After Motani, 1998, 1999, Nicholls & Manabe, 2001, Chen, Motani, Cheng, Jiang & Rieppel, 2014b, and Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014
Family-level taxonomy of ichthyopterygians
<==o †Ichthyosauromorpha Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014 (ichthyosaurs and relatives; kalaliskomaiset matelijat) |?- †Wumengosaurus delictomandibularis Jiang, Rieppel, Motani, Hao, W-C. Sun, Schmitz, & Z-Y. Sun, 2008 [originally treated as nothosauroid sauropterygian] |--o †Hupehsuchia Carroll & Dong, 1991 | |-- †Nanchungosaurus suni Wang, 1959; L.Trias. SEAs. [Nanchangosauridae Wang, 1959] | `--+-- †Eohupehsuchus brevicollis Chen, Motani, Cheng, Jiang & Rieppel, 2014c | `--o †Hupehsuchidae Young, 1972 | |-- †Hupehsuchus nanchengensis Young, 1972; L.Trias. SEAs. | `--o †Parahupehsuchinae Chen, Motani, Cheng, Jiang & Rieppel, 2015 | |-- †Eretmorhipis carrolldongi Chen, Motani, Cheng, Jiang & Rieppel, 2015 [IVPP.V4070; WGSC.V26020] | `-- †Parahupehsuchus longus Chen, Motani, Cheng, Jiang & Rieppel, 2014a; L.Trias. SEAs. `--o †Ichthyosauriformes Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014 |--o †Nasorostra Jiang, Motani, Huang, Tintori, Hu, Rieppel, Fraser, Ji, Kelley, Fu & Zhang, 2016 | |-- †Cartorhynchus lenticarpus Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014 | `-- †Sclerocormus parviceps Jiang, Motani, Huang, Tintori, Hu, Rieppel, Fraser, Ji, Kelley, Fu & Zhang, 2016 `--o †Ichthyopterygia Owen, 1840 |?- †Thaisaurus chonglakmanii Mazin, Suteethorn, Buffenaut, Jaeger & Helmcke-Ingavat, 1991 |?- †Isfjordosaurus minor Motani, 1999 [Pessopteryx] |-- †Utatsusaurus hataii Shikama, Kamei & Murata, 1978 |-- †Parvinatator wapitiensis Nicholls & Brickman, 1995 `--o Eoichthyosauria Motani, 1999b [Ichthyosauria sensu lato] |--o †Grippidae Wiman, 1929 [Grippidia Wiman, 1929] | |?- †Gulosaurus helmiCuthbertson, Russell & Anderson, 2013 | |-- †Grippia longirostris Wiman, 1929 | `--o †Chaohusaurus Young & Dong, 1972 [Anhuisaurus Chen, 1985; Chensaurus (Chen, 1985) Mazin et al., 1991] | |-- †C. geishanensis Young & Dong, 1972 [Anhuisaurus faciles Chen, 1985; Chensaurus faciles (Chen, 1985) Mazin et al., 1991] | |?- †C. chaoxianensis (Chen, 1985) [Anhuisaurus chaoxianensis Chen, 1985; Chensaurus chaoxianensis (Chen, 1985) Mazin et al., 1991] | `-- †C. zhangjiawanensis Chen, Sander, Long & Wang, 2013 `-- †Ichthyosauria Blainville, 1835
Ichthyopterygia incertae sedis:
†Protoichthyosauridae Appleby, 1979 †Protoichthyosaurus prosostealis Appleby, 1979 Nomina dubia: †Svalbardosaurus crassidens Mazin, 1981 †Shastasaurus altispinus Merriam, 1902 †Shastasaurus carinthiacus Huene, 1925 †Shastasaurus careyi Merriam, 1902 †Pessopteryx arctica Wiman, 1910 †Pessopteryx nisseri Wiman, 1910 †Pessopteryx pinguis Wiman, 1910
| After Jiang, Motani, Huang, Tintori, Hu, Rieppel, Fraser, Ji, Kelley, Fu & Zhang, 2016<==o †Ichthyosauromorpha |-- †Hupehsuchia `--o †Ichthyosauriformes |--o †Nasorostra | |-- †Cartorhynchus | `-- †Sclerocormus `--o †Ichthyopterygia |-- †Chaohusaurus geishanensis |-- †Chaohusaurus chaoxianensis |-- †Chaohusaurus zhangjiawanensis `--+-- †Utatsusaurus |-- †Parvinatator |--o †Grippidia | |-- †Grippia | `-- †Gulosaurus `-- †Ichthyosauria 
 | After Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014 (C. Ji et al., in prep.)<==o †Ichthyosauromorpha |-- †Hupehsuchia `--o †Ichthyosauriformes |-- †Cartorhynchus `--o †Ichthyopterygia |--o †Chaohusaurus | |-- †C. geishanensis | |-- †C. chaoxianensis | `-- †C. zhangjiawanensis `--+--o †Grippidia | |-- †Utatsusaurus | |-- †Parvinatator | `--+-- †Grippia | `-- †Gulosaurus `-- †Ichthyosauria 
 | After Chen, Motani, Cheng, Jiang & Rieppel, 2014b<==o |-- †Thalattosauriformes `--+-- †Wumengosaurus `--+-- †Ichthyopterygia `--+-- †Nanchungosaurus `-- †Hupehsuchus 
 | 
Reference(s):
- Bardet, N. & Fernandez, M., 2000: A new ichthyosaur from the Upper Jurassic lithographic limestones of Bavaria.
