 1839 |-- Fojia bumui Greer & Simon, 1982 |--o Geomyersia | |-- G. coggeri Greer, 1982 | `-- G. glabra Greer & Parker, 1968 |-- Geoscincus haraldmeieri Böhme, 1976 |--o Gongylus | |-- G. androvandii Dúmeril & Bibron 1839 | |-- G. brachypoda Dúmeril & Bibron 1839 | `-- G. crassicaudum Dúmeril & Dúmeril 1851 |-- Graciliscincus shonae Sadlier 1987 |?- Grandidierina |-- Haackgreerius miopus Greer & Haacke 1982 |-- Hemisphaeriodon gerrardii Gray, 1845 |--o Isopachys | |-- I. anguinoides Boulenger 1914 | |-- I. borealis Lang & Böhme 1990 | |-- I. gyldenstolpei Lönnberg 1916 | `-- I. roulei Angel 1920 |?- Lacertaspis |-- Lacertoides pardalis Sadlier, Shea & Bauer, 1997 |?- Lacertus |--o Lankascincus | |-- L. deignani Taylor, 1950 | |-- L. deraniyagalae Greer, 1991 | |-- L. fallax Peters, 1860 | |-- L. gansi Greer, 1991 | |-- L. taprobanensis Kelaart, 1854 | `-- L. taylori Greer, 1991 |--o Larutia | |-- L. larutense Boulenger, 1900 | |-- L. miodactyla Boulenger, 1903 | |-- L. sumatrensis Bleeker, 1860 | `-- L. trifasciata Tweedie, 1940 |--o Leptoseps | |-- L. osellai Böhme, 1981 | `-- L. poilani Bourret, 1937 |--o Leptosiaphos | |-- L. aloysiiabaudiae Peracca, 1907 | |-- L. amieti Perret, 1973 | |-- L. blochmanni Tornier, 1903 | |-- L. dewittei Loveridge, 1934 | |-- L. fuhni Perret, 1973 | |-- L. gemmiventris Sjöstedt, 1897 | |-- L. graueri Sternfeld, 1912 | |-- L. hackarsi Witte, 1941 | |-- L. hylophilus Laurent, 1982 | |-- L. ianthinoxantha Böhme, 1975 | |-- L. kilimensis Stejneger, 1891 | |-- L. lepesmei Angel, 1940 | |-- L. luberoensis de Witte, 1933 | |-- L. meleagris Boulenger, 1907 | |-- L. pauliani Angel, 1940 | |-- L. rhodurus Laurent, 1952 | |-- L. rhomboidalis Broadley, 1989 | `-- L. vigintiserierum Sjöstedt, 1897 |--o Lioscincus | |-- L. greeri Böhme, 1979 | |-- L. maruia Sadlier, Whitaker & Bauer, 1998 | |-- L. nigrofasciolatum Peters, 1869 | |-- L. novaecaledoniae Parker, 1926 | |-- L. steindachneri Bocage, 1873 | `-- L. tillieri Ineich & Sadlier, 1991 |--o Lobulia | |-- L. brongersmai Zweifel, 1972 | `-- L. elegans Boulenger, 1897 |?- Lubuya |--o Lygisaurus | |-- L. aeratus Garman, 1901 | |-- L. foliorum de Vis, 1884 | |-- L. laevis Oudemans, 1894 | |-- L. macfarlani Günther, 1877 | |-- L. novaeguineae Meyer, 1874 | |-- L. rococo Ingram & Covacevich, 1988 | |-- L. sesbrauna Ingram & Covacevich, 1988 | |-- L. tanneri Ingram & Covacevich, 1988 | `-- L. zuma Couper, 1993 |-- Macroscincus coctei Dúmeril & Bibron, 1839 |--o Marmorosphax | |-- M. euryotis Werner, 1909 | `-- M. tricolor Bavay, 1869 |?