After Xu, Zhou & Wang, 2000, Xu, Norell, Wang, Makovicky & Wu, 2002, Clark, Norell & Makovicky, 2002, Hwang, Norell, Ji & Gao, 2002, Makovicky, Norell, Clark & Rowe, 2003, Senter, Barsbold, Britt & Burnham, 2004, Xu & Zhang, 2005, Makovicky, Apesteguía & Agnolín, 2005, Turner, Pol, Clarke, Erickson & Norell, 2007, and Novas, Pol, Canale, Porfiri & Calvo, 2009
<==o Paraves sensu Xu, Norell, Wang, Makovicky & Wu, 2002 (birds and near-birds; linnut ja sirppikynnet)
`--o Eumaniraptora
|-- †Pedopenna daohugouensis Xu & Zhang, 2005
|--o Avialae Gauthier 1986 sensu Chiappe, 1995, Chiappe, Ji, Ji & Norell, 1999 (lentävät teropodit, linnut; flying theropods, birds)
| |?- †Avialae genera et species indet. [Mongolia]
| |--o †Scansoriopterygidae Czerkas & Yuan, 2002 sensu Zhang, Zhou, Xu, Wang & Sullivan, 2008
| | |-- †Epidentrosaurus ningchengensis Zhang, Zhou, Xu & Wang, 2002 sensu Xu & Zhang, 2005 [Scansoriopteryx heilmanni Czerkas & Yuan, 2002 (invalid sr. synon.)]
| | `-- †Epidexipteryx hui Zhang, Zhou, Xu, Wang & Sullivan, 2008
| `-- Aves Linnaeus, 1758 sensu lato (linnut, birds)
`--o †Deinonychosauria sensu Xu, Norell, Wang, Makovicky & Wu, 2002
|--o †Troodontidae Gilmore, 1924d [Saurornithoididae Barsbold, 1974]
| |-- †Anchiornis huxleyi Xu et al., 2009; E.Cret. EAs.
| |--+-- †Mei long Xu & Norell, 2004; E.Cret. EAs.
| | `-- †Sinovenator changii Xu, Norell, Wang, Makovicky & Wu, 2002
| `--+-- Troodontidae genera et species indet. [EK troodontid sensu Makovicky, Norell, Clark & Rowe, 2003]
| |-- †Jinfengopteryx elegans Q. Ji, S. Ji, Lu, You, Chen, Liu, & Liu, 2005
| `--+-- †Sinornithoides youngi Russell & Dong, 1994
| |--+?- †Urbacodon itemirensis Averianov & Sues, 2007
| | |?- †Xixiasaurus henanensis Lü, Xu, Liu, Zhang, Jia & Ji, 2010
| | `-- †Byronosaurus jaffei Norell, Makovicky & Clark, 2000
| `--o †"Troodontinae" Gilmore, 1924d
| |-- †Troodon formosus Leidy, 1856 [Polyodontosaurus grandis Gilmore, 1932b, Stenonychosaurus inequalis Sternberg, 1932a, Pectinodon bakkeri Carpenter, 1982b]; L.Cret. WNAm.
| |-- †Linhevenator tani Xu, Tan, Sullivan, Han & Xiao, 2011; L.Cret. EAs.
| `--+-- †Saurornithoides mongoliensis Osborn, 1924a; L.Cret. EAs.
| `-- †Zanabazar junior (Barsbold, 1974) Norell, Makovicky, Bever, Balanoff, Clark, Barsbold & Rowe, 2009 [†Saurornithoides junior Barsbold, 1974; Zanabazer (lapsus calami)]; L.Cret. EAs.
`--o †Dromaeosauridae Matthew & Brown, 1922 sensu Makovicky, Apesteguía & Agnolín, 2005 and Turner, Pol, Clarke, Erickson & Norell, 2007
|-- †Mahakala omnogovae Turner, Pol, Clarke, Erickson & Norell, 2007 [IGM 100/1033]
|--o †Unenlagiinae Bonaparte, 1999 sensu Turner, Pol, Clarke, Erickson & Norell, 2007, and Novas, Pol, Canale, Porfiri & Calvo, 2009
| |-- †Shanag ashile Turner, Hwang & Norell, 2007 [IGM 100/1119]
| `--o †Unenlagiinae Bonaparte, 1999 sensu Makovicky, Apesteguía & Agnolín, 2005
| |-- †Buitreraptor gonzalezorum Makovicky, Apesteguía & Agnolín, 2005
| `--+-- †Austroraptor cabazai Novas, Pol, Canale, Porfiri & Calvo, 2009
| |-- †Rahonavis ostromi (Forster, Sampson, Chiappe & Krause, 1998a) [Rahona ostromi Forster, Sampson, Chiappe & Krause, 1998a]
| `--o †Unenlagia Novas & Puerta, 1997 [incl.? Neuquenraptor argentinus Novas & Pol, 2005]
| |-- †U. comahuensis Novas & Puerta, 1997
| `-- †U. paynemili Calvo, Porfiri & Kellner, 2004
`--o †Dromaeosauridae Matthew & Brown, 1922 sensu Xu, Zhou & Wang, 2000
|--o †Microraptoria Senter, Barsbold, Britt & Burnham, 2004 [Microraptorinae Burnham, Senter, Barsbold & Britt, 2004 sensu Makovicky, Apesteguía & Agnolín, 2005, and Turner, Pol, Clarke, Erickson & Norell, 2007]
| |-- †Bambiraptor feinbergorum Burnham, Derstler, Currie, Bakker, Zhou & Ostrom, 2000 emend. Olshevsky, 2000
| |?- †Tianyuraptor ostromi Zheng, Xu, You, Zhao & Dong, 2009; E.Cret. EAs. [feathered]
| `--+--o †Sinornithosaurus Xu, Wang & Wu, 1999; E.Cret. EAs. [feathered]
| | |-- †S. millenii Xu, Wang & Wu, 1999
| | `-- †S. haoiana Liu, Ji, Tang & Gao, 2004
| `--+-- †Graciliraptor lujiatunensis Xu & Wang, 2004; E.Cret. EAs. [feathered]
| `--o †Microraptor Xu, Zhou & Wang, 2000; E.Cret. EAs. [feathered]
| |-- †M. zhaoianus Xu, Zhou & Wang, 2000 [feathered]
| `-- †M. gui Xu, Zhou, Wang, Kuang, Zhang & Du, 2003 [Cryptovolans pauli (nomen nudum?)] (four-wing dinosaur) [feathered]
`--o †Dromaeosauridae Matthew & Brown, 1922 sensu Senter, Barsbold, Britt & Burnham, 2004
|-- †Atrociraptor marshalli Currie & Varricchio, 2004
|--+-- †Linheraptor exquisitus Xu, Choiniere, Pittman, Q. Tan, Xiao, Li, L. Tan, Clark, Norell, Hone & Sullivan, 2010; L.Cret. EAs.
| `-- †Tsaagan mangas Norell, Clark, Turner, Makovicky, Barsbold & Rowe, 2006 [IGM 100/1015]; L.Cret. EAs.
|--o †Velociraptorinae
| |-- †Velociraptor mongoliensis Osborn, 1924a; L.Cret. EAs.
| `-- †Balaur bondoc Csiki, Vremirb, Brusatte. & Norell, 2010; L.Cret. EEu.
`--o †Dromaeosaurinae
|-- †Adasaurus mongoliensis Barsbold, 1983; L.Cret. EAs.
`--+--o †Saurornitholestes Sues, 1978a; L.Cret. WNAm.
| |-- †S. langstoni Sues, 1978a
| `-- †S. robustus Sullivan, 2006
`--+-- †Deinonychus antirrhopus Ostrom, 1969a; L.Cret. WNAm.
`--+-- †Achillobator giganticus Perle, Norell, & Clark, 1999; L.Cret. WNAm.
`--+-- †Utahraptor ostrommaysi Kirkland, Gaston & Burge, 1993 [U. ostrommaysorum]; L.Cret. WNAm.
`-- †Dromaeosaurus albertensis Matthew & Brown, 1922; L.Cret. WNAm.
Paraves incertae sedis:
†Euronychodon
†E. asiaticus
†E. portucalensis Antunes & Sigogneau-Russell, 1991
†Kitadanisaurus
†Koreanosaurus
†Nuthetes destructor
†Ornithodesmus cluniculus [also treated as a troodontid]
Dromaeosauridae incertae sedis:
†Dromaeosauridae genera et species indet. [BPM 1 3-13]
†Dromaeosauridae genera et species indet. [M. Jur. England] [Evans & Milner, 1994] [teeth]
†Hulsanpes perlei Osmóska, 1982
†Paronychodon lacustris Cope, 1876a
†Pyroraptor olympius Allain & Taquet, 2000
†Variraptor mechinorum [nomen dubium?]
†Dromaeosaurus cristatus
†Dromaeosaurus gracilis
†Dromaeosaurus explanatus
†Dromaeosauroides bornholmensis Christiansen & Bonde, 2003 [tooth-taxa]
†Velociratorinae:
†Velociratorinae genera et species indet. [U. Jur. Portugal] [Rahut & Zinke, 1995] [teeth]
†Velociratorinae genera et species indet. [L. Cret. England] [Sweetman, 2004] [teeth] [IWCMS.2002.1., 2., 3., 4.; BMNH R 16510]
†Nuthetes destructor sensu Milner, 2002 [tooth-taxa]
Troodontidae incertae sedis:
†Borogovia gracilicrus Osmólska, 1987
†Elopteryx nopscai Andrews, 1913
†Heptasteornis andrewsi Harrison & Walker, 1975
†Koparion douglassi Chure, 1994
†Sinusonasus magnodens Xu & Wang, 2004
†Tochisaurus nemegtensis Kurzanov & Osmólska, 1991
†Tripriodon caperatus Marsh, 1889d
†Zapsalis abradens Cope, 1876b
†Troodontidae genera et species indet. [IGM 100/44]
Afternative phylogenies of Dromaeosauridae:
After Turner, Pol, Clarke, Erickson & Norell, 2007
<==o †Dromaeosauridae
|-- †Mahakala
`--+--+--o †Microraptorinae
| | |-- †Bambiraptor
| | `--+-- †Sinornithosaurus
| | `--+-- †Graciliraptor
| | `-- †Microraptor
| `--o †Unenlagiinae
| |-- †Shanag
| `--+-- †Buitreraptor
| `--+-- †Rahonavis
| `-- †Unenlagia
`--+?- †Atrociraptor
|--o †Dromaeosaurinae
| |-- †Adasaurus
| |-- †Dromaeosaurus
| `--+-- †Achillobator
| `-- †Utahraptor
`--o †Velociraptorinae
|-- †Tsaagan
`--+-- †Saurornitholestes
`--+-- †Velociraptor
`-- †Deinonychus
|
After Novas, Pol, Canale, Porfiri & Calvo, 2009
<==o †Dromaeosauridae
|-- †Mahakala
`--+--+--o †Dromaeosaurinae
| | |-- †Achillobator
| | |-- †Adasaurus
| | |-- †Dromaeosaurus
| | `-- †Utahraptor
| `--o †Velociraptorinae
| |-- †Tsaagan
| `--+-- †Saurornitholestes
| `--+-- †Velociraptor
| `-- †Deinonychus
`--o †Microraptorinae [paraphyletic]
|-- †Sinornithosaurus
|--+-- †Graciliraptor
| `-- †Microraptor
`--o †Unenlagiinae
|-- †Shanag
`--+-- †Buitreraptor
`--+-- †Austroraptor
|-- †Rahonavis
`-- †Unenlagia
|
After Zheng, Xu, You, Zhao & Dong, 2009
<==o †Dromaeosauridae
|--o †Unenlagiinae
| |-- †Buitreraptor
| |-- †Rahonavis
| `-- †Unenlagia
`--+-- †Bambiraptor
|-- †Tianyuraptor
|-- †Tsaagan
|-- †Adasaurus
|--+-- †Sinornithosaurus
| `--+-- †Microraptor
| `-- NGMC91
`--+-- †Velociraptor
|-- †Saurornitholestes
`--+-- †Deinonychus
`--+-- †Atrociraptor
`--+-- †Achillobator
`--+-- †Dromaeosaurus
`-- †Utahraptor
|
Afternative phylogenies of Troodontidae:
After Xu, Tan, Sullivan, Han & Xiao, 2011
<==o †Troodontidae
|-- †Sinovenator
`--+-- †Anchiornis
`--+-- †Mei
`--+--†Byronosaurus
`--+-- IGM100/44
`--+-- †Sinornithoides
`--+-- †Troodon formosus Leidy, 1856 [Polyodontosaurus grandis Gilmore, 1932b, Stenonychosaurus inequalis Sternberg, 1932a, Pectinodon bakkeri Carpenter, 1982b]; L.Cret. WNAm.
|-- †Linhevenator tani Xu, Tan, Sullivan, Han & Xiao, 2011; L.Cret. EAs.
`--+-- †Saurornithoides mongoliensis Osborn, 1924a; L.Cret. EAs.
`-- †Zanabazar junior (Barsbold, 1974) Norell, Makovicky, Bever, Balanoff, Clark, Barsbold & Rowe, 2009 [†Saurornithoides junior Barsbold, 1974; Zanabazer (lapsus calami)]; L.Cret. EAs.
|
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