Mikko's Phylogeny Archive

†Ankylosauridae – armored dinosaurs

After Carpenter, 2001, and Lü, Ji, Gao & Li, 2007

<==o †Ankylosauridae Brown, 1908 (nuijahäntäpanssariselät; club-tailed ankylosaurs)
   |?- †Minotaurasaurus ramachandrani C. A. Miles & C. J. Miles, 2009; L.Cret. CAs.
   |?- †Tatankacephalus cooneyorum W. L. Parsons & K. M. Parsons, 2009; E.Cret. WNAm.
   |?- †Oohkotokia horneri
   |?- †Zuul crurivastator
   |?-o †Anodontosaurus
   |  |-- †A. inceptus
   |  `-- †A. lambei
   |?-o †Scolosaurus
   |  |-- †S. cutleri
   |  `-- †S. thronus
   |--o Shamosaurinae Tumanova, 1983
   |  |?- †Aletopelta coombsi Ford & Kirkland, in Carpenter, 2001
   |  |?- †Cederpelta bilbayhallorum Carpenter, Kirkland, Burge & Bird, in Carpenter, 2001
   |  `--+-- †Gobisaurus domoculus Vickaryous, Russell, Currie & Zhao, 2001
   |     `-- †Shamosaurus scutatus Tumanova, 1983
   `--o †Ankylosaurinae Brown, 1908
      |-- †Ankylosaurus magniventris Brown, 1908
      `--+?-o †Crichtonsaurus Dong, 2002 sensu Lü, Ji, Gao & Li, 2007
         |  |-- †D. bohlini Dong, 2002
         |  `-- †C. benxiensis Lü, Ji, Gao & Li, 2007
         `--+--+-- †Tarchia gigantea (Maleev, 1956) Marya´nska, 1977 [Dyoplosaurus giganteus Maleev, 1956; incl. Tarchia kielanae Marya´nska, 1977]
            |  `--+-- †Saichania chulsanensis Marya´nska, 1977 [Tianzhenosaurus youngi Pang & Cheng, 1998]
            |     `-- †Tsagantegia longicranialis Tumanova, 1993
            `--+-- †Talarurus plicatospineus Maleev, 1952b
               `--+-- †Euoplocephalus tutus (Lambe, 1902) Lambe, 1910 [Synonyms]
                  `--+-- †Nodocephalosaurus kirtlandensis Sullivan, 1999
                     |?- †Tianzhenosaurus youngi Peng & Cheng, 1998 sensu Lü, Ji, Gao & Li, 2007
                     `--o †Pinacosaurus Gilmore, 1933b [Syrmosaurus Maleev, 1952a]
                        |-- †P. grangeri [Syrmosaurus viminiculatus Maleev, 1952a; Pinacosaurus ninghsiensis Young, 1935b]
                        `-- †P. mephistocephalus Godefroit, Pereda Suberbiola & Dong, 1999

Alternative phylogenies of Ankylosauridae:

<==o †Ankylosauridae Brown, 1908
   |-- †Gargoyleosaurus
   `--+-- †Gastonia
      `--+-- †Gobisaurus
         `--+-- †Minmi
            `--+-- †Pinacosaurus
               `--+-- †Talarurus
                  `--+--o Shamosaurinae
                     |  |-- †Shamosaurus
                     |  `-- †Tsagantegia
                     `--o †Ankylosaurinae
                        |--+-- †Tarchia
                        |  `-- †Saichania
                        |-- †Tianzhenosaurus
                        |-- †Nodocephalosaurus
                        |-- †Shanxia
                        |-- †Ankylosaurus
                        `-- †Euoplocephalus

<==o †Ankylosauridae
   |--o Shamosaurinae
   |  |?- †Aletopelta
   |  |?- †Cederpelta
   |  `--+-- †Gobisaurus
   |     `-- †Shamosaurus
   `--o †Ankylosaurinae
      |-- †Ankylosaurus
      `--+-- †Crichtonsaurus
         `--+--+-- †Tarchia
            |  `--+-- †Saichania
            |     `-- †Tsagantegia
            `--+-- †Talarurus
               `--+-- †Euoplocephalus
                  `--+-- †Nodocephalosaurus
                     |-- †Tianzhenosaurus
                     `-- †Pinacosaurus

<==o †Ankylosauridae
   |-- †Cederpelta
   `--+-- †Minmi
      `--+-- †Shamosaurus
         |-- †Gobisaurus
         `--+-- †Tsagantegia
            |--+-- †Gastonia
            |  `-- †Tatankacephalus
            `--+== †Pinacosaurus
               |-- †Tianzhenosaurus
               |-- †Talarurus
               |-- †Tarchia
               |-- †Saichania
               `--+-- †Ankylosaurus
                  `-- †Euoplocephalus

Reference(s):

• Barrett, P. M., You, H., Upchurch, P. & Burton, A. C., 1998: A new ankylosaurian dinosaur (Ornithischia: Ankylosauria) from the Upper Cretaceous of Shanxi Province, People's Republic of China.
  –Journal of Vertebrate Paleontology: Vol. 18, #2, pp. 376-384
• Carpenter, K., 2001: Phylogenetic analysis of the Ankylosauria. xv-526
  in Carpenter, K., (ed.) 2001: The Armored Dinosaurs.
  –Indiana University Press, Bloomington & Indianapolis, 2001, pp. xv-526
• Carpenter, K., Kirkland, J. I., Burge, D. & Bird, J., 2001: Disarticulated skull of a new primitive ankylosaurid from the Lower Cretaceous of Eastern Utah. 211-238
  in Carpenter, K., (eds.) 2001: The Armored Dinosaurs.
  –Indiana University Press, Bloomington & Indianapolis, 2001, pp. xv-526
• Carpenter, K., Miles, C. & Cloward, K., 1998: Skull of a Jurassic ankylosaur (Dinosauria).
  –Nature: Vol. 393, 25 June, pp. 782-783
• Coombs, W. P. Jr., & Marya´nska, T., 1990: Ankylosauria. 456-483
  in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
  –University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733
• Ford, T. L. & Kirkland, J. I., 2001: Carlsbad ankylosaur (Ornithichia, Ankylosauria): an ankylosaurid and not a nodosaurid. 239-260
  in Carpenter, K., (eds.) 2001: The Armored Dinosaurs.
  –Indiana University Press, Bloomington & Indianapolis, 2001, pp. xv-526
• Hill, R. V., Witmer, L. M. & Norell, M. A., 2003: A New Specimen of Pinacosaurus grangeri (Dinosauria: Ornithischia) from the Late Cretaceous of Mongolia: Ontogeny and Phylogeny of Ankylosaurs.
  –American Museum Novitates: #3395, pp. 1-29
• Keesey, T. M., 1996-2000: The Dinosauricon
  –iNet: http://dinosauricon.com/
• Lü, J.-C., Ji, Q., Gao, Y. & Li, Z., 2007: A new species of the ankylosaurid dinosaur Crichtonsaurus (Ankylosauridae: Ankylosauria) from the Cretaceous of Liaoning Province, China.
  –Acta Geologica Sinica (English Edition): Vol. 81, #6, pp. 883-897
• Miles, C. A. & Miles, C. J., 2009: Skull of Minotaurasaurus ramachandrani, a new Cretaceous ankylosaur from the Gobi Desert.
  –Current Science: Vol. 96, #1, pp. 65-70
• Parsons, W. L. & Parsons, K. M., 2009: A new ankylosaur (Dinosauria: Ankylosauria) from the Lower Cretaceous Cloverly Formation of central Montana.
  –Canadian Journal of Earth Sciencies: Vol. 46, pp. 721-738 [doi: 10.1139/E09-045]
• Sullivan, R. M., 1999: Nodocephalosaurus kirtlandensis, gen. et sp. nov., a new ankylosaurid dinosaur (Ornithischia: Ankylosauria) from the Upper Cretaceous Kirtland Formation (Upper Campanian), San Juan Basin, New Mexico.
  –Journal of Vertebrate Paleontology: Vol. 19, #1, pp. 126-139
• Vickaryous, M. K., Russell, A. P., Currie, P. J. & Zhao, X., 2001: A new ankylosaur (Dinosauria: Ankylosauria) from the Lower Cretaceous of China, with comments on ankylosaurian relationships.
  –Canadian Journal of Earth Sciencies: Vol. 38, #12, pp. 1767-1780