- Malacontias |--o Menetia | |-- M. alanae Rankin, 1979 | |-- M. amaura Storr, 1978 | |-- M. concinna Sadlier, 1984 | |-- M. greyii Gray, 1845 | |-- M. koshlandae Greer, 1991 | |-- M. maini Storr, 1976 | |-- M. sadlieri Greer, 1991 | |-- M. surda Storr, 1976 | `-- M. timlowi Ingram, 1977 |--o Mochlus | |-- M. brevicaudis Greer et al., 1985 | |-- M. fernandi Burton, 1836 | |-- M. guineensis Peters, 1879 | `-- M. sundevalli Smith, 1849 |-- Morethia | |-- M. adelaidensis Peters, 1874 | |-- M. boulengeri Ogilby, 1890 | |-- M. butleri Storr, 1963 | |-- M. lineoocellata Dúmeril & Bibron, 1839 | |-- M. obscura Storr, 1972 | |-- M. ruficauda Lucas & Frost, 1895 | |-- M. storri Greer, 1980 | `-- M. taeniopleura Peters, 1874 |--o Nannoscincus | |-- N. gracilis Bavay, 1869 | |-- N. greeri Sadlier, 1987 | |-- N. maccoyi Lucas & Frost, 1894 | |-- N. mariei Bavay, 1869 | |-- N. rankini Sadlier, 1987 | `-- N. slevini Loveridge, 1941 |-- Neoseps reynoldsi Stejneger, 1910 |--o Novoeumeces | |-- N. algeriensis Peters, 1864 | |-- N. blythianus Anderson, 1871 | `-- N. schneideri Daudin, 1802 |--o Panaspis | |-- P. africana Gray, 1845 | |-- P. annobonensis Fuhn, 1972 | |-- P. breviceps Peters, 1873 | |-- P. burgeoni Witte | |-- P. cabindae Bocage, 1866 | |-- P. chriswildi Böhme & Schmitz, 1996 | |-- P. helleri Loveridge | |-- P. kitsoni Boulenger, 1913 | |-- P. maculicollis Jacobson & Broadley 2000 | |-- P. megalurus Nieden, 1913 | |-- P. nimbaensis Angel, 1944 | |-- P. quattuordigitata Sternfeld, 1912 | |-- P. reichenowi Peters, 1874 | |-- P. rohdei Müller, 1910 | |-- P. thomasi Tornier | |-- P. togoensis Werner, 1902 | `-- P. wahlbergi Smith 1849 |--o Papuascincus | |-- P. buergersi Vogt, 1932 | |-- P. morokanus Parker, 1936 | |-- P. phaeodes Vogt, 1932 | `-- P. stanleyanus Boulenger, 1897 |-- Parachalcides socotranus Boulenger, 1899 |-- Paralipinia rara Darevsky & Orlov, 1997 |--o Parvoscincus | |-- P. palawanensis Brown & Alcala, 1961 | `-- P. sisoni Ferner, Brown & Greer, 1997 |--o Phoboscincus | |-- P. bocourti Brocchi, 1876 | `-- P. garnieri Bavay 1869 |--o Proablepharus | |-- P. kinghorni Copland, 1947 | |-- P. reginae Glauert, 1960 | `-- P. tenuis Broom, 1896 |--o Pseudemoia | |-- P. baudini Dúmeril & Bibron, 1839 | |-- P. cryodroma Hutchinson & Donnellan, 1992 | |-- P. entrecasteauxii Dúmeril & Bibron, 1839 | |-- P. pagenstecheri Lindholm, 1901 | |-- P. rawlinsoni Hutchinson & Donnellan, 1988 | `-- P. spenceri Lucas & Frost, 1894 |--o Riopa | |-- R. bampfyldei Bartlett, 1895 | |-- R. herberti Smith, 1916 | `-- R. opisthorhodum Werner, 1910 |--o Ristella | |-- R. beddomii Boulenger 1887 | |-- R. guentheri Boulenger 1887 | |-- R. rurkii Gray 1839 | `-- R. travancorica Beddome 1870 |--o Saproscincus | |-- S. basiliscus Ingram & Rawlinson, 1981 | |-- S. challengeri Boulenger, 1887 | |-- S. czechurai Ingram & Rawlinson, 1981 | |-- S. hannahae Couper & Keim, 1998 | |-- S. lewisi Couper & Keim, 1998 | |-- S. mustelinus O'Shaughnessy, 1874 | |-- S. rosei Wells & Wellington, 1985 | |-- S. spectabilis de Vis, 1888 | `-- S. tetradactylus Greer & Kluge, 1980 |--o Sigaloseps | |-- S. deplanchei Bavay, 1869 | `-- S. ruficauda Sadlier & Bauer, 1999 |-- Simiscincus aurantiacus Sadlier & Bauer, 1997 |-- Tachygia microlepis Dúmeril & Bibron, 1839 |-- Trachydosaurus rugosus Gray, 1825 |-- Tropidophorus [20] |--o Tropidoscincus | |-- T. aubrianus Bocage 1873 | |-- T. rohsii Andersson 1908 | `-- T. variabilis Bavay 1869 |-- Vietnascincus rugosus Darevsky & Orlov, 1994 |--o Acontinae | |--o Acontias [incl. Typhlosaurus in partim; Microacontias] | | |-- A. aurantiacus (Peters, 1854) | | |-- A. bicolor (Hewitt, 1929) | | |-- A. breviceps Essex, 1925 | | |-- A. cregoi (Boulenger, 1903) | | |-- A. gariepensis (FitzSimons, 1941) | | |-- A. gracilicauda Essex, 1925 | | |-- A. jappi (Broadley, 1968) | | |-- A. kgalagadi (Lamb, Biswas & Bauer, 2010) | | |-- A. lineatus Peters, 1879 | | |-- A. litoralis Broadley & Greer, 1969 | | |-- A. meleagris Linnaeus, 1758 | | |-- A. namaquensis Hewitt, 1938 | | |-- A. orientalis Hewitt, 1938 | | |-- A. percivali Loveridge, 1935 | | |-- A. plumbeus Bianconi, 1849 | | |-- A. poecilus Bourquin & Lambiris, 1996 | | |-- A. richardi (Jacobsen, 1987) | | |-- A. rieppeli (Lamb, Biswas & Bauer, 2010) | | `-- A. tristis Werner, 1911 | `--o Typhlosaurus | |-- T. braini Haacke, 1964 | |-- T. caecus Cuvier, 1817 | |?- T. lineatus Boulenger, 1887 | |-- T. lomiae Haacke, 1986 [T. lomii] | |-- T. meyeri Boettger, 1894 | `-- T. vermis Boulenger, 1887 |-- Scincinae `--o Lygosominae Mittleman, 1952 |--o Sphenomorphus-group | |?- Lipinia | |?-o Tribolonotus (Spinose Crocodile Skinks) | | |-- T. annectens Zweifel, 1966 | | |-- T. blanchardi Burt, 1930 | | |-- T. brongersmai Cogger, 1973 | | |-- T. gracilis de Rooij, 1909 | | |-- T. novaeguineae Schlegel, 1834 | | |-- T. ponceleti Kinghorn, 1937 | | |-- T. pseudoponceleti Greer & Parker, 1968 | | `-- T. schmidti Burt, 1930 | |--+-- Scincella | | `--o Prasinohaema | | |-- P. flavipes Parker, 1936 | | |-- P. parkeri Smith, 1937 | | |-- P. prehensicauda Loveridge, 1945 | | |-- P. semoni Oudemans, 1894 | | `-- P. virens Peters, 1881 | `--+-- Sphenomorphus | `--+--o Notoscincus | | |-- N. butleri Storr, 1979 | | `-- N. ornatus Broom, 1896 | `--+--+--o Anomalopus | | | |-- A. brevicollis Greer & Cogger, 1985 | | | |-- A. gowi Greer & Cogger, 1985 | | | |-- A. leuckartii Weinland, 1862 | | | |-- A. mackayi Greer & Cogger, 1985 | | | |-- A. pluto Ingram, 1977 | | | |-- A. swansoni Greer & Cogger, 1985 | | | `-- A. verreauxi Dúmeril & Dúmeril, 1851 | | `--+--o Calyptotis | | | |-- C. lepidorostrum Greer, 1983 | | | |-- C. ruficauda Greer, 1983 | | | |-- C. scutirostrum Peters, 1874 | | | |-- C. temporalis Greer, 1983 | | | `-- C. thorntonensis Greer, 1983 | | `--+--+-- Nangura spinosa Covacevich, Couper & James, 1993 | | | `--+-- Gnypetoscincus queenslandiae de Vis, 1890 | | | `==o Eulamprus | | | |-- E. amplus Covacevich & McDonald, 1980 | | | |-- E. brachyosoma Lönnberg & Andersson, 1915 | | | |-- E. frerei Greer, 1992 | | | |-- E. heatwolei Wells & Wellington, 1983 | | | |-- E. kosciuskoi Kinghorn, 1932 | | | |-- E. leuraensis Wells & Wellington, 1983 | | | |-- E. luteilateralis Covacevich & McDonald, 1980 | | | |-- E. martini Wells & Wellington, 1985 | | | |-- E. sokosoma Greer, 1992 | | | |-- E. tenuis Gray, 1831 | | | |-- E. tigrinus