After Frost et al., 2006, with Recent species from Frost, 2000-2002 [“AMNH Amphibian species of the World”]
<==o Leptodactylidae Werner, 1896 “1838” [taxonomic history] |--o Edalorhina Jiménez de la Espada, 1871 “1870” | |-- E. nasuta | `-- E. perezi |-- Engystomops Jiménez de la Espada, 1872 |-- Eupemphix Steindachner, 1863 |-- Hydrolaetare schmidti Gallardo, 1963 |-- Leptodactylus Fitzinger, 1826 [incl. Adenomera Steindachner, 1867; Lithodytes Fitzinger, 1843; Vanzolinius Heyer, 1974] |--o Paratelmatobius Lutz & Carvalho, 1958 | |-- P. cardosoi | |-- P. gaigeae | |-- P. lutzii | |-- P. mantiqueira | |-- P. pictiventris | `-- P. poecilogaster |-- Physalaemus Fitzinger, 1826 |--o Pleurodema Tschudi, 1838 | |-- P. bibroni | |-- P. borellii | |-- P. brachyops | |-- P. bufonina | |-- P. cinerea | |-- P. diplolistris | |-- P. guayapae | |-- P. kriegi | |-- P. marmorata | |-- P. nebulosa | |-- P. thaul | `-- P. tucumana |--o Pseudopaludicola Miranda-Ribeiro, 1926 | |-- P. boliviana | |-- P. ceratophryes | |-- P. falcipes | |-- P. llanera | |-- P. mirandae | |-- P. mineira | |-- P. mystacalis | |-- P. pusilla | |-- P. riopiedadensis | |-- P. saltica | `-- P. ternetzi |-- Scythrophrys sawayae Lynch, 1971 `-- Somuncuria somuncurensis Lynch, 1978
Reference(s):
- Frost, D. R., 2000: Amphibian species of the World: An Online Reference.
–iNet: American Museum of Natural History, Department of Herpetology: http://research.amnh.org/herpetology/amphibia/index.html - Frost, D. R., Grant, T., Faivovich, J., Bain, R. H., Haas, A., Haddad, C. F. B., de Sá, R. O., Channing, A., Wilkinson, M., Donnellan, S. C., Raxworthy, C. J., Campbell, J. A., Blotto, B. L., Moler, P., Drewes, R. C., Nussbaum, R. A., Lynch, J. D., Green, D. M. & Wheeler, W. C., 2006: The amphibian tree of life.
–Bulletin of the American Museum of Natural History: #297, pp. 1-370 - Grant, T., Frost, D. R., Caldwell, J. P., Gagliardo, R., Haddad, C. F. B., Kok, P. J. R., Means, D. B., Noonan, B. P., Schargel, W. E. & Wheeler, W. C., 2006: Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae).
–Bulletin of the American Museum of Natural History: Vol. #299, pp. 1-262