 –Journal of Paleontology: Vol. 74, #3, pp. 503-511
- Chen, X.-h., Motani, R., Cheng, L., Jiang, D.-y. & Rieppel, O., 2015: A New Specimen of Carroll's Mystery Hupehsuchian from the Lower Triassic of China.
 –Plos One: Vol. 10, #5, pp. e0126024 (1-15) [doi: 10.1371/journal.pone.0126024]
- Chen, X.-h., Motani, R., Cheng, L., Jiang, D.-y. & Rieppel, O., 2014a: A Carapace-Like Bony ‘Body Tube’ in an Early Triassic Marine Reptile and the Onset of Marine Tetrapod Predation.
 –Plos One: Vol. 9, #4, pp. e94396 (1-8) [doi: 10.1371/journal.pone.0094396]
- Chen, X.-h., Motani, R., Cheng, L., Jiang, D.-y. & Rieppel, O., 2014b: The Enigmatic Marine Reptile Nanchangosaurus from the Lower Triassic of Hubei, China and the Phylogenetic Affinities of Hupehsuchia.
 –Plos One: Vol. 9, #7, pp. e102361 (1-13) [doi: 10.1371/journal.pone.0102361]
- Chen, X.-h., Motani, R., Cheng, L., Jiang, D.-y. & Rieppel, O., 2014c: A Small Short-Necked Hupehsuchian from the Lower Triassic of Hubei Province, China.
 –Plos One: Vol. 9, #12, pp. e115244 [doi: 10.1371/journal.pone.0115244]
- Chen, X., Sander, P. M., Long, C. & Wang, X., 2013: A new Triassic primitive ichthyosaur from Yuanan, South China.
 –Acta Geologica Sinica - English Edition: Vol. 87, #3, pp. 672-677 [doi: 10.1111/1755-6724.12078]
- Cuthbertson, R. S., Russell, A. P. & Anderson, J. S., 2013: Cranial morphology and relationships of a new grippidian (Ichthyopterygia) from the Vega-Phroso Siltstone Member (Lower Triassic) of British Columbia, Canada.
 –Journal of Vertebrate Paleontology: Vol. 33, #4, pp. 831-847 [doi: 10.1080/02724634.2013.755989]
- Fernández, M. S., 1999: a new ichthyosaur from the Los Molles Formation (Early Bajocian), Neuquen Basin, Argentina.
 –Journal of Paleontology: Vol. 73, #4, pp. 677-681
- Jiang, D.-Y., Motani, R., Huang, J.-D., Tintori, A., Hu, Y.-C., Rieppel, O., Fraser, N. C., Ji, C., Kelley, N. P., Fu, W.-L. & Zhang, R., 2016: A large aberrant stem ichthyosauriform indicating early rise and demise of ichthyosauromorphs in the wake of the end-Permian extinction.
 –Scientific Reports: Vol. 6, Art. nr. 26232 [doi: 10.1038/srep26232]
- Jiang, D.-Y., Rieppel, O., Motani, R., Hao, W.-C., Sun, Y.-L., Schmitz, L. & Sun, Z.-Y., 2008: A New Middle Triassic Eosauropterygian (Reptilia, Sauropterygia) from Southwestern China.
