Alternative phylogeny of Basal Aves after Hou et al., 1996, Hou et al., 1999 & Feduccia, 1996 <==o AVIALAE sensu Gauthier 1986 |?- †Avialae genera et species indet. [Mongolia] |?- †Unenlagia comahuensis Novas & Puerta, 1997 |?- †Bambiraptor feinbergorum Burnham, Derstler, Currie, Bakker, Zhou & Ostrom, 2000 emended Olshevsky, 2000 `--o AVES (linnut, birds) |?- †Wyleyia valdensis Harrison & Walker, 1973 [nomen dubium] |--o †Archaeopterygidae [ARCHAEOPTERYGIFORMES] | |?- †Rahonavis ostromi (Forster et al., 1998a) [Rahona ostromi Forster et al., 1998a] | `-- †Archaeopteryx lithographica Meyer, 1861 [Synonyms] (liskolintu; urvogel) `--+?- †Protoavis texensis Chatterjee, 1991 sensu Chatterjee, 1997 `--o METORNITHES Chiappe, 1995 |?- †Alvarezsauridae sensu Chiappe, 1995, 1998 `--o PYGOSTYLIA (pygostyliset linnut) |--o †Confuciusornithidae Hou et al., 1995 | |-- †Changchengornis hengdaoziensis Ji et al., 1998 | `-- †Confuciusornis sanctus Hou et al., 1995 [Synonyms] `--o ORNITHOTHORACES (modern rib-cased birds) |-- †ENANTIORNITHES Walker, 1981 [Enantiornithomorpha Chiappe et al. 1999] ("opposite birds") `-- ORNITHUROMORPHA Chiappe et al. 1999 [Ornithurae sensu Feduccia 1996] (modernlike birds)
Note: The phylogeny of basal Avialae is murky. The cause of this murkiness are alvarezsaurids, which are currently of disputed phylogenetic position. I have here decided to follow the view of Chiappe, 1995, 1998. However a recent analysis of Paul C. Sereno places alvarezsaurids as very specialized ornithomimids. If this view is upholded, will group Pygostylia be dumbed as it then will be a junior synonym of Metornithes.
[Metornithes = All Aves closer to Neornithes than Archaeopteryx]
[Pygostylia = Last common ancester of Confuciusornis and Neornithes, and all it's descents]
Reference(s): Britt, B. B., Makovicky, P. J., Gauthier, J. & Bonde, N. 1998: Postcranial pneumatization in Archaeopteryx. –Nature: vol. 395, 24 September, pp. 374-376. Chiappe, L. M. 1995: The first 85 million years of avian evolution. –Nature: vol. 378, 23 November, pp. 349-355. Chiappe, L. M., Norell, M. A. & Clark, J. M. 1998: The skull of a relative of the stem-group bird Mononykus. –Nature: vol. 392, 19 March, pp. 275-278. Chiappe, L. M., Ji, Q. & Ji, S. 1998: Temporal morphology of the early bird Confuciusornis and its bearing on the evolution of avian cranial kinesis. –Journal of Vertebrate Paleontology: vol. 18, #3, suppl. to #3, pp. 33A Chiappe, L. M., Ji, Shu'an, Ji, Qiang & Norell, M. A., 1999: Anatomy and systematics of the Confuciusornithidae (Theropoda: Aves) from the Late Mesozoic of Northeastern China. --Bulletin of American Museum of Natural History: n:o 242, 1-89 Clarke, J. A., 2000: A new avialan from the Late Cretaceous of Mongolia –Journal of Vertebrate Paleontology: Vol. 20, #3, suppl. to #3, 36A Feduccia, A., 1980: The age of birds. --Harvard University Press, Cembridge, Massachusetts & London, England, 1980. IX-196. Feduccia, A. 1993: Evidence from claw geometry indicating arboreal habits of Archaeopteryx. --Science: vol. 259, 5 February, pp. 790-793. Feduccia, A. 1995: Explosive evolution in Tertiary birds and mammals. --Science: vol. 267, 3 February, pp. 637-638. Feduccia, A. 1996: The origin and evolution of birds. --Yale University Press, New Haven and London, 1996. xii-420. Gibbons, A. 1996: Early birds rise from China fossil beds. --Science: vol. 274, 15 November, p. 1083. Hou, L., Martin, L. D., Zhou, Z. & Feduccia, A. 1996: Early adaptive radiation of birds: Evidence from fossils from Northwestern China. --Science: vol. 274, 15 November, pp. 1164-1167. Hou, L., Martin, L. D., Zhou, Z., Feduccia, A. & Zhang, F., 1999: A diapsid skull in a new species of the primitive bird Confuciusornis. –Nature: vol. 399, 17 June, pp. 679-682 Keesey, T. M., 1996-2000: The Dinosauricon --Internet: http://dinosauricon.com/ Milner, A. C. 1993: Ground rules for early birds. –Nature: vol. 362, 15 April, p. 589. Morell, V. 1993: Archaeopteryx: Early bird catches a can of worms. --Science: vol. 256, 5 February, pp. 764-765. Novas, F. E. & Puerta, P. F. 1997: New evidence concerning avian origins from the Late Cretaceous of Patagonia. –Nature: vol. 387, 22 May, pp. 390-392. Padian, K. 1996: Early bird in slow motion. –Nature: vol. 382, 1 August, pp. 400-401. Padian, K. 1998: When is a bird not a bird? –Nature: vol. 393, 25 June, pp. 729-730. Sanz, J. L., Chiappe, L. M., Pérez-Moreno, B. P., Buscalioni, A. D., Moratalia, J. J., Ortega, F. & Poyato-Ariza, F. J. 1996: An Early Cretaceous bird from spain and it's implications for the evolution of avian flight. –Nature: vol. 382, 1 August, pp. 442-445. Sereno, P. C. & Chenggang, R. 1992: Early evolution af Avian fligth and perching: new evidence from the Lower Cretaceous of China. --Science: vol. 255, 14 February, pp. 845-848. Whetstone, K. N. 1983: Braincase of Mesozoic birds: I. New Preparation of the "London" Archaeopteryx. –Journal of Vertebrate Paleontology: vol. 2, #4,, pp. 439-452.