de Vis, 1888 | | | |-- E. tryoni Longman, 1918 | | | `-- E. tympanum Lönnberg & Andersson, 1915 | | `--+-- Eulamprus murrayi Boulenger, 1887 | | `--+--o Ophioscincus | | | |-- O. cooloolensis Greer & Cogger, 1985 | | | |-- O. ophioscincus Boulenger, 1887 | | | `-- O. truncatus Peters, 1876 | | `--+-- Saiphos equalis Gray 1825 | | `--o Coeranoscincus | | |-- C. frontalis de Vis, 1888 | | `-- C. reticulatus Günther, 1873 | `--+--+--o Eremiascincus Greer, 1979 sensu Mecke, Doughty & Donnellan, 2009 | | | |-- E. antoniorum (Smith, 1927) (Antoni) | | | |-- E. brongersmai (Storr, 1972) (Brown-sided Bar-lipped Skink) | | | |-- E. butlerorum (Aplin, How & Boeadi, 1993) | | | |-- E. douglasi (Storr, 1967) (Orange-sided Bar-lipped Skink) | | | |-- E. emigrans (Lidth de Jeude, 1895) | | | |-- E. fasciolatus (Günther, 1867) (Narrow-banded Sand-swimmer) | | | |-- E. isolepis (Boulenger, 1887) (Northern Bar-lipped Skink) | | | |-- E. musivus Mecke, Doughty & Donnellan, 2009 (Mosaic Desert Skink) | | | |-- E. pardalis (MacLeay, 1877) (Lowlands Bar-lipped Skink) | | | |-- E. richardsonii (Gray, 1845) (Broad-banded Sand-swimmer) | | | `-- E. timorensis (Greer, 1990) | | `--+==o Glaphyromorphus Wells & Wellington, 1983 | | | |-- G. cracens Greer, 1985 | | | |-- G. darwiniensis Storr, 1967 | | | |-- G. fuscicaudis Greer, 1979 | | | |-- G. gracilipes Steindachner, 1870 | | | |-- G. mjobergi Lönnberg & Andersson, 1915 | | | |-- G. nigricaudis Macleay, 1877 | | | |-- G. pumilus Boulenger, 1887 | | | `-- G. punctulatus Peters, 1871 | | `--o Hemiergis | | |-- H. decresiensis Cuvier 1829 | | |-- H. initialis Werner 1910 | | |-- H. millewae Coventry 1976 | | |-- H. peronii Fitzinger 1826 | | `-- H. quadrilineatum Dúmeril & Bibron 1839 | `--+--+-- Glaphyromorphus arnhemicus Storr, 1967 [Glaphyromorphus darwiniensis arnhemicus Storr, 1967] | | `-- em>Eulamprus quoyii Quoy & Gaimard, 1824 | `--+-- Lerista | `-- Ctenotus `--+--o Egernia-group | |-- Corucia zebrata Gray, 1855 (jättiläiskinkki) | |--o Cyclodomorphus | | |-- C. branchialis Günther, 1867 | | |-- C. casuarinae Dúmeril & Bibron, 1839 | | |-- C. celatus Shea & Miller, 1995 | | |-- C. maxima Storr, 1976 | | |-- C. melanops Stirling & Zietz, 1893 | | |-- C. michaeli Wells & Wellington, 1984 | | |-- C. praealtus Shea, 1995 | | `-- C. venustus Shea & Miller, 1995 | |-- Egernia (piikkiskinkit) | `--o Tiliqua (sinikieliskinkit) | |-- T. adelaidensis Peters, 1863 | |-- T. gigas Schneider, 1801 | |-- T. multifasciata Sternfeld, 1919 | |-- T. nigrolutea Quoy & Gaimard, 1824 | |-- T. occipitalis Peters, 1863 | `-- T. scincoides White, 1790 `--+--o Lygosoma-group | |--o Lamprolepis | | |-- L. leucosticta Müller, 1923 | | |-- L. nieuwenhuisi Lidth de Jeude, 1905 | | |-- L. smaragdina Lesson, 1826 | | `-- L. vyneri Shelfford, 1905 | `-- Lygosoma (hoikkaskinkit) `--+--o Eugongylus-group | |--o Bassiana | | |-- B. duperreyi Gray, 1838 | | |-- B. platynota Peters, 1881 | | `-- B. trilineata Gray, 