After the ”Tree of Life“, Steward & Rothwell, 1993, and Hassler & Swale, 2001, with Finnish species from Hämet-Ahti et al., 1984
<==o Huperzia Bernhardi, 1800 (ketunlieot) |-- H. acerosa |-- H. acifolia |-- H. acuta |-- H. acutifolia |-- H. affinis |-- H. afromontana |-- H. albescens |-- H. aloifolia |-- H. amentacea |-- H. andina |-- H. apolinari-mariae |-- H. appalachiana |-- H. aqualupiana |-- H. archboldiana |-- H. arcuata |-- H. aristei |-- H. ascendens |-- H. attenuata |-- H. australiana |-- H. australis |-- H. austroecuadorica |-- H. austrosinica |-- H. badiniana |-- H. balansae |-- H. bampsiana |-- H. banayanica |-- H. beccarii |-- H. beiteliana |-- H. beitelii |-- H. biformis Huperzia binervia Huperzia bolanica Huperzia boliviana Huperzia bonapartei Huperzia borneensis Huperzia brachiata Huperzia brachystachys Huperzia bradeorum Huperzia brasiliana Huperzia brassii Huperzia brevifolia Huperzia brongniartii Huperzia bucahwangensis Huperzia campestris Huperzia campiana Huperzia cancellata Huperzia capellae Huperzia capillaris Huperzia carinata Huperzia catacachiensis Huperzia catharinae Huperzia cavifolia Huperzia ceylanica Huperzia chamaeleon Huperzia chinensis Huperzia chiricana Huperzia chishuiensis Huperzia christii Huperzia cipoensis Huperzia clarkiana Huperzia clausenii Huperzia colanensis Huperzia columnaris Huperzia comans Huperzia compacta Huperzia copelandiana Huperzia copelandii Huperzia coralium Huperzia coriacea Huperzia costaricensis Huperzia crassa Huperzia crebra Huperzia crispata Huperzia crucis-australis Huperzia cruenta Huperzia cryptomeriana Huperzia cubana Huperzia cuernavacensis Huperzia cumingii Huperzia cuneifolia Huperzia curiosa Huperzia curvifolia Huperzia dacrydioides Huperzia dalhousiana Huperzia darwiniana Huperzia delavayi Huperzia delbrueckii Huperzia deminuens Huperzia dentata Huperzia desvauxii Huperzia dianae Huperzia dichaeoides Huperzia dichotoma Huperzia dielsii Huperzia dura Huperzia echinata Huperzia ellenbeckii Huperzia elmeri Huperzia emeiensis Huperzia engleri Huperzia eremorum Huperzia ericifolia Huperzia erosa Huperzia erythrocaulon Huperzia espinosana Huperzia everettii Huperzia eversa Huperzia fargesii Huperzia feejeensis Huperzia filiformis Huperzia firma Huperzia flagellacea Huperzia foliacea Huperzia foliosa Huperzia fontinaloides Huperzia fordii Huperzia formosana Huperzia friburgensis Huperzia fuegiana Huperzia funiformis Huperzia gagnepainiana Huperzia galapagensis Huperzia gedeana Huperzia gigantea Huperzia glauca Huperzia gnidioides Huperzia goebelii Huperzia goliathensis Huperzia goudotii Huperzia guangdongensis Huperzia gunturensis Huperzia haeckelii Huperzia haitensis Huperzia haleakalae Huperzia hamiltonii Huperzia hartwegiana Huperzia hastata Huperzia hellwigii Huperzia hemleri Huperzia henryi Huperzia herteriana Huperzia heterocarpon Huperzia heteroclita Huperzia hexasticha Huperzia hildebrandtii Huperzia hilliana Huperzia hippuridea Huperzia hippuris Huperzia hochreutineri Huperzia hoffmannii Huperzia hohenackeri Huperzia holstii Huperzia homocarpa Huperzia hookeri Huperzia horizontalis Huperzia huberi Huperzia humbertii-henrici Huperzia hupehensis Huperzia hypogaea Huperzia hystrix Huperzia iajouensis Huperzia ignambiensis Huperzia innocentia Huperzia insularis Huperzia intermedia Huperzia itambensis Huperzia jaegeri Huperzia juniperistachya Huperzia kajewskii Huperzia kamaensis Huperzia kangdingensis Huperzia kaysseri Huperzia kesselringii Huperzia keysseri Huperzia killipii Huperzia kuesteri Huperzia kunmingensis Huperzia laipoensis Huperzia lajouensis Huperzia lancifolia Huperzia lauterbachii Huperzia laxa Huperzia lechleri Huperzia ledermannii Huperzia leipoensis Huperzia leishanensis Huperzia leitzii Huperzia leptodon Huperzia liangshanica Huperzia lignosa Huperzia lindenii Huperzia linifolia Huperzia llanganatensis Huperzia lockyeri Huperzia loefgreniana Huperzia longa Huperzia longyangensis Huperzia loxensis Huperzia lucidula Huperzia luederwaldtii Huperzia macbridei Huperzia macgregorii Huperzia macrostachys Huperzia maerhkangensis Huperzia mafuluensis Huperzia magnifica Huperzia magnusiana Huperzia mannii Huperzia marsupiiformis Huperzia martii Huperzia medogensis Huperzia megastachya Huperzia melanesica Huperzia merrillii Huperzia mesoamericana Huperzia mexicana Huperzia mildbraedii Huperzia mingcheensis Huperzia miniata Huperzia minima Huperzia minimadenta Huperzia minutifolia Huperzia mira Huperzia miyoshiana (Makino, 1981) Ching; Acta Bot. Yunnan 3(3): 303, 304 () Distribution [complete]: E-Siberia / Amur / Ussuri, Japan, Korea, S-Alaska, Canada (W-British Columbia, Newfoundland), USA (Washington) Remarks: selago agg. = Huperzia selago var. miyoshiana (Makino) Taylor & MacBryde; Canadian Journal of Botany 56(2): 185 (1978) = Lycopodium miyoshianum Makino; Bot. Mag. (Tokyo) 12: 36 (1898) = Lycopodium selago ssp. miyoshianum (Makino) Calder & Taylor; Canadian Journal of Botany 43(11): 1388 (1965) = Lycopodium selago var. miyoshianum (Makino) Makino; Botanical Magazine 16(189): 199 (1902) = Lycopodium tenuifolium Herter; Bot. Jahrb. Syst. 43(1, Beibl. 98): 41 (1909) = Urostachys miyoshianum (Makino) Herter ex Nessel; Die Bärlappgewächse 28, Abb. 4, f. 4 (1939) var. coreana (Hayata) Ching; Acta Botanica Yunnanica 3(3): 304 (1981) Distribution: Korea = Lycopodium miyoshianum var. coreanum Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 5: 255 (1915) = Urostachys coreanus Herter; Index Lyc. 57 (1949) = Urostachys cryptomerinus (Maxim.) Nessel var. coreanus Herter ex Nessel; [Bärlappgewächse 72, t. 11, f. 8. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 160. 1940 = Urostachys miyoshiana var. coreanus (Hayata) Herter ex Nessel; Die Bärlappgewächse 28, Abb. 4, f. 5 (1939) Huperzia mollicoma (Spring) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Costa Rica, Panama, Colombia, Venezuela, Ecuador, ?Peru, Bolivia, SE-Brazil, C-Brazil Remarks: verticillata agg. = Huperzia flaccida (Fée) Holub; Folia Geobot. Phytotax. 20: 72 (1985) = Lycopodium dichotomum ssp. mollicomum Spring; Flora 21(1): 162 (1838) = Lycopodium flaccidum Fée; Crypt. Vasc. Bras. 2: 92, t. 106, f. 1 (1872)-73 [nom. illeg., non Lycopodium flaccidum Bory (1833) = Lycopodium mollicomum (Spring) Spring; Flora Brasiliensis 1(2): 113 (1840) = Lycopodium setaceum Lam. var. brasiliense Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 43 (1842) = Lycopodium williamsii Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33(2): 112 (1906) = Urostachys flaccidus (Fée) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923) = Urostachys mollicomus (Spring) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 407, t. 25 (1927) = Urostachys neptuni Herter; Estudios Botánicos en la Región Uruguaya 20: 72 (1949) [nom. illeg. superfl. = Urostachys williamsii (Underw. & Lloyd) Herter ex Nessel; Die Bärlappgewächse 164 (1939) = Urostachys mollicomus (Spring) Nessel var. major Nessel; Repert. Spec. Nov. Regni Veg. 36: 183, t. 172. 1934 Huperzia molongensis (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Ecuador, Peru, Venezuela, Colombia Remarks: phlegmaria agg. = Lycopodium molongense Herter; Bot. Jahrb. Syst. 43: Beibl. 98.- 51 (1909) = Urostachys molongensis (Herter) Herter ex Nessel; Bärlappgewächse 240 (1939) Huperzia montana (L. Underw. & F. Lloyd) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Jamaica, Cuba, Hispaniola Remarks: brongniartii agg. = Lycopodium hippurideum var. montanum (Underw. & F.E. Lloyd) Proctor; Amer. Fern Journal 72: 107 (1982) = Lycopodium montanum Underw. & F.E. Lloyd; Bulletin of the Torrey Botanical Club 33: 107 (1906) = Urostachys hippurideus var. montanus (Underw. & F.E. Lloyd) Herter ex Nessel; Die Bärlappgewächse 88 (1939) = Urostachys montanus (Underw. & F.E. Lloyd) Herter; Estudios Botánicos en la Región Uruguaya 20: 70 (1949), Alt. title: Index Lycopodiarum 70 (1949) Huperzia monticola (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: Sumatra Remarks: unverified name, selago agg. = Lycopodium monticolum Alderw.; Malayan ferns & fern allies, Suppl. 1, corr.: 65. 1917 [Bull. Jard. Bot. Buitenzorg, ser. 2, 28: 43, t. 7 (1918]) = Urostachys monticolus (Alderw.) Herter ex Nessel; Bärlappgewächse 120 (1939) Huperzia mooreana (Baker) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: Brazil (Bahia) Remarks: brongniartii agg. = Lycopodium luetzelburgii Rosenstock; Repert. Spec. Nov. Regni Veg. 20: 95 (1924) = Huperzia sydowiora (Herter) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21(100): 157 (1988) = Lycopodium sydowiorum Herter; Repert. Spec. Nov. Regn. Veg. 13: 296 (1914) = Urostachys sydowiorum (Herter) Herter ex Nessel; Archivos de Botânica do São Paulo 1: 390 (1927) = Lycopodium mooreanum Hort. Sander ex Baker; Gard. Chron., ser. 3, 12: 582 (1892) = Urostachys mooreanus (Baker) Herter; Index Lyc. 70 (1949) Huperzia mortonii (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: Guatemala Remarks: unverified name, dichotoma agg. = Urostachys mortonii Herter; F. Morton; Repert. Spec. Nov. Regni Veg. 28: 108 (1930) Huperzia multidichotoma Ching; Acta Bot. Yunnanica 3: 295 (1981) Distribution: Sichuan Remarks: selago agg. Huperzia multifaria (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: Ternate Remarks: unverified name, hippuris agg. = Lycopodium multifarium Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 226 (1922) = Urostachys multifarius (Alderw.) Herter ex Nessel; Bärlappgewächse 207 (1939) Huperzia myriophyllifolia (Hayata) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: Taiwan Remarks: selago agg. = Lycopodium serratum Thunb. ex Murray var. myriophyllifolium Hayata; Ic. pl. formos. 