 –Journal of Vertebrate Paleontology: Vol. 28, #4, pp. 1055-1062 [doi: 10.1671/0272-4634-28.4.1055]
- McGowan, C., 1994: A new species of Shastasaurus (Reptilia: Ichthyosauria) from the Triassic of British Colombia: The most complete exemplar of the genus.
 –Journal of Vertebrate Paleontology: Vol. 14, #2, pp. 168-179
- McGowan, C., 1997: The taxonomic status of the Late Jurassic ichthyosaur Grendelius mordax: A preliminary report.
 –Journal of Vertebrate Paleontology: Vol. 17, #2, pp. 428-430
- McGowan, C. & Milner, A. C., 1999: A new Pliensbachian ichthyosaur from Dorset, England.
 –Palaeontology: Vol. 42, #5, pp. 761-768
- Motani, R., 1996: Redescription of the dental features of an Early Triassic ichthyosaur, Utatsusaurus hataii.
 –Journal of Vertebrate Paleontology: Vol. 16, #3, pp. 396-402
- Motani, R., 1997: Redescription of the dentition of Grippia longirostris (Ichthyosauria) with a comparison with Utatsusaurus hataii.
 –Journal of Vertebrate Paleontology: Vol. 17, #1, pp. 39-44
- Motani, R., 1999: The skull and taxonomy of Mixosaurus (Ichthyopterygia).
 –Journal of Paleontology: Vol. 73, #5, pp. 924-935
- Motani, R., 1999: On the evolution and homologies of ichthyopterygian forefins.
 –Journal Of Vertebrate Paleontology: Vol. 19, #1, pp. 28-41.
- Motani, R., 1999: Phylogeny of the Ichthyopterygia.
 –Journal Of Vertebrate Paleontology: Vol. 19, #3, pp. 473-496.
- Motani, R., 2000: Skull of Grippia longirostris: No contradiction with a diapsid affinity for the Ichthyosauria.
 –Palaeontology: Vol. 43, #1, pp. 1-14
- Motani, R., 2000: Is Omphalosaurus ichthyopterygian? - A phylogenetic perspective.
 –Journal Of Vertebrate Paleontology: Vol. 20, #2, pp. 295-301
- Motani, R., Jiang, D.-Y., Chen, G.-B., Tintori, A., Rieppel, O., Ji, C. & Huang, J.-D., 2014: A basal ichthyosauriform with a short snout from the Lower Triassic of China.
 –Nature: in press [doi: 10.1038/nature13866]
- Motani, R., Jiang, D.-Y., Tintori, A., Rieppel, O., Chen, G.-B. & You, H., 2015: Status of Chaohusaurus chaoxianensis (Chen, 1985).
 –Journal of Vertebrate Paleontology: Vol. 35, #1, pp. e892011 (1-8) [doi: 10.1080/02724634.2014.892011]
- Motani, R., Minoura, N. & Ando, T., 1998: Ichthyosaurian relationships illuminated by new primitive skeletons from Japan.
 –Nature: Vol. 393, 21 May. pp. 255-257
- Motani, R. & You, H., 1998: Taxonomy and limb ontogeny of Chaohusaurus geishanensis (Ichthyosauria), with a note on the allometric equation.
 –Journal of Vertebrate Paleontology: Vol. 18, #3, pp. 533-540
- Motani, R., You, H., & McGowan, C., 1996: Eel-like swimming in the earliest ichthyosaurs.
 –Nature: Vol. 382, 25 July, pp. 347-348
- Nicholls, E. L. & Manabe, M., 2001: A new genus of ichthyosaur from the Late Triassic Pardonet Formation of British Colombia: bridging the Triassic Jurassic gap.
 –Canadian Journal of Earth Sciencies: Vol. 38, #6, pp. 983-1002
- Sander, P. M., 1989: The large ichthyosaur Cymbospondylus buchseri, sp. nov., from the Middle Triassic of Monte Giorgio (Switzerland), with a survay of the genus in Europe.
 –Journal of Vertebrate Paleontology: Vol. 9, #2, pp. 163-173
- Storrs, G. W., Arkhangel'skii, M. S. & Efimov, V. R., 2000: Mezosoic marine reptiles of Russia and other former Soviet republics. 187-210
 in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
 –Cambridge University Press, Cambridge, 2000 xxxix-696
 