1838 | |-- Carlia | |-- Cryptoblepharus | |-- Emoia | |--o Eugongylus | | |-- E. albofasciolatus Günther, 1872 | | |-- E. microlepis Dúmeril & Bibron, 1851 | | |-- E. rufescens Shaw, 1802 | | |-- E. sulaensis Kopstein, 1927 | | `-- E. unilineatus de Rooij, 1915 | |--o Lampropholis | | |-- L. adonis Ingram, 1991 | | |-- L. amicula Ingram & Rawlinson, 1981 | | |-- L. caligula Ingram & Rawlinson, 1981 | | |-- L. coggeri Ingram, 1991 | | |-- L. colossus Ingram, 1991 | | |-- L. couperi Ingram, 1991 | | |-- L. delicata de Vis, 1888 | | |-- L. elongata Greer, 1997 | | |-- L. guichenoti Dúmeril & Bibron, 1839 | | |-- L. mirabilis Ingram & Rawlinson, 1981 | | `-- L. robertsi Ingram, 1991 | |--o Leiolopisma | | |-- L. alazon Zug, 1985 | | |-- L. bardensis | | |-- L. eulepis Fitzinger, 1853 | | |-- L. fasciolare Girard, 1858 | | |-- L. lioscincus Bocage, 1857 | | |-- L. mauritiana Günther, 1877 | | |-- L. paronae | | |-- L. rhomboidalis Peters | | `-- L. telfairii des Jardin, 1831 | |--o Niveoscincus | | |-- N. coventryi Rawlinson, 1975 | | |-- N. greeni Rawlinson, 1975 | | |-- N. metallicus Gray, 1845 | | |-- N. microlepidotus O'Shaughnessy, 1874 | | |-- N. ocellatus Gray, 1845 | | |-- N. orocryptus Hutchinson et al., 1988 | | |-- N. palfreymani Rawlinson, 1974 | | `-- N. pretiosus O'Shaughnessy, 1874 | `-- Oligosoma `--o Mabuya-group |-- Apterygodon vittatum Edeling, 1865 |?-o Chioninia [Mabuya in partim] (Cape Verde Mabuyas) | |-- C. delalandii (Duméril & Bibron, 1839) | |-- C. fogoensis (O'Shaughnessy, 1874) | |-- C. geisthardti Joger, 1993 | |-- C. spinalis (Boulenger, 1906) | |-- C. stangeri (Gray, 1845) | `-- C. vaillantii (Boulenger, 1885) (Vaillant's Mabuya) |--o Dasia | |-- D. griffini Taylor, 1915 | |-- D. grisea Gray, 1845 | |-- D. haliana Haly & Nevill, 1887 | |-- D. nicobarensis Biswas & Sanyal, 1977 | |-- D. nieuwenhuisi Lidth de Jeude | |-- D. olivacea Gray, 1839 | |-- D. semicincta Peters, 1867 | `-- D. subcaerulea Boulenger, 1891 `-- Mabuya (mabuijat)
Reference(s):
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–Zootaxa: Vol. #2246, pp. 1-20 - Reeder, T. W., 2003: A phylogeny of the Australian Sphenomorphus group (Scincidae: Squamata) and the phylogenetic placement of the crocodile skinks (Tribolonotus): Bayesian approaches to assessing congruence and obtaining confidence in maximum likelihood inferred relationships.
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–Cambridge University Press, New York, 1994 - Sullivan, R. M., 1997: Estescincosaurus cooki (Estes, 1964) new genus (Squamata: Scincomorpha): Replacement name for Sauriscus Lawrence, 1949 (Arachnida: Trombiculidae).
–Journal of Vertebrate Paleontology: Vol. 17, #2, pp. 447 - Sullivan, R. M. & Lucas, S. G., 1996: Palaeoscincosaurus middletoni, new genus and species (Squamata: ?Scincidae) from the Early Paleocene (Puercan) Denver Formation, Colorado.
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–iNet: http://www.reptile-database.org/