4: 133 (1914) = Urostachys myriophyllifolius (Hayata) Herter; Index Lyc. 71 (1949) Huperzia myrsinites (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874) Distribution [complete]: Mexico (Chiapas, Oaxaca), Guatemala, Honduras, El Salvador, Nicaragua, Costa Rica, Colombia, Venezuela, Ecuador, Peru, Hispaniola, Trinidad, British Guyana, Bolivia Remarks: myrsinites agg. = Lycopodium myrsinites Lam.; Encyclopédie Méthodique, Botanique 3: 654. [1789] (1792) = Lycopodium patens Willd. ex Spreng.; Systema Vegetabilium ed. 14 4: 12 (1827) = Lycopodium roraimense Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 115 (1906) = Lycopodium skutchii Maxon; Proc. Biol. Soc. Wash. 46: 159 (1933) = Plananthus myrsinites (Lam.) P. Beauv.; Prodr. Aetheog. 111 (1805) = Urostachys myrsinites (Lam.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 166 (1923) = Urostachys patens (Willd. ex Spreng.) Herter ex Nessel; Die Bärlappgewächse 245, t. 59, f. 25 (1939) = Urostachys roraimensis (Underw. & Lloyd) Herter ex Nessel; Die Bärlappgewächse 246 (1939) Huperzia myrtifolia (G. Forster) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Queensland, Tasmania Remarks: myrtifolia agg. = Lycopodium myrtifolium G. Forster; Fl. ins. austr. 87 (1786) = Stachygynandrum myrtifolium (G. Forster) P. Beauv.; Prodr. aethéogam. 113 (1805) = Urostachys myrtifolius (G. Forster) Herter; Index Lyc. 71 (1949) Huperzia myrtuosa (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution: Venezuela Remarks: brevifolia agg. = Huperzia lellingeri (Rolleri) Holub; Folia Geobot. Phytotax. 20: 74 (1985) = Lycopodium lellingeri Rolleri; Hickenia 1 (26): 143 (1978) = Lycopodium myrtuosum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 9 (1849) = Urostachys myrtuosus (Spring) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 387 (1927) Huperzia nana (C.Y. Ma) comb. ined.; [nom. illeg., non Herter 1814 Distribution: China = Phlegmariurus nanus C.Y. Ma (1990) Huperzia nanchuanensis (Ching & H.S. Kung) Ching & H.S. Kung; Acta Botanica Yunnanica 3(3): 302 (1981) Distribution: Sichuan Remarks: selago agg. = Lycopodium nanchuanense Ching & Kung ex Kung; Acta Phytotax. Sin. 18 (2): 235, f. 5, t. 6, f. 2 (1980) Huperzia neocaledonica (Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: New Caledonia Remarks: hippuris agg. = Urostachys neocaledonicus Nessel; Repert. Spec. Nov. Regni Veg. 36: 187, t. 175 (1934) Huperzia nesselii (Brause ex Nessel) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21(100): 156 (1988) Distribution [complete]: Peru Remarks: saururus agg. = Urostachys nesselii Brause ex Nessel; [Bärlappgewächse 78, f. 12. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 161 (1940) = Lycopodium nesselii (Nessel) Rolleri; Revista Mus. La Plata (N. S.) Bot. 13: 77 (1981) Huperzia nilagirica (Spring) Dixit; J. Bombay Nat. Hist. Soc. 77 (3): 541 (1981) Distribution: India (Nilgiri Hills, Tamil Nadu, Kerala) Remarks: unverified name, hamiltonii agg. = Lycopodium niligaricum Spring; Bull. Acad. Sci. Belg. 15 (1): 58 (1843) Huperzia nuda (Nessel) B. Øllg. & Wind.; Bradea 5: 14 (1987) Distribution: Brazil Remarks: brongniartii agg. = Urostachys nudiusculus Herter; Index Lyc. 72. 1949 [nom. superfl. for Urostachys nudus Nessel nudus Christ ex Nessel, Arch. Bot. Sao Paulo 1: 396, t. 14 (1927) Huperzia nummulariifolia (Blume) Chambers, Jermy & Crabbe; Brit. Fern Gaz. 10: 176 (1971) Distribution: peninsular Malaysia (Kedah, Penang Isl., Kelantan, Pahan, Johor), Java, Sumatra, Thailand, Nicobars, Fiji (Viti Levu, Ovalau, Taveuni) Remarks: nummulariifolia agg. = Lycopodium nummulariifolium Blume; Enum. pl. Javae 2: 263 (1828) = Phlegmariurus nummulariifolius (Blume) Ching; Acta Bot. Yunnanica 3: 298 (1981) = Urostachys nummulariifolius (Blume) Herter; Philipp. Sci. 22: 182 (1923) = Lycopodium rotundifolium Roxb.; Calcutta J. Nat. Hist. 4: 473 (1844) Huperzia nutans (Brackenr.) Rothm.; Feddes Repert. 54: 62 (1944) Distribution [complete]: Hawaii (Kauai, Oahu) Remarks: squarrosa agg. Conservation status: IUCN E = Lycopodium nutans Brackenr.; U. S. Expl. exped., Filic. 16: 327, t. 46 (1854) = Urostachys nutans (Brack.) Herter ex Nessel; Bärlappgewächse 208. 1939 = Huperzia schlechteri (E. Pritzel) Holub; Folia Geobot. Phytotax. 20: 76 (1985) = Lycopodium schlechteri E. Pritzel; Schlechter, Bot. Jahrb. Syst. 39: 14 (1906) = Urostachys schlechteri (E. Pritzel) Herter ex Nessel; Bärlappgewächse 207 (1939) Huperzia nylamensis Ching & S.K. Wu; Acta Botanica Yunnanica 3(3): 305 (1981) Distribution: Tibet Remarks: unverified name, selago agg. = Phlegmariurus nylamensis (Ching & S.K. Wu) H.S. Kung & L.B. Zhang; Acta Phytotaxonomica Sinica 37(1): 52 (1999) = Huperzia nylamensis Ching & S.K. Wu; Flora Xizangica 1: 9, t. 2, f. 1-3, t. 1, f. 4-6 (1983) [nom. illeg., superfl. Huperzia obovalifolia (Bonap.) comb. ined. Distribution: Vietnam Remarks: unverified name, obtusifolia agg. = Lycopodium obovalifolium Bonap.; Notes Pteridol. 14: 190 (1923) = Urostachys obovalifolius (Bonap.) Nessel; Bärlappgewächse 249 (1939) Huperzia obscure-denticulata Ching; Acta Bot. Yunnanica 3: 295 (1981) Distribution: NW-Sichuan Remarks: unverified name, selago agg. Huperzia obtusifolia (P. Beauv.) Rothm.; Feddes Repert. 54: 61 (1944) Distribution [complete]: Madagascar, Réunion Remarks: obtusifolia agg. = Lepidotis obtusifolia P. Beauv.; Prodr. aetheogam. 109 (1805) = Lycopodium obtusifolium (P. Beauv.) Sw.; Syn. fil. 177 (1806) = Urostachys obtusifolius (P. Beauv.) Herter ex Nessel; Bärlappgewächse 190 (1939) var. pachyphylla (Kuhn ex Herter) C. Chr.; Catalogue des Plantes de Madagascar, Pterid. 69 (1931) Distribution [complete]: Madagascar = Huperzia pachyphylla (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985) = Lycopodium pachyphyllum Kuhn ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 51. 1909 [Kuhn ex E. Pritzel, in Engler & Prantl., Natürl. Pflanzenfam. l(4): 599 (1901 [nom. inval.]) = Urostachys pachyphyllus (Herter) Herter ex Nessel; Bärlappgewächse 190 (1939) Huperzia ocanana (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Colombia Remarks: saururus agg. = Huperzia moritzii (Nessel) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) = Urostachys moritzii Herter ex Nessel; [Bärlappgewächse 70. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 1601 t. 9, f. 29 (1940) = Lycopodium ocananum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909) = Urostachys ocananus (Herter) Herter ex Nessel; Bärlappgewächse 101 (1939) = Huperzia arthurii (Herter) Holub; Folia Geobot. Phytotax. 20: 70 (1985) = Urostachys arthurii Maxon ex Herter; Revista Sudamer. Bot. 10: 114 (1953 [1954]) = Lycopodium arthurii (Herter) C. Morton; Amer. Fern J. 54: 72 (1964) = Huperzia trachyloma (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 157 (1988) = Lycopodium trachyloma (Herter) C. Morton; Amer. Fern J. 54: 73 (1964) = Urostachys trachyloma Maxon ex Herter; Revista Sudamer. Bot. 10: 113 (1953 [1954]) Huperzia occidentalis (Clute) Kartesz & Gandhi; Phytologia 70: 201 (1991) Distribution [complete]: Canada (SW-Yukon, W-British Columbia, Alberta), USA (Washington, Alberta, Montana, N-Idaho) = Huperzia occidentalis (Clute) Beitel; Amer. Fern Journal 82(2): 46 (1992) [nom. illeg. = Lycopodium lucidulum Michaux f. occidentale Clute; Fern Bull. 11: 13 (1903) = Lycopodium lucidulum var. occidentale (Clute) L. R. Wilson; Rhodora 34: 170 (1932) = Lycopodium selago var. occidentale (Clute) B. Boivin; Le Naturaliste Canadien 93: 359 (1966) Huperzia oceaniana (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: Vanuatu Remarks: unverified name, myrtifolia agg. = Lycopodium oceanianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 52 (1909) = Urostachys oceanianus (Herter) Herter ex Nessel; Bärlappgewächse 237 (1939) Huperzia oltmannsii (Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Society Isl. (Tahiti) Remarks: phlegmaria agg. = Urostachys oltmannsii Herter ex Nessel; [Bärlappgewächse 231. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 168, t. 14, f. 71 (1940) Huperzia ophioglossoides (Lam.) Rothm.; Feddes Repert. 54: 62 (1944) Distribution: Transvaal, Zimbabwe, Mozambique, Malawi, Bioko Isl. (Fernando Poo), Madagascar, Réunion, Mauritius, Comores, Seychelles, Cameroun, Uganda, Kenya, Tanzania, Sudan, Sao Tomé, Equatorial Guinea, Annobon Isl., Congo, Zaire, Ethiopia Remarks: ophioglossoides agg. = Lepidotis longifolia P. Beauv.; Prodr. aethéogam. 109 (1805) = Lycopodium longifolium (Beauv.) Sw.; Syn. Fil. 177 (1806) = Lycopodium ophioglossoides Lam.; Encyclopédie Méthodique, Botanique 3: 646 (1789) = Urostachys ophioglossoides (Lam.) Herter ex Nessel; Die Bärlappgewächse 238 (1939) Huperzia orgaosana (Herter ex Nessel) comb. ined. Distribution: Brazil Remarks: unverified name, reflexa agg. = Urostachys orgaosanus Herter ex Nessel; Arch. Bot. Sao Paulo 1: 380 (1927) Huperzia orizabae (L. Underw. & F. Lloyd) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Mexico (Chiapas, Veracruz, Oaxaca), Guatemala, Honduras, El Salvador Remarks: linifolia agg. = Lycopodium orizabae Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 110 (1906) = Urostachys orizabae (Underw. & Lloyd) Herter ex Nessel; Die Bärlappgewächse 175 (1939) Huperzia ovatifolia Ching; Acta Bot. Yunnanica 3: 298 (1981) Distribution: SE-Yunnan Remarks: unverified name, hamiltonii agg. = Phlegmariurus ovatifolius (Ching) W.M. Chu; Acta Phytotaxonomica Sinica 37(1): 52 (1999) Huperzia parksii (Copel.) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Fiji (Viti Levu) Remarks: verticillata agg. = Lycopodium parksii Copel.; Bernice P. Bishop. Mus. Bull. 59: 8, t. 2 (1929) = Urostachys parksii (Copel.) Herter ex Nessel; Bärlappgewächse 124 (1939) Huperzia parvifolia (Nessel) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988) Distribution: Brazil Remarks: reflexa agg. = Urostachys parvifolius Raddi ex Nessel; Arch. Bot. Sao Paulo 1: 395, t. 10, f. 2 (1927) Huperzia patentissima (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: New Guinea Remarks: unverified name, phlegmaria agg. = Lycopodium patentissimum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 4 (1917) = Urostachys patentissimus (Alderw.) Herter ex Nessel; Bärlappgewächse 226 (1939) Huperzia pearcei (Baker) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Bolivia, Colombia Remarks: reflexa agg. = Lycopodium pearcei Baker; Handbook of the Fern-Allies 14 (1887) = Urostachys pearcei (Baker) Herter ex Nessel; Bärlappgewächse 73 (1939) Huperzia pecten (Baker) Tard.; Adansonia 10: 18 (1970) Distribution [complete]: Madagascar Remarks: pecten agg. = Lycopodium pecten Baker; J. Linn. Soc. Bot. 15: 421 (1876) = Urostachys pecten (Baker) Herter ex Nessel; Bärlappgewächse 67 (1939) Huperzia perrieriana Tard.; Adansonia 10: 20, t. 2, fig. 14 (1970) Distribution [complete]: Madagascar Remarks: phlegmaria agg. Huperzia petiolata (C. B. Clarke) Dixit; J. Bombay Nat. Hist. Soc. 77 (3): 541 (1981) Distribution: India (Meghalaya, Khasia), Yunnan Remarks: hamiltonii agg. = Lycopodium hamiltonii Sprengel var. petiolatum C. B. Clarke; Trans. Linn. Soc. London, ser. 2; Bot. 1: 590 (1880 (Aug.)) = Lycopodium petiolatum (C. B. Clarke) Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 36. 1909, by reference to Baker (Handb. fern-allies 10 (1887)) = Lycopodium petiolatum (C.B. Clarke) Christ; Bulletin de l'Académie Internationale de Géographie, Botanique 11: 271., H. Christ? (1902) = Phlegmariurus hamiltonii var. petiolatus (C.B. Clarke) Ching; Acta Botanica Yunnanica 4(2): 126-127 (1982) = Phlegmariurus petiolatus (C.B. Clarke) H.S. Kung & L.B. Zhang; Acta Phytotaxonomica Sinica 37(1): 45-47 (1999) = Urostachys hamiltonii var. petiolatus (C.B. Clarke) Herter ex Nessel; Die Bärlappgewächse 68 (1939) = Urostachys petiolatus (C.B. Clarke) Herter ex Nessel; Die Bärlappgewächse 196, see Engl. Bot Jahrb. 34, 98 (1909). 9to check basinym, Nessel lists as Baker, Handb (1887)0 Huperzia petrovii Siplivinsky; Novosti Sist. Vyssh. Rast. 10: 346 (1973) Distribution: Russia, W-Siberia, C-Siberia, E-Siberia / Amur / Ussuri Remarks: selago agg. = Lycopodium appressum (Desv.) V. Petrov; Flora Iakutiae 37 (1930) excl. basionymo & pl. arct. [non Lloyd & Underw. p. p. = Lycopodium selago L. ssp. appressum (Desv.) Hult.; [p. p. Huperzia pflanzii (Nessel) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100) 156 (1988) Distribution [complete]: Costa Rica, Bolivia Remarks: brevifolia agg. = Lycopodium pflanzii (Nessel) Lellinger; Proc. Biol. Soc. Wash. 89 (61): 717 (1977) = Urostachys pflanzii Nessel; Repert. Spec. Nov. Regn. Veg. 36(12-15): 181, t. 171 (1934) Huperzia phlegmaria (L.) Rothm.; Feddes Repert. 54: 62 (1944) Distribution: India (West Bengal, Sikkim, Assam, Arunachal Pradesh, Kerala), Andamans, Nicobars, peninsular Malaysia, Borneo (Mt. Kinabalu, etc.), Malesia, Sri Lanka, Taiwan, Japan, Ryukyu Isl., China, Queensland, Christmas Isl. (Austr.), Sumatra, Thailand, Philippines, Palawan, Palau Isl. (Babeldoab, Koror, Malakal), Tonga (Late Isl., 'Eua), Western Samoa (Aleipata Isl., Savaii, Upolu), Niue, Cook Isl. (Rarotonga), American Samoa (Manua Isl., Tutuila, Ta'u), Society Isl. (Tahiti), Marquesas Isl. (Nuku Hiva, Fatu Hiva), Principe Isl., Sao Tomé, Uganda, Tanzania, Malawi, Guinea, Liberia, Cameroun, Equatorial Guinea, Principe Isl., Gabon, ?Zaire, Bioko Isl. (Fernando Poo) Remarks: phlegmaria agg. = ?Lycopodium phlegmaria var. marchionicum E. Br. (1931) = Huperzia edanoi (Copel.) Holub; Folia Geobot. Phytotax. 20: 72 (1985) = Huperzia harmsii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985) = Lepidotis phlegmaria (L.) P. Beauv.; Prodr. aethéogam. 109 (1805) = Lycopodium edanoi Copel.; Philipp. J. Sci. 46: 209 (1931) = Lycopodium filiforme Roxb.; Calcutta J. Nat. Hist. 4: 473. 1844 [non Sw. 1800 = Lycopodium phlegmaria L.; Sp. pl. 1101 (1753) = Lycopodium phlegmaria var. longibracteatum Domin (1913) = Phlegmariurus phlegmaria (L.) Holub; Preslia 36(1): 21 (1964) = Phlegmariurus phlegmaria (L.) Sen & Sen; Fern Gaz. 11 (6): 421. f. 4 a (1978) = Urostachys edanoi (Copel.) Nessel; Bärlappgewächse 230 (1939) = Urostachys harmsii Herter ex Nessel; [Bärlappgewächse 223. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 166 (1940) = Urostachys phlegmaria (L.) Herter ex Nessel; Die Bärlappgewächse 215 (1939) var. divaricatum Blume; Enum. pl. Javae 262 (1828) Distribution: Java = Lycopodium divergens Alderw.; Malayan fern allies 45 (1915) var. tardieuae (Herter) Tardieu; Adansonia ser. 2 10: 18 (1970) Distribution [complete]: Madagascar = Urostachys tardieuae Herter; Notulae Systematicae. Herbier du Museum de Paris 15: 355 (1958) = Huperzia tardieuae (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) var. longifolium Spring Distribution: Southern Marianas (Rota Isl., Guam), Palau Isl. (Babeldaob, Aulupse'el), Micronesia (Nukuoro, Pohnpei, Kosrae), New Caledonia, Vietnam, Austral Isl. (Tubuai) Conservation status: IUCN E = Huperzia mirabilis (Willd.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) = Lycopodium mirabile Willd.; Sp. pl. 5: 11 (1810) = Urostachys mirabilis (Willd.) Herter ex Nessel; Bärlappgewächse 225 (1939) = Lycopodium phlegmaria var. mirabile (Willd.) R. Fosb. Huperzia phlegmarioides (Gaudich.) Rothm.; Feddes Repert. 54: 62 (1944) Distribution: NE-Queensland, New Caledonia, Moluccas, Fiji (Viti Levu, Vanua Levu, Ovalau), Society Isl. (Tahiti), Western Samoa (Upolu), Austral Isl. (Raivavae) Remarks: phlegmarioides agg. = Huperzia kandavuensis (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985) = Huperzia pseudophlegmaria (Kuhn) Holub; Folia Geobot. Phytotax. 20: 76 (1985) = Lycopodium phlegmarioides Gaudich.; Voy. Uranie 1: 281, t. 23 (1828 (16 Aug)) = Lycopodium pseudophlegmaria Kuhn; Forschungsr. Gazelle 4: 16 (1889) = Lycopodium tetrapterygium (Herter) F.M. Bailey; Proc. Roy. Soc. Queensland 1: 150. 1884 = Urostachys kandavuensis Nessel; Repert. Spec. Nov. Regni Veg. 39: 67, t. 192a (1935) = Urostachys phlegmarioides (Gaudich.) Herter ex Nessel; Bärlappgewächse 247 (1939) = Urostachys pseudophlegmaria (Kuhn) Herter ex Nessel; Bärlappgewächse 237 (1939) = Urostachys tetrapterygius (Bailey) Herter; Index Lyc. 86 (1949) Huperzia phylicifolia (Desv. ex Poir.) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution [complete]: Ecuador, Chile, Galapagos, Peru, Bolivia, N-Argentina, SE-Brazil, Colombia, Venezuela Remarks: myrsinites agg. = Lycopodium nubigenum Herzog; Meded. Rijks-Herb. 27: 2 (1915) = Urostachys nubigenus (Herzog) Herter ex Nessel; Bärlappgewächse 246 (1939) = Lycopodium congestifolium Spring; Bull. Acad. Roy. Sci. Belg. 8: 518 (1841) = Lycopodium phylicifolium Desv. ex Poiret; Encycl. suppl. 3: 546 (1813 [1814] ("phylicaefolium")) = Lycopodium subulatum var. pastoense Baker; Handbook of the Fern-Allies 21 (1887) = Urostachys phylicifolius (Poiret) Herter ex Nessel; Bärlappgewächse 246 (1939) Huperzia phyllantha (Hook. & Arn.) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: India (Uttar Pradesh, Arunachal Pradesh, West Bengal, Sikkim, Assam, Manipur, Meghalaya, Kerala, Tamil Nadu), Nicobars, Andamans, Sri Lanka, trop. America, Cape Prov., Mauritius, Réunion, peninsular Malaysia (Penang Isl., Kelantan, Perak, Negeri Sembilan, Singapore), Borneo (Mt. Kinabalu, etc.), Hawaii (Kauai, Oahu, Molokai, Maui, Lanai, Hawaii Isl.), Fiji (Viti Levu), Micronesia (Pohnpei), New Caledonia, Western Samoa (Savaii) Remarks: phyllantha agg. = Lycopodium phyllanthum Hook. & Arn.; Bot. Beechey Voy. 102 (1832 (Oct-Nov.)) = Phlegmariurus phyllanthus (Hook. & Arn.) Dixit; J. Bombay Nat. Hist. Soc. 77 (3): 541 (1980 [1981]) = Urostachys phyllanthum (Hook. & Arn.) Herter; Philipp. J. Sci. 22: 181. 1923 = Huperzia parvifolia (Raddi ex Nessel) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21(100): 156 (1988) = Huperzia parvifolia (Raddi) Rothm.; Feddes Repert. 54: 60 (1944) [nom. inval. = Lycopodium parvifolium "Raddi" (Herter 1909a: 34, Rolleri 1981: 81) [nom. inval. = Lycopodium pachystachyon Spring; Bull. Acad. Roy. Sci. Belg. 8: 518. 1841 [non Lycopodium pachystachyum Desv. ex Poiret (1814) = Lycopodium parvifolium Raddi ex Rolleri; Notas Mus. La Plata, Bot. 13(71): 81 (1981), nom. inval. = Urostachys pachystachyon (Spring) Herter ex Nessel; Bärlappgewächse 211 (1939) = Urostachys pachystachyon (Spring) Rothm.; Feddes Repert. 54: 62 (1944) = Urostachys parvifolius Raddi ex Nessel; Archivos de Botânica do São Paulo 1: 395, t. 10, f. 2 (1927) Huperzia picardae (H. Christ ex Krug) Holub; Folia Geobot. Phytotax. 20: 75 (1985) Distribution: Hispaniola (Haiti) Remarks: taxifolia agg. = Lycopodium picardae Christ ex Krug; Urban; Bot. Jahrb. Syst. 24: 148 (1897) = Urostachys picardae (Krug) Herter; Repert. Spec. Nov. Regni Veg. 19: 166 (1923) Huperzia pichiana Tard.; Adansonia 10: 21, t. 2, fig. 5-8 (1970) Distribution [complete]: Madagascar Remarks: verticillata agg. Huperzia pinifolia Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874) [nom. nov. Distribution: Borneo (Mt. Kinabalu, etc.) Remarks: hippuris agg. = Lycopodium pinifolium Blume; Enum. pl. javae 2: 264. 1828 [non Kaulf (1824) = Urostachys pinifolius (Blume) Herter; Bot. Arch. 3: 16 (1923) Huperzia piscium (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution [complete]: Thailand, Java = Lycopodium piscium (Herter) Tag. & Iwatsuki; Acta Phytotax. Geobot. 22: 103 (1967) = Urostachys piscium Herter; Index Lyc. 75 (1949) Huperzia pithyoides (Schldl. & Cham.) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Costa Rica, Mexico (Chiapas, Jalisco, Oaxaca, Veracruz, San Luis Potosi, Tamaulipas), Guatemala, Honduras, El Salvador, Nicaragua, Colombia, Venezuela, Cuba, Hispaniola Remarks: wilsonii agg. = Lycopodium gigas Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 50 (1909) = Lycopodium mandiocanum Raddi var. mexicanum Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 45 (1842) = Lycopodium pithyoides Schlecht. & Cham.; Linnaea 5: 623 (1830) = Urostachys gigas (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923) = Urostachys pithyoides (Schlecht. & Cham.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923) Huperzia pittieri (H. Christ) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Isla del Coco Remarks: linifolia agg. = Lycopodium pittieri H. Christ; Bulletin de la Société Botanique de Genève 1(5): 236 (1909) = Urostachys pittieri (H. Christ) Herter; Estudios Botánicos en la Región Uruguaya 20: 75 (1949) Huperzia polaris (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Antipodes Isl. Remarks: varia agg. = Urostachys varius (R. Br.) Nessel var. polaris Herter ex Nessel; Revista Sudamer. Bot. 6: 165, t. 12, fig. 59. 1940 [non Lycopodium varium R. Br. var. polaris Kirk (18849) = Urostachys polaris (Nessel) Herter; Index Lyc. 76 (1949) Huperzia polycarpos (Kunze) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Costa Rica, Panama, Colombia, Ecuador, Peru Remarks: dichotoma agg. = Lycopodium cuatrecasasii (Herter) C. V. Morton; Amer. Fern Journal 54: 72 (1964) = Lycopodium polycarpum Kunze; Linnaea 9: 5., orth. var (1834) = Urostachys cuatrecasasii Herter; Revista Sudamericana de Botánica 10: 123 (1953) = Urostachys polycarpos (Kunze) Herter; Estudios Botánicos en la Región Uruguaya 20: 76 (1949) Huperzia polyclada (Sodiro) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988) Distribution [complete]: Ecuador Remarks: saururus agg. = Lycopodium polycladum Sodiro; Crypt. vasc. quit. 561. 1893 [Anales Univ. Quito 12 (81): 316 (1895]) = Urostachys polycladus (Sodiro) Herter ex Nessel; Bärlappgewächse 73 (1939) Huperzia polydactyla B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 64, f. 11B (1988) Distribution [complete]: Ecuador Remarks: brevifolia agg. Huperzia polylepidetorum B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 42, f. 6B (1988) Distribution [complete]: Ecuador, Peru Remarks: saururus agg. Huperzia porophila (F.W. Lloyd & L. Underwood) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: USA (Alabama, Illinois, Indiana, Iowa, Kentucky, Minnesota, Missouri, Ohio, Pennsylvania, Tennessee, Virginia, West Virginia, Wisconsin) Trivial names: Rock Club-Moss Remarks: selago agg. = Huperzia selago (L.) Bernh. ex Schrank & Mart. ssp. lucidula var. porophila (Lloyd & Underw.) Love & Love = Lycopodium lucidulum Michx. var. porophilum (Lloyd & Underwood) Clute; The Fern Allies 262 (1905) = Lycopodium porophilum Lloyd & Underw.; Bull. Torrey Bot. Club 27(4): 150 (1900) = Lycopodium selago L. var. porophila (Lloyd & Underwood) Clute = Urostachys lucidulus (Michx.) Herter var. porophilus (Lloyd & Underwood) Nessel, non Lam. = Urostachys porophilus (Lloyd & Underwood) Herter Huperzia pringlei (Underw. & F.E. Lloyd) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Mexico (Chiapas, Sinaloa, Jalisco, Michoacan, Mexico State, Morelos, Oaxaca, Veracruz, Hidalgo), Guatemala, Honduras, El Salvador Remarks: verticillata agg. = Lycopodium fontinaloides var. mexicanum Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 49 (1842) = Lycopodium pringlei Underw. & F.E. Lloyd; Bulletin of the Torrey Botanical Club 33: 109 (1906) = Urostachys pringlei (Underw. & Lloyd) Herter ex Nessel; Die Bärlappgewächse 131 (1939) Huperzia prolifera (Blume) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution: Java, trop. Asia, NE-Queensland Remarks: hippuris agg. = Lycopodium proliferum Blume; Enum. pl. Javae 2: 265 (1828) = Urostachys prolifer (Blume) Herter; Index Lyc. 77 (1949) Huperzia pruinosa (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Peru, Colombia Remarks: phlegmaria agg. Conservation status: IUCN R = Lycopodium pruinosum Hieron. et Herter ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 52 (1909) = Urostachys pruinosus (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 420 (1927) = Huperzia durissima (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985) = Lycopodium durissimum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 52 (1909) = Urostachys durissimum (Herter) Herter ex Nessel; Bärlappgewächse 240 (1939) Huperzia pseudovaria (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution [complete]: New Caledonia Remarks: myrtifolia agg. Conservation status: IUCN R = Lycopodium pseudovarium Brownlie; Ptéridophytes, in Aubréville, Fl. Nouv.-Calédonie 3.- 23, t. 1, f. 13-15 (1969) Huperzia pulcherrima (Hook.& Grev.) Pichi-Serm.; Webbia 24: 719 (1970) Distribution: India (NW-Himalayas, E-India), Sri Lanka, Nepal, Bhutan, China, Taiwan, Japan Remarks: hippuris agg. = Huperzia pulcherrima (Wall. ex Hook. & Grev.) Sen & Sen; Fern Gaz. 11 (6): 419. f 2 i-r (1978) = Huperzia setacea Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) = Huperzia taiwanensis (Ching) Holub; Folia Geobot. Phytotax. 20(1): 77 (1985) = Lycopodium pulcherrimum Wall. ex Hook. & Grev.; Icones Filicum 1(2): t. 38 (1827) = Lycopodium setaceum Buch.-Ham. ex D. Don; Prodromus Florae Nepalensis 18 (1825) [nom. illeg., non Lycopodium setaceum Lam. (1789) = Phlegmariurus pulcherrimus (Hook. & Grev.) Löve & Löve; Taxon 26: 324 (1977) = Phlegmariurus taiwanensis Ching; Acta Bot. Yunnanica 4: 124 (1982 [nom. superfl. for Lycopodium pulcherrimum Hook. & Grev. by reference to Hayata (1914: 132)) = Urostachys amphorae Herter; Index Lyc. 51 (1949 [nom. superfl. for Urostachys pulcherrimus (Hook. & Grev.) Nessel) = Urostachys pulcherrimus (Hook. & Grev.) Herter ex Nessel; Bärlappgewächse 72 (1939) = Urostachys setaceus (Hamilt. ex D. Don) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 408. 1927 [Bärlappgewächse 141 (19399) Huperzia pungentifolia (Silveira) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution: Brazil (Minas Gerais, Santa Catarina), Haiti Remarks: brongniartii agg. = Urostachys hennebergorum Nessel; Repert. Sp. Nov. Regni Veg. 48: 169, t. 316, f. 1 (1940) = Lycopodium ouropretanum Christ; Schwacke, Pl. Nov. Mineiras 2: 39, t. 4. 1900 [sens. Bull. Herb. Boissier, ser. 2, 2: 702 (1902]) = Lycopodium pungentifolium Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2, 5: 119, t. 4 (1898) = Urostachys pungentifolius (Silveira) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 394 (1927) Huperzia quadrifariata (Bory) Rothm.; Feddes Repert. 54: 60 (1944) Distribution: Brazil (Santa Catarina, Rio de Janeiro) Remarks: unverified name, quadrifariata agg. = Lycopodium quadrangulare Spring, in. C. Martius; Fl. Bras. l(2): 112-113, t. 5, f. 1 (1840) = Urostachys quadrangularis (Spring) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 404. 1927 = Lycopodium quadrifariatum Bory; Duperrey, Voy. monde; Bot. 1: 245 (1829 (Aug.)) = Urostachys quadrifariatus (Bory) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 405 (1927) = Huperzia aschersonii (Herter) Holub; Folia Geobot. Phytotax. 20: 70 (1985) = Lycopodium aschersonii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 53 (1909) = Urostachys aschersonii (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1. 422 (1927) Huperzia quasipolytrichoides (Hayata) Ching; Acta Botanica Yunnanica 3(3): 299 (1981) Distribution: Taiwan Remarks: selago agg. = Huperzia reflexo-integra (Hayata) Holub; Folia Geobot. Phytotax. 20: 76 (1985) = Lycopodium quasipolytrichoides Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 5: 252-254, f. 89 (1915) = Lycopodium reflexo-integrum Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 5: 254-255, f. 90 (1915) = Urostachys quasipolytrichoides (Hayata) Herter ex Nessel; Die Bärlappgewächse 64-65, Abb. 8, f. 37 (1939) = Urostachys reflexo-integer (Hayata) Herter ex Nessel; Bärlappgewächse 64 (1939) var. rectifolia (J.F. Cheng) H.S. Kung & L.B. Zhang; Acta Phytotaxonomica Sinica 36(6): 528 (1998) Distribution: Distribution unknown = Huperzia whangshanensis var. rectifolia J.F. Cheng; Fl. Jiangxi 1: 27, 505, f. 12 (1993) Huperzia recurvifolia Rolleri; Notas del Museo de la Plata, Botánica 21 (10, f. 1-33): 210 (1989) Distribution [complete]: Venezuela, Brazil (Bahia), British Guyana Huperzia reflexa (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Puerto Rico, Costa Rica, Galapagos, Cuba, Hispaniola, Jamaica, Mexico (Jalisco, Guerrero, Mexico State, Oaxaca, Veracruz, Puebla, Hidalgo, Chiapas), Guatemala, Honduras, El Salvador, Paraguay, Bolivia, Nicaragua, Colombia, Venezuela, Ecuador, Galapagos, Peru, Brazil, Martinique, Guadeloupe, Dominica, St. Vincent, British Guyana (Mt. Roraima) Remarks: reflexa agg. = Huperzia bifida (Humb. & Bonpl. ex Willd.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) = Huperzia densifolia (Baker) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988) = Huperzia mexiae (Copel.) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988) = Huperzia reflexa (Lam.) Rothm.; Repert. Spec. Nov. Regn. Veg. 54: 60., hom. illeg. superfl (1944) = Huperzia reflexa var. bifida (Humb. & Bonpl. ex Willd.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) = Lycopodium bifidum Humb. & Bonpl. ex Willd.; Sp. Pl. Editio quarta 5: 53 (1810) = Lycopodium densifolium (Baker) Underw. & F.E. Lloyd; Bulletin of the Torrey Botanical Club 33: 106 (1906) = Lycopodium mexiae Copel.; Univ. California Publ. Bot. 19: 294, t. 47 (1941) = Lycopodium reflexum Lam.; Encyclopédie Méthodique, Botanique 3: 653 (1792) = Lycopodium reflexum var. caesium Vareschi; Flora de Venezuela 1(1): 31 (1968), nom. inval. (no type indicated) = Lycopodium reflexum var. densifolium Baker; Handbook of the Fern-Allies 11 (1887) = Lycopodium reflexum var. majus Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 26 (1842) = Lycopodium reversum C. Presl; Reliquiae Haenkeanae 1(1): 82 (1825) = Lycopodium rigidum Sw.; J. Bot. (Schrader) 1800 (2): 114. 1801 [Syn. fil. 176, 400. 1806] [non J. Gmelin (1791) = Lycopodium squarrosum Sw.; Prodr. 137. 1788 [non G. Forster 1786, nec Larn (1792) = Lycopodium stellae-polaris (Herter) C.V. Morton; Amer. Fern Journal 54: 72 (1964) = Plananthus reflexus (Lam.) P. Beauv.; Prodr. Aetheog. 100 (1805) = Plananthus squarrosus (Sw., Lam., J. Gmelin) P. Beauv.; Prod. aethéogam. 112 (1805) [nom. confus., combination based on different basionyms = Urostachys bifidus (Humb. & Bonpl. ex Willd.) Herter ex Nessel; Die Bärlappgewächse 110, t. 24, f. 4 (1939) = Urostachys densifolius (Baker) Herter; Index Lyc. 58. 1949 = Urostachys jergii Nessel; Repert. Spec. Nov. Regn. Veg. 39: 70, t. 194a (1935) = Urostachys reflexus (Lam.) Herter; Beihefte zum Botanischen Centralblatt 39: 249 (1922) = Urostachys reflexus var. densifolius (Baker) Herter ex Nessel & Hoehne; Archivos de Botânica do São Paulo 1(4): 394 (1927) = Urostachys reversus (C. Presl) Herter; Estudios Botánicos en la Región Uruguaya 20: 78 (1949), Alt. title: Index Lycopodiarum 78 (1949) = Urostachys stellae-polaris Herter; Revista Sudamericana de Botánica 10: 121 (1954) var. minor (Spring) B. Øllg.; Flora of Ecuador 33: 26 (1988) Distribution [complete]: Honduras, Costa Rica, Ecuador, Peru, Bolivia, Brazil, Trinidad = Lycopodium brutum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 47 (1909) = Lycopodium reflexum var. minus Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 26 (1842) = Urostachys brutus (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923) = Urostachys reflexus (Lam.) Herter var. brutus (Herter) Herter ex Nessel; Die Bärlappgewächse 116 (1939) = Urostachys reflexus var. minor (Spring) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 393 (1927) Huperzia regnellii (Maxon) B. Øllg. & Wind.; Bradea 5: 16 (1987) Distribution: Brazil (Minas Gerais) Remarks: brongniartii agg. Conservation status: IUCN R = Lycopodium regnellii Maxon; Contr. U. S. Natl. Herb. 17 (4): 424, t. 23 (1914) = Urostachys regnellii (Maxon) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 381 (1927) Huperzia ribourtii (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Society Isl. (Tahiti), Cook Isl. (Rarotonga) Remarks: obtusifolia agg. = Lycopodium ribourtii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 53 (1909) = Urostachys ribourtii (Herter) Herter ex Nessel; Bärlappgewächse 250 (1939) Huperzia rigida (J. Gmelin) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Hispaniola, Jamaica, Guadeloupe, Martinique, St. Vincent Remarks: reflexa agg. = Lycopodium rigidum J. Gmelin; Syst. nat. II [Syst. Veg.1 (2): 128] (1791) = Plananthus rigidus (J. Gmelin) P. Beauv.; Prod. aethéogam. 111 (1805) = Urostachys rigidus (J. Gmelin) Herter; Repert. Spec. Nov. Regni Veg. 19: 163 (1923) Huperzia riobambensis (Herter) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Colombia Remarks: reflexa agg. = Urostachys affinis (Grev. & Hook.) Nessel var. riobambensis Nessel; [Bärlappgewächse 98 (1939] (Revista Sudamericana de Botánica 6(5-6): 162, t. 10, f. 38 (1940) [nom. inval. = Urostachys riobambensis (Nessel) Herter; Index Lyc. 79 (1949) = Lycopodium castoris (Herter) C. Morton; Amer. Fern J. 54: 72 (1964) = Urostachys castoris Herter; Revista Sudamer. Bot. 10: 111 (1953 [1954]) Huperzia robusta (Klotzsch) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Venezuela Remarks: phlegmaria agg. = Huperzia phelpsii (Vareschi) Holub; Folia Geobot. Phytotax. 20: 75 (1985) = Lycopodium phelpsii Vareschi; Acta Botanica Venezuelica 1: 84 (1966) = Lycopodium robustum Klotzsch; Linnaea 18: 518 (1844) = Urostachys robustus (Klotzsch) Herter ex Nessel; Bärlappgewächse 212 (1939) Huperzia rosenstockiana (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution [complete]: Ecuador, Peru, Colombia Remarks: taxifolia agg. = Lycopodium cassandrae (Herter) C. Morton; Amer. Fern J. 54: 72 (1964) = Lycopodium rosenstockianum Herter; Hedwigia 49: 90 (1909) = Urostachys cassandrae Herter; Revista Sudamer. Bot. 10: 116 (1953 [1954]) = Urostachys rosenstockianus (Herter) Herter ex Nessel; Bärlappgewächse 174 (1939) Huperzia rostrifolia (Silveira) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution: Brazil (Minas Gerais) Remarks: unverified name, reflexa agg. = Lycopodium rostrifolium Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2, 5: 118, t. 2 (1898) = Urostachys rostrifolius (Silveira) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 379 (1927) Huperzia rubicaulis S.K. Wu & X. Cheng; Acta Phytotaxonomica Sinica 23(5): 400, f. 1 (1985) Distribution: Qinghai, Tibet, Yunnan Remarks: selago agg. Huperzia rubra (Cham.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874) Distribution: Brazil Remarks: unverified name, brongniartii agg. = Lycopodium rubrum Cham.; Linnaea 8: 389 (1833) = Urostachys ruber (Cham.) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 381 (1927) Huperzia rubrica (C. Chr.) Tardieu; Adansonia ser. 2 10: 18 (1970) Distribution [complete]: Madagascar Remarks: ophioglossoides agg. = Lycopodium ophioglossoides Lam. var. rubricaule C. Chr.; Dansk Bot. Ark. 7: 192, t. 77, f. 9-14 (1932) = Urostachys rubricus Herter; Revista Sudamer. Bot. 8: 25 (1949) Huperzia rubricaulis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution: New Guinea Remarks: unverified name, selago agg. = Lycopodium rubricaule Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 3 (1917) = Urostachys rubricaulis (Alderw.) Herter ex Nessel; Bärlappgewächse 44 (1939) Huperzia rufescens (Hook.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874) Distribution [complete]: Ecuador, Colombia Remarks: brevifolia agg. = Lycopodium rufescens Hook.; Icones Plantarum 1: tab 36 (1837) = Urostachys rufescens (Hook.) Nessel; Die Bärlappgewächse 103 (1939) Huperzia rupicola (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution: Brazil Remarks: unverified name, squarrosa agg. = Lycopodium rupicolum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 16: 39 (1914) = Urostachys rupicolus (Alderw.) Herter ex Nessel; Bärlappgewächse 203 (1939) Huperzia sagasteguiana B. Øllg.; Fieldiana, Bot., New Ser. 34: 37 (1994) Distribution [complete]: Ecuador, Peru, Colombia Huperzia salvinioides (Herter) Holub; Folia Geobot. Phytotax. 20. 76 (1985) Distribution [complete]: Philippines, Taiwan, Japan Remarks: nummulariifolia agg. = Lycopodium salvinioides (Herter) Tag.; Acta Phytotax. Geobot. 8: 20 (1939) = Phlegmariurus salvinioides (Herter) Ching; Acta Bot. Yunnanica 4: 122 (1982) = Urostachys formosanus Herter ex Nessel; [Bärlappgewächse 229. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 167, excl. fig. 1940 = Urostachys formosanus Herter ex Nessel; Die Bärlappgewächse 229 (1939) = Urostachys salvinioides Herter; Bot. Arch. 3. 18 (1923) Huperzia samoana (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution: Western Samoa Remarks: hippuris agg. = Urostachys samoanus Herter ex Nessel; [Bärlappgewächse 125, f. 29e 5. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 163 (1940) Huperzia sarmentosa (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Ecuador, Peru Remarks: verticillata agg. = Lycopodium sarmentosum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 13 (1849) = Urostachys sarmentosus (Spring) Herter; Repert. Spec. Nov. Regni veg. 19: 165 (1923) Huperzia saururoides (Bory & d'Urv.) Rothm.; Feddes Repert. 54: 59 (1951) Distribution [complete]: Falkland Isl. Remarks: selago agg. = Lycopodium selago L. var. saururoides Bory & D'Urville ex Baker; Handb. fern-allies 9. 1887 [Bory in D'Urville, Mém. Soc. Linn. Paris 4: 597, pl. 4 (1825 [1826]) [nom. inval. = Urostachys saururoides (Bory et D'Urville ex Baker) Herter ex Nessel; Bärlappgewächse 30 (1939) Huperzia saururus (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874) Distribution: Marion Isl., Prince Edward Isl., Tristan da Cunha, Gough Isl., Iles Crozet, Iles Kerguelen, New Amsterdam Isl., Réunion, Madagascar, Comores, Mauritius, Rodrigues, St. Helena, Ascension Isl., Nightingale Isl., Natal, Cape Prov., Lesotho, Zimbabwe, Malawi, Uganda, Kenya, Tanzania, Ethiopia, Cameroun, Zaire, Rwanda, Argentina (Jujuy, Buenos Aires, Catamarca, Cordoba, Salta, San Luis Prov., Tucuman), Peru, Bolivia, Uruguay, Colombia, ?Ecuador Conservation status: IUCN I, Ex/E (Ascension), V (St. Helena), I (Tristan da Cunha) Remarks: saururus agg. = ?Lycopodium breve Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909) = ?Urostachys brevis (Herter) Herter ex Nessel; Bärlappgewächse 82 (1939) = Huperzia axillaris (Roxb.) Rothm.; Feddes Repert. 54: 60 (1944) = Huperzia sanctae-barbarae (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988) = Lycopodium axillare Roxb.; Beatson, Tracts St. Helena 312 (1816) = Lycopodium elongatum Lagasca ex Sw.; Syn. fil. 175 (1806) = Lycopodium sanctae-barbarae Rolleri; Darwiniana 16: 129, f. l(D-E)o t. 11 t. 3(B) (1970) = Lycopodium saururus Lam.; Encycl. 3: 653 (1789 [1792]) = Plananthus saururus (Lam.) P. Beauv.; Prod. aethéogam. 111 (1805) = Selago indica Dill. (Herter 1949a: 49) [nom. inval. (polynom.) = Urostachys axillaris (Roxb.) Herter ex Nessel; Bärlappgewächse 82 (1939) = Urostachys elongatus (Sw.) Herter; Estudios Botánicos en la Región Uruguaya 20: 60 (1949) = Urostachys saururus (Lam.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923) Huperzia scabrida B. Øllg.; Harling & Andersson, Fl. Ecuador 33. 60, f. 8A (1988) Distribution [complete]: Ecuador Remarks: saururus agg. Huperzia schlimii (Herter) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Colombia Remarks: saururus agg. = Lycopodium schlimii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909) = Urostachys schlimii (Herter) Herter ex Nessel; Bärlappgewächse 91 (1939) Huperzia schmidtchenii (Hieron.) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution: South America Remarks: unverified name, brevifolia agg. = Lycopodium schmidtchenii Hieron.; Bot. Jahrb. Syst. 34: 570 (1905) = Urostachys schmidtchenii (Hieron.) Herter ex Nessel; Bärlappgewächse 103 (1939) Huperzia selago (L.) Bernhardi ex Schrank & C. F. P. Martius; Hort. Reg. Monac.: 3 (1829) ssp. selago (L.) Bernhardi ex Schrank & C. F. P. Martius; Hort. Reg. Monac.: 3 (1829) Trivial names: Tannenbärlapp, Northern Firmoss, Fir Clubmoss Distribution: Albania, Austria, Azores, Belgium, England, Bulgaria, former Czechoslovakia, Denmark, Faroer Isl., Finland, France, Germany, Ireland, Switzerland, Netherlands, Spain, Hungary, Italy, Iceland, former Yugoslavia, Malta, Norway, Poland, Romania, Sweden, Jan Mayen, asiatic Turkey (Anatolia), Pakistan, Gilgit, Kashmir, Baltic States, C-European Russia, E-European Russia, N-European Russia, W-European Russia, Belarus, Ukraine, Caucasus/Transcaucasus, W-Siberia, C-Siberia, E-Siberia / Amur / Ussuri, C-Asia, Taiwan, Japan, Yakushima Isl., Sikkim, NW-India, NE-India, Nepal, St. Pierre & Miquelon, Canada (Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, Labrador, N.W.Territories, Nova Scotia, Ontario, Prince Edward Isl., Quebec, Saskatchewan, Yukon, Nunavut), USA (Connecticut, Massachusetts, Maine, Michigan, Minnesota, New Hampshire, New York, Ohio, Rhode Island, Vermont, Wisconsin), Victoria, Tasmania, mountains of C-Brazil, Réunion Remarks: selago agg. = Huperzia dixitii Mandal & S. Ghosh; [fide Fraser-Jenkins (1997) = Huperzia selago (L.) Bernh.; Journal für die Botanik 2: 121-35 (1801) = Lycopodium abietiforme S. F. Gray; Brit. Pl. 2: 22 (1821) = Lycopodium densum Lam.; Fl. franc. 1 (Meth. anal.): (33) (1779) = Lycopodium recurvum Kitaibel ex Willd.; Sp. pl. 5: 50 (1810) = Lycopodium selago L.; Sp. Pl.: 1102 (1753) = Mirmau selago (L.) H. P. Fuchs; Verh. Naturf. Ges. Basel 66: 43-44 (1955) = Plananthus fastigiatus Opiz; Seznam 202. 1852, et Böh. Gew. 114 (1823) (Adelbert MS) [nom. superfl. = Plananthus recurvus (Kitaibel ex Willd.) Opiz; Seznam 202 (1852) = Plananthus selago (L.) P. Beauv.; Prod. aetheogam. 112 (1805) = Selago vulgaris Dill. (Herter 1949a: 49) [nom. inval. = Selago vulgaris Schur; Enum pl. Transsilv. 825 (1866) = Urostachys selago (L.) Herter; Botanisches Centralblatt 39(2): 249 (1922) [nom. illegit. var. sulcata Ching; Acta Botanica Yunnanica 3(3): 303 (1981) Distribution: Distribution unknown var. patens (Beauv.) Trevisan Distribution: Japan, North America = Lycopodium selago ssp. patens (Beauv.) Calder & Taylor = Lycopodium selago var. patens (Beauv.) Desv. = Plananthus patens P. Beauv. (1805) ssp. arctica (Tolm.) A. & D. Löve; Bot. Not. (Lund) 114: 35 (1961) Distribution [complete]: Spitsbergen, N-European Russia, C-Siberia, E-Siberia / Amur / Ussuri, Japan, Yakutia, S-Kuril Isl., Caucasus = Huperzia appressa (Desv.) A. Löve & D. Löve; Bot. Not. 114: 34 (1961) = Huperzia arctica (Tolm.) Sipliv.; Novosti Sist. Vyssh. Rast. 10: 347 (1973) = Huperzia selago f. reducta H. Christ; Bolletino della Società Botanica Italiana 1898: 184 (1898) = Huperzia selago f. reductum-angustinum H. Christ; Bolletino della Società Botanica Italiana 1898: 184 (1898) = Huperzia selago ssp. appressum (Desv.) D. Löve = Huperzia selago var. appressa (Desv.) Ching; Acta Botanica Yunnanica 3(3): 303 (1981) = Huperzia selago var. densa Trevisan = Lycopodium appressum (Desv.) V. Petrov; Flora Iakutiae 37. [p. p. excl. basionymo & pl., non arct., non Lloyd & Underw (1930) = Lycopodium arcticum (Tolm.) Sipl. = Lycopodium arcticum Grossh.; [nom. invalid. = Lycopodium selago L. ssp. appressum (Desv.) Hult.; [p. p. = Lycopodium selago L. ssp. arcticum Grossh. ex Tolm.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 20: 39, f. 1-2 (1960) = Lycopodium selago L. var. appressa Desv.; Mem. Soc. Linn. Paris 6: 180 (1827) = Lycopodium selago ssp. appressum (Desv.) Hultén = Urostachys selago f. reductus (H. Christ) Herter; Die Bärlappgewächse 37 (1939) = Urostachys selago f. reductus-angustinus (H. Christ) Herter ex Nessel; Die Bärlappgewächse 37 (1939) Huperzia sellifolia B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 68, f. 12C (1988) Distribution [complete]: Ecuador, Colombia, Peru Remarks: brevifolia agg. = Urostachys compactus f. minor Herter; Revista Sudamericana de Botánica 10: 119-120 (1954) Huperzia sellowiana (Herter) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution: Brazil Remarks: brongniartii agg. = Lycopodium sellowianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 44 (1909) = Urostachys sellowianus (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 385 (1927) Huperzia serpentiformis (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988) Distribution [complete]: Colombia Remarks: brevifolia agg. = Lycopodium serpentiforme Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 46 (1909) = Urostachys serpentiformis (Herter) Herter ex Nessel; Bärlappgewächse 104 (1939) Huperzia serrata (Thunb. ex Murray) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875) Distribution: India (West Bengal, Sikkim, Arunachal Pradesh, Assam, Meghalaya, Manipur, Tamil Nadu), Nepal, Burma, Sri Lanka, NE-Thailand, peninsular Thailand, Fiji (Viti Levu), Western Samoa (Savaii), peninsular Malaysia (Pahan), Borneo (Mt. Kinabalu, etc.), E-Siberia / Amur / Ussuri, Taiwan, Cuba, Hispaniola (Haiti), Mexico (Hidalgo, Oaxaca), Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hubei, Hunan, Jiangxi, Jilin, Liaoning, Shaanxi, Sichuan, Japan, Ryukyu Isl., Korea, Vietnam, Hawaii (Kauai, Oahu, Molokai, Lanai, Hawaii Isl.), New Caledonia, NE-Queensland Remarks: selago agg. = Huperzia selago (L.) Bernh. ex Schrank & C. Mart. ssp. serrata (Thunb.) A. & D. Löve = Huperzia serrata (Thunb.) Rothm.; Feddes Repert. 54 (1): 59 (1944) = Huperzia serrata f. longipetiolata (Spring) Ching; Acta Botanica Yunnanica 3(3): 294 (1981) = Huperzia shangsiensis (Chun Y. Yang) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) = Lycopodium serratum Thunb. ex Murray; Systemat Vegetabilium. Editio decima quarta 944 (1784), also Thunb., Fl. Jap. 341, t. 38 (1784) = Lycopodium serratum var. javanicum (Sw.) Makino; Botanical Magazine 12(131): 13 (1898) = Lycopodium serratum var. longipetiolatum Spring; Monogr. Lycop. 2: 18 (1850) = Lycopodium serratum var. myriophyllifolium Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 4: 133 (1914) = Lycopodium serratum var. thunbergii Makino; Botanical Magazine 12(131): 12 (1898) = Phlegmariurus shangsiensis Chun Y. Yang; Acta Phytotaxonomica Sinica 22(1): 87-88, f. 1 (1984) = Plananthus serratus (Thunb. ex Murray) P. Beauv.; Prod. aethéogam. 100, 111 (1805) = Urostachys javanicus (Sw.) Herter; Index Lyc. 66 (1949) = Urostachys myriophyllifolius (Hayata) Herter; Estudios Botánicos en la Región Uruguaya 20: 71 (1949) = Urostachys serratus (Thunb. ex Murray) Herter; Bot. Arch. 3: 13 (1923) = Urostachys serratus (Thunb. ex Murray) Herter var. sachalinensis Nessel; Repert. Spec. Nov. Regni Veg. 36: 179, t. 171. 1934 = Urostachys serratus var. japonica-neotropicus Herter ex Nessel (1940) = Urostachys sachalinensis (Nessel) Herter; Index Lyc. 80. 1949 = Lycopodium sargassifolium Liebm.; Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1847: 41 (1847) f. intermedia (Nakai) Ching; Acta Botanica Yunnanica 3(3): 294 (1981) Distribution: China, Japan = Lycopodium serratum f. intermedium Nakai; Botanical Magazine 39(463): 196-197 (1925) var. integrifolium (Matsuda) comb ined. Distribution: China = Lycopodium serratum Thunb. ex Murray var. integrifolium Matsuda; Bot. Mag. (Tokyo) 25: 124 (1911) = Lycopodium integrifolium (Matsuda) Matsuda & Nakai ex Nakai; Bot. Mag. (Tokyo) 28: 102. 1914 [non Goldie (1822) var. longipetiolatum Spring Distribution: SE-Asia, common, Japan = Lycopodium javanicum Sw.; J. Bot. (Schrader) 1800 (2): 114. 1801 [Syn. Fil. 175, 399 (1806]) var. myriophyllifolium Hayata Distribution: Distribution unknown Huperzia setifolia (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution: Borneo Remarks: unverified name, phlegmaria agg. = Lycopodium setifolium Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 16: 40 (1914) = Urostachys setifolius (Alderw.) Herter ex Nessel; Bärlappgewächse 233 (1939) Huperzia sieberiana (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Martinique, St. Kitts, St. Vincent, ?Haiti Remarks: reflexa agg. = Lycopodium sieberianum Spring; Flora 21: 153 (1838) = Lycopodium squarrosum Lam.; Encycl. 3. 653. 1792 [non G. Forster 1786, nec Sw. (1788) = Urostachys sieberianus (Spring) Herter; Repert. Spec. Nov. Regni Veg. 19: 162 (1923) Huperzia sieboldii (Miq.) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution: Taiwan, Japan, Korea (Cheju Isl.), China, Ryukyu Isl. Remarks: cancellata agg. = Lycopodium sieboldii Miq.; Ann. Mus. Bot. Lugduno-Batavum 3: 184 (1867) = Phlegmariurus sieboldii (Miq.) Ching; Acta Bot. Yunnanica 4: 121(1982) = Phlegmariurus sieboldii (Miq.) Ching; Flora Xizangica 1: 12 (1983), nom superfl. [nom. illeg., non Lycopodium squarrosum G. Forst (1786) = Urostachys sieboldii (Miq.) Herter ex Nessel; Bärlappgewächse 138 (1939) Huperzia silveirae (Nessel) B. Øllg. & Wind.; Bradea 5: 17 (1987) Distribution: Brazil (Minas Gerais) Remarks: brongniartii agg. = Urostachys silveirae Herter ex Nessel; Arch. Bot. Sao Paulo 1: 386 (1927) Huperzia simonii (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985) Distribution: Bismarck Arch. Remarks: unverified name, phlegmaria agg. = Urostachys simonii Nessel; Revista Sudamer. Bot. 6: 167, excl. fig (1940) Huperzia sintenisii (Nessel) Holub; Folia Geobet. Phytotax. 20: 76 (1985) Distribution [complete]: Puerto Rico, Hispaniola (Haiti) Remarks: reflexa agg. = Lycopodium sintenisii (Herter) Maxon ex C. Chr.; Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16 (2): 91 (1937) = Urostachys sintenisii Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 161 (1923) Huperzia societensis (J. Moore) comb. ined. Distribution [complete]: Society Isl. (Raiatea) = Lycopodium societense J. Moore; Bernice P. Bishop Mus. Bull. 102: 12 (1933) Huperzia socratis (Herter) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21(100): 157 (1988) Distribution [complete]: Colombia Remarks: brongniartii agg. = Lycopodium socratis (Herter) C. Morton; Amer. Fern 1. 54: 72 (1964) = Urostachys socratis Herter; Revista Sudamer. Bot. 10: 117 (1953 [1954]) Huperzia somai (Hayata) Ching; Acta Bot. Yunnanica 3: 301 (1981) Distribution [complete]: Taiwan, Japan, Philippines (N-Luzon) Remarks: selago agg. = Lycopodium chinense var. somai (Hayata) Masam.; Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 4: 114 (1934) = Lycopodium selago var. somai (Hayata) Masam.; Short Fl. Form. 32 (1936) = Lycopodium somai Hayata; Ic. pl. formos. 5: 255, f. 91 (1915) = Urostachys somai (Hayata) Herter ex Nessel; Bärlappgewächse 54 (1939) Huperzia sooiana (Lawalrée) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution [complete]: Zaire Remarks: phlegmaria agg. = Phlegmariurus sooianus LawaIrée; Acta Bot. Acad. Sci. Hung. 19: 195, f. 1 (1973) Huperzia sotae (Rolleri) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Argentina (Jujuy, Salta, Tucuman), Bolivia Remarks: taxifolia agg. = Lycopodium sotae Rolleri; Darwiniana 16: 130, f. 1 (A-C), t. 2, t. 3(A) (1970) Huperzia spongiosa (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 157 (1988) Distribution [complete]: Colombia Remarks: reflexa agg. = Lycopodium spongiosum Rolleri; Revista Mus. La Plata (N. S.) Bot. 14 (87): 6, f. 3-4 (1985) Huperzia sprengeri (Nessel) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution [complete]: Society Isl. (Tahiti) Remarks: selago agg. = Urostachys sprengeri Nessel; Repert. Spec. Nov. Regni Veg. 36: 179, t. 171 (1934) Huperzia squarrosa (C.Forster) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874) Distribution: India (West Bengal, Sikkim, Meghalaya, Assam, Kerala), Nepal, Burma, Bangladesh, Sri Lanka, Malesia, Philippines, peninsular Malaysia (Kedah, Kelantan, Perak, Pahan), Borneo (Mt. Kinabalu, etc.), Java, Thailand (widespread), China, Taiwan, Japan, NE-Queensland, Madagascar, Mauritius, Réunion, Seychelles, Palau Isl. (Babeldaob), Micronesia (Pohnpei, Kosrae), Tonga (Late Isl., Tafahi), New Caledonia, Society Isl. (Tahiti), Cook Isl. (Rarotonga), American Samoa (Manua Isl., Tutuila, Ta'u, Olosega), Western Samoa (Savaii, Upolu), Austral Isl. (Rapa Iti, Tubuai) Remarks: squarrosa agg. = Huperzia cunninghamioides (Hayata) Holub; Folia Geobot. Phytotax. 20(1): 72 (1985) = Huperzia epiceifolia (Desv. ex Poiret) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) = Huperzia lecomteana (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985) = Huperzia squarrosa (G. Forst.) Rothm.; Repert. Spec. Nov. Regn. Veg. 54: 62 (1944) = Lycopodium cunninghamioides Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 4: 131 (1914) = Lycopodium epiceifolium Desv. ex Poiret; Encycl. suppl. 3: 559 (1813 [1814]) = Lycopodium forsteri Poiret; Encycl. suppl. 3: 545 (1813 [1814]) [nom. superfl. = Lycopodium gnidioides Blanco; Fl. Filip. 824. 1837, ed. 2.569. 1845, ed. 3 (3): 239 (1879) = Lycopodium madagascariense Desv.; [nom. inval. (Spring 1842: 34) = Lycopodium pseudo-squarrosum Pampan.; Bull. R. Soc. Tosc. Orticult. 111, 13: 99, t. 2 (1908) = Lycopodium squarrosum G. Forster; Fl. ins. austr. 479 (1786) = Phlegmariurus cunninghamioides (Hayata) Ching; Acta Botanica Yunnanica 4(2): 125 (1982) = Phlegmariurus squarrosus (G. Forster) Löve & Löve; Taxon 26: 324 (1977) = Plananthus squarrosa (Sw.) P. Beauv.; Prodr. Aetheog. 112 (1805) = Urostachys cunninghamioides (Hayata) Herter ex Nessel; Die Bärlappgewächse 206 (1939) = Urostachys epiceaefolius (Desv.) Herter ex Nessel; Die Bärlappgewächse 202 (1939) = Urostachys epiceaefolius var. acutifolius (Desv.) Herter ex Nessel; Die Bärlappgewächse 292 (1939) = Urostachys epiceifolius (Poiret) Herter ex Nessel; Bärlappgewächse 202 (1939) = Urostachys lecomteanus Nessel; Repert. Spec. Nov. Regni Veg. 36: 1871 t. 174 (1934) = Urostachys madagascariensis (Desv. ex Nessel) Herter; Estudios Botánicos en la Región Uruguaya 20: 68 (1949), Alt. title: Index Lycopodiarum 68 (1949) = Urostachys madagascariensis (Nessel) Herter; Index Lyc. 68 (1949) = Urostachys squarrosus (G. Forst.) Herter; Botanisches Archiv 3: 14 (1923) = Urostachys squarrosus (G. Forster) Herter var. prolifer (Blume) Nessel f. madagascariensis Nessel = Urostachys squarrosus var. proliferus (Blume) Herter ex Nessel; Bärlappgewächse 200 (1939) Huperzia staudtii (Nessel) Pichi-Serm.; Webbia 23: 163 (1968) Distribution [complete]: Cameroun, Ivory Coast, Ghana, Nigeria, Angola, Central African Republic, Gabon Remarks: phlegmaria agg. = Lycopodium staudtii (Nessel) C. Adams & Alston; Bull. Brit. Mus. (Nat. Hist.) Bot. 1 (6): 183 (1955) = Urostachys adolfi-friedericii Herter ex Nessel; Die Bärlappgewächse 226 (1939) [nom. inval., without Latin description = Urostachys staudtii Nessel; Repert. Spec. Nov. Regni Veg. 36: 189, t. 175 (1934) Huperzia stemmermanniae (Medeiros & Wagner) comb. ined. Distribution [complete]: Hawaii = Phlegmariurus stemmermanniae Medeiros & Wagner; Amer. Fern Journ. 86(3): 89-97 (1996) Huperzia suberecta (Lowe) Tard.; Adansonia 10: 20 (1970) Distribution [complete]: Madeira, Madagascar, Réunion Remarks: selago agg. = Huperzia selago ssp. suberecta (Lowe) Toledo & Vasc.; Bol. Soc. Brot., ser. 2 41: 23 (1967) = Lycopodium selago ssp. suberectum (Lowe) Romariz; Revista da Facultade de Ciências : 2.a Serie : C : Ciências Naturais 3: 71, t. 1, f. 3 (1953) = Lycopodium selago var. suberectum (Lowe) C. Chr.; Dansk Botanisk Arkiv 7: 188 (1932) = Lycopodium suberectum Lowe; Trans. Catnbridge Philos. Soc. 4 (1): 9 (1831) = Urostachys suberectus (Lowe) Herter ex Nessel; Bärlappgewächse 41 (1939) Huperzia subfalciformis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: New Guinea Remarks: unverified name, phlegmarioides agg. = Lycopodium subfalciforme Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 28: 44 (1918) = Urostachys subfalciformis (Alderw.) Herter ex Nessel; Bärlappgewächse 227 (1939) Huperzia subintegra (Hillebr.) Beitel & W.H. Wagner Distribution [complete]: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.) Huperzia submoniliformis (Alderw.) comb. ined. Distribution: New Guinea Remarks: unverified name, phlegmaria agg. = Lycopodium horizontale Alderw. var. submoniliforme Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 4 (1917) = Urostachys submoniliformis (Alderw) Herter; Index Lyc. 84 (1949) Huperzia subtrifoliata (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution [complete]: Fiji (Viti Levu, Vanua Levu) Remarks: phlegmaria agg. = Lycopodium subtrifoliatum Brownlie; Beih. Nova Hedwigia 55: 24 (1977) Huperzia subtubulosa (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution [complete]: Hispaniola Remarks: taxifolia agg. = Urostachys subtubulosus Herter; Repert. Spec. Nov. Regni Veg. 19: 166 (1923) Huperzia subulata (Desv. ex Poir.) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution [complete]: Costa Rica, Honduras, Panama, Colombia, Ecuador, Peru, Argentina (Salta, Tucuman), Bolivia, Venezuela Remarks: myrsinites agg. = Lycopodium ewanii (Herter) C. V. Morton; Amer. Fern Journal 54: 72 (1964) = Lycopodium subulatum Desv. ex Poir.; Encyclopedie Methodique. Botanique . Supplement 3: 544. [1813] (1814) = Urostachys ewanii Herter; Revista Sudamericana de Botánica 10: ,126-127 (1954) = Urostachys subulatus (Desv. ex Poir.) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 420 (1927) Huperzia subulifolia (Hook. & Grev.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution: India (West Bengal, Sikkim, Meghalaya, Uttar Pradesh), Nepal, Sri Lanka Remarks: verticillata agg. = Lycopodium subulifolium Wall. ex Hook. & Grev.; lcon. Fil. t. 49 (1831) = Urostachys subulifolius (Hook. & Grev.) Herter; Index Lyc. 84 (1949) Huperzia sumatrana (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Sumatra Remarks: unverified name, selago agg. = Lycopodium sumatranum Alderw.; Bull. Jard. Bot. Buitenzorg ser. 2, 28: 44 (1918) = Urostachys sumatranus (Alderw.) Herter ex Nessel; Bärlappgewächse 66 (1939) Huperzia sutchueniana (Herter) Ching; Acta Bot. Yunnanica 3: 297 (1981) Distribution: E-China Remarks: unverified name, selago agg. = Lycopodium serratum var. alpestre H. Christ; Bulletin de l'Académie Internationale de Géographie, Botanique 16(199-201): 141 (1906) = Lycopodium sutchuenianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909) = Urostachys sutchuenianus (Herter) Herter ex Nessel; Bärlappgewächse 53 (1939) Huperzia tafanensis (Nessel) comb. ined. Distribution: Papua New Guinea Remarks: unverified name, = Urostachys tafanensis Nessel; Repert. Spec. Nov. Regni Veg. 39: 61, t. 189b (1935) Huperzia tahkuanensis Ching; Acta Botanica Yunnanica 3(3): 297 (1981) Distribution: China Remarks: selago agg. Huperzia takingensis Ching; Acta Botanica Yunnanica 3(3): 300-301 (1981) Distribution: China Remarks: selago agg. Huperzia talamancana B. Øllg.; Novon 3(1): 67, f. 2 (1993) Distribution [complete]: Costa Rica, Panama Huperzia talamauana (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Sumatra Remarks: unverified name, phyllantha agg. = Lycopodium talamauanum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2,28: 45 (1918) = Urostachys talamauanus (Alderw.) Herter ex Nessel; Bärlappgewächse 212 (1939) Huperzia talpiphila B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 44, f. 9A (1988) Distribution [complete]: Ecuador Remarks: saururus agg. Huperzia tauri (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution: Java Remarks: unverified name, phlegmaria agg. = Lycopodium apiculatum Spring; Miquel, Pl. Jungh. 3: 272 1854 [non Poiret 1814 = Urostachys tauri Herter; Index Lyc. 85 (1949) = Urostachys apiculatus (Spring) Herter ex Nessel; Bärlappgewächse 227 (1939) Huperzia taxifolia (Sw.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Puerto Rico, Cuba, Hispaniola, Jamaica, Costa Rica, Nicaragua, Mexico (Chiapas, Sinaloa, Michoacan, Tamaulipas, San Luis Potosi, Distrito Federal, Veracruz, Puebla, Oaxaca), Belize, Guatemala, Honduras, El Salvador, Colombia, Venezuela, Ecuador, Galapagos, Saba Isl., St. Kitts, Nevis, Montserrat, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent, Grenada, British Guyana, Surinam, French Guiana, Peru, Bolivia Remarks: taxifolia agg. = Huperzia passerinoides (Kunth) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) = Huperzia passerinoides var. nitens (Schlecht. & Cham.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) = Lycopodium americanum Desv.; [herb. Vaillant = Lycopodium herminieri Spring; Bull. Acad. Roy. Sci. Belg. 8514. 1841 = Lycopodium nitens Schlecht. & Cham.; Linnaea 5: 623 (1830) = Lycopodium passerinoides Kunth; Nova Genera & Sp. Pl. 1: 41 (1815)[1816] = Lycopodium schwendenerii Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 50 (1909) = Lycopodium taxifolium Sw.; Nova Genera & Sp. Pl. seu Prodromus 138 (1788) = Lycopodium taxifolium var. passerinoides (Kunth) Baker; Handbook of the Fern-Allies 16 (1887) = Phlegmariurus taxifolius (Sm.) Love & Love; Taxon 26(2-3): 324 (1977) = Plananthus taxifolius (Sw.) P. Beauv.; Prod. aethéogam. 112 (1805) = Urostachys bruelkei Nessel; Repert. Spec. Nov. Regn. Veg. 36(12-15): 184, t. 173 (1934) = Urostachys herminieri (Spring) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923) = Urostachys nitens (Schltdl. & Cham.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923) = Urostachys passerinoides (Kunth) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 417 (1927) = Urostachys passerinoides var. nitens (Schlecht. & Cham.) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 418 (1927) = Urostachys rubiginosus Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 162, t. 8, f. 36 (1940) = Urostachys schwendeneri (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 165 (1923) = Urostachys schwendeneri (Herter) Herter var. spongiosus Nessel; Revista Sudamer. Bot. 6: 165 (1940) = Urostachys schwendeneri var. nitens (Schlecht. & Cham.) Herter ex Nessel; Die Bärlappgewächse 166 (1939) = Urostachys spongiosus (Nessel) Herter; Index Lyc. 83 (1949) = Urostachys taxifolius (Sm.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923) Huperzia tenuicaulis (Underw. & Lloyd) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Puerto Rico, Guadeloupe, Dominica, British Guyana Remarks: linifolia agg. = Urostachys tenuicaulis (Underwood & Lloyd) Herter; Repert. Spec. Nov. Regni Veg. 19: 165 (1923) = Lycopodium tenuicaule Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 113 (1906) Huperzia tenuis (Humb. & Bonpl. ex Willd.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Costa Rica, Panama, Colombia, Venezuela, Ecuador, N-Peru Remarks: verticillata agg. = Lycopodium tenue Humb. & Bonpl. ex Willd.; Sp. Pl., Editio quarta 5: 55 (1810) = Urostachys pax-hominibus Herter; Index Lyc. 74 (1949) [nom. superfl. for Lycopodium tenue Willd. = Urostachys tenuis (Humb. & Bonpl. ex Willd.) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 401 (1927) Huperzia terrae-guilelmii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Irian Jaya Remarks: unverified name, phlegmaria agg. = Lycopodium terrae-guilelmii Herter; Bot. Jahrb. Syst. 54: 229 (1916) = Urostachys terrae-guilelmii (Herter) Herter ex Nessel; Bärlappgewächse 231 (1939) Huperzia tetragona (Hook. & Grev.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Ecuador, Peru, Colombia, Bolivia Remarks: saururus agg. = Lycopodium catharticum Hook.; Annals of Natural Hisotry 1: 430-431, t. 14 (1838) = Lycopodium myrsinites var. minus Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 29 (1842) = Lycopodium tetragonum Hook. & Grev.; Icon. filic. 1: t. 109 (1829) = Lycopodium tetragonum var. patulum Spring; Mémoires de l'Académie Royales des Sciences, Lettres & Beaux Arts de Belgique 24: 12 (1849) = Urostachys catharticus (Hook.) Herter ex Nessel; Bärlappgewächse 135 (1939) = Urostachys tetragonus (Hook. & Grev.) Herter ex Nessel; Bärlappgewächse 135 (1939) Huperzia tetrasticha (Kunze) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Java, peninsular Malaysia (Kelantan, Pahan, Negeri Sembilan), Sulawesi Remarks: carinata agg. = Lycopodium tetrastichum Kunze; Bot. Zeit. (Berlin) 6: 99 (1848) = Lycopodium sarasinorum Christ; Verh. Naturf. Ges. Basel 11: 34, t. 2o f. 11-12 (1894) = Urostachys sarasinorum (Christ) Herter ex Nessel; Bärlappgewächse 182 (1939) = Urostachys tetrastichus (Kunze) Herter ex Nessel; Bärlappgewächse 183 (1939, by reference to Alderw (1915a: 35)) Huperzia thwaitesii (Nessel) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Java Remarks: unverified name, selago agg. = Urostachys thwaitesii Herter ex Nessel; [Bärlappgewächse 52. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 158, t. 8, f. 14 (1940) Huperzia tibetica (Ching) Ching; Acta Bot. Yunnanica 3: 304 (1981) Distribution: Yunnan, Tibet Remarks: selago agg. = Lycopodium tibeticum Ching; Bull. Fan Mem. Inst. Biol. Bot. Ser. 10: 17 (1940) = Huperzia tibetica (Ching) Ching; Flora Xizangica 1: 7 (1983) [nom. superfl. Huperzia toppingii (Herter) Holub; Folia Geobot. Phytotax. 20: 77(1985) Distribution: Philippines Remarks: unverified name, squarrosa agg. = Urostachys toppingii Herter; Bot. Arch. 3: 15 (1923) Huperzia tournayana (LawaIrée) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution [complete]: Zaire Remarks: phlegmaria agg. = Phlegmariurus tournayanus Lawalrée; Acta Bot. Acad. Sci. Hung. 19: 197, f. 2 (1973) Huperzia transiens (Alderw.) comb. ined. Distribution: Sumatra Remarks: unverified name, phlegmaria agg. = Lycopodium transiens Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 227 (1922) = Urostachys transiens (Alderw.) Herter ex Nessel; Bärlappgewächse 143 (1939) Huperzia transilla (Baker) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Ecuador Remarks: brevifolia agg. = Lycopodium transilla Sodiro ex Baker; J. Bot. 15: 168 (1877) = Urostachys transilla (Baker) Herter ex Nessel; Bärlappgewächse 106 (1939) Huperzia treitubensis (Silveira) B. Øllg.; Opera Bot. 92: 170 (1987) Distribution: Brazil (Minas Gerais) Remarks: brongniartii agg. = Lycopodium treitubense Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2, 5: 118, t. 3 (1898) = Urostachys treitubensis (Silveira) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 380 (1927) = Lycopodium hoehnei (Nessel) Rolleri; Revista Mus. La Plata (N. S.) Bot. 13: 77 (1981) = Urostachys hoehnei Nessel; Arch. Bot. Sao Paulo 1- 383, t. 6 (1927) = Lycopodium inflexum Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2,5: 118, t. 2 (1898) [non (P. Beauv.) Sw. (1806) = Urostachys inflexus Herter ex Nessel; Arch. Bot. Sao Paulo 1: 384. 1927 [nom. nov. for Lycopodium inflexum Silveira [non (P. Beauv.) Sw. (1806]) = Urostachys capri Herter; Index Lyc. 55 (1949) [nom. superfl. Huperzia trifoliata (Copel.) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution [complete]: Fiji (Viti Levu, Vanua Levu, Kadavu, Ovalau, Taveuni) Remarks: phlegmaria agg. = Lycopodium trifoliatum Copel.; Bernice P. Bishop Mus. Bull. 59: 7, t. 1 (1929) = Urostachys trifoliatus (Copel.) Herter ex Nessel; Bärlappgewächse 224 (1939) Huperzia trigona (C. Chr.) Tard.; Adansonia 10: 20 (1970) Distribution [complete]: Madagascar Remarks: obtusifolia agg. = Lycopodium trigonum C. Chr.; Dansk Bot. Ark. 7.191, t. 76, f. 1-5 (1932) = Urostachys humbertii Nessel; Repert. Sp. Nov. Regni Veg. 48: 170, t. 317 ("307"), f. 3 (1940) = Urostachys trigonus (C. Chr.) Herter; Index Lyc. 87 (1949) Huperzia tubulosa (Maxon) B. Øllg.; Annals of the Missouri Botanical Garden 79: 697 (1992) Distribution [complete]: Costa Rica, Panama, Venezuela Remarks: taxifolia agg. = Lycopodium tubulosum Maxon; Contributions from the U. S. National Herbarium 17(2): 178, t. 10 (1913) = Urostachys tubulosus (Maxon) Herter ex Nessel; Die Bärlappgewächse 177 (1939) Huperzia ulei (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Brazil (Rio de Janeiro) Remarks: unverified name, brongniartii agg. = Urostachys ulei Herter; Repert. Spec. Nov. Regni Veg. 19162 (1923) Huperzia ulicifolia (Sw.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution: Distribution unknown Remarks: unverified name, squarrosa agg. = Lycopodium ulicifolium Sw.; Syn. fil. 177 (1806) = Urostachys ulicifolius (Sw.) Herter ex Nessel; Bärlappgewächse 204 (1939) Huperzia ulixis (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution [complete]: Colombia Remarks: saururus agg. = Urostachys ulixis Herter; Revista Sudamer. Bot. 10: 115 (1953 [1954]) = Lycopodium ulixis (Herter) C. Morton; Amer. Fern 1. 54: 73 (1964) Huperzia unguiculata B. Øllg.; Harling & Andersson: Fl. Ecuador 33: 26, f. 2B (1988) Distribution: Ecuador Remarks: reflexa agg. Huperzia urbanii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution [complete]: Ecuador, Colombia Remarks: reflexa agg. = Lycopodium urbanii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 47(1909) = Urostachys urbanii (Herter) Herter ex Nessel; Bärlappgewächse 110 (1939) Huperzia vanikorensis (Copel.) Holub; Folia Geobot. Phytotax. 20: 77 (1985) Distribution: Santa Cruz Islands, Solