After Frost et al., 2006
Alternative phylogeny after Grant et al., 2006
<==o Ceratophryidae Tschudi, 1838 [taxonomic history] (sarvikonnat, sarvisammakot) |-- †Watoelia |-- Telmatobius Wiegmann, 1834 [Telmatobiinae Fitzinger, 1843] `--o Ceratophryinae Tschudi, 1838 |--o Batrachylini Gallardo, 1965 sensu Frost et al, 2006 | |-- Atelognathus Lynch, 1978 | |-- Batrachyla Bell, 1843 | `?- Insuetophrynus acarpicus Barrio, 1970 `--o Ceratophryini Tschudi, 1838 sensu Frost et al, 2006 |-- Ceratophrys Wied-Neuwied, 1824 |-- Chacophrys Reig & Limeses, 1963 `-- Lepidobatrachus Budgett, 1899
After Grant et al., 2006, with species from Frost, 2000-2002 [“AMNH Amphibian species of the World”]
<==o Ceratophryidae Tschudi, 1838 [taxonomic history] (sarvikonnat, sarvisammakot) |-- †Watoelia |?- Insuetophrynus acarpicus Barrio, 1970 |--o Batrachylinae Gallardo, 1965 sensu Grant et al., 2006 [Ceratophryinae: Batrachylini Gallardo, 1965 sensu Frost et al, 2006 | |--o Atelognathus Lynch, 1978 | | |-- A. ceii | | |-- A. grandisonae | | |-- A. nitoi | | |-- A. patagonicus | | |-- A. praebasalticus | | |-- A. reverberii | | |-- A. salai | | `-- A. solitarius | `--o Batrachyla Bell, 1843 | |-- B. antartandica | |-- B. fitzroya | |-- B. leptopus | |-- B. nibaldoi | `-- B. taeniata `--+-- Telmatobius Wiegmann, 1834 [Telmatobiinae Fitzinger, 1843] `--o Ceratophryinae Tschudi, 1838 sensu Grant et al., 2006 [Ceratophryinae: Ceratophryini Tschudi, 1838 sensu Frost et al., 2006] |--+-- †Beelzebufo ampinga Evans, Jones & Krause, 2008; U.Cret. Mad. | `--o Ceratophrys Wied-Neuwied, 1824 | |-- C. aurita | |-- C. calcarata | |-- C. cornuta | |-- C. cranwelli | |-- C. joazeirensis | |-- C. ornata | |-- C. stolzmanni | `-- C. testudo `--+--o Lepidobatrachus Budgett, 1899 | |-- L. asper | |-- L. laevis | `-- L. llanensis `--+-- †Baurubatrachus pricei Báez & Peri, 1989; U.Cret. SAm. |-- †Wawelia gerholdi; Mioc. SAm. `-- Chacophrys pierottii Reig & Limeses, 1963
Reference(s):
- Evans, S. E., Jones, M. E. H. & Krause, D. W., 2008: A giant frog with South American affinities from the Late Cretaceous of Madagascar.
–Proceedings of the National Academy of Science [PNAS]: Vol. 105, #8, pp. 2951-2956 [doi: 10-1073/pnas.0707599105] [http://www.pnas.org/cgi/doi/10.1073/pnas.0707599105] - Frost, D. R., 2000: Amphibian species of the World: An Online Reference.
–iNet: American Museum of Natural History, Department of Herpetology: http://research.amnh.org/herpetology/amphibia/index.html - Frost, D. R., Grant, T., Faivovich, J., Bain, R. H., Haas, A., Haddad, C. F. B., de Sá, R. O., Channing, A., Wilkinson, M., Donnellan, S. C., Raxworthy, C. J., Campbell, J. A., Blotto, B. L., Moler, P., Drewes, R. C., Nussbaum, R. A., Lynch, J. D., Green, D. M. & Wheeler, W. C., 2006: The amphibian tree of life.
–Bulletin of the American Museum of Natural History: #297, pp. 1-370 - Grant, T., Frost, D. R., Caldwell, J. P., Gagliardo, R., Haddad, C. F. B., Kok, P. J. R., Means, D. B., Noonan, B. P., Schargel, W. E. & Wheeler, W. C., 2006: Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae).
–Bulletin of the American Museum of Natural History: Vol. #299, pp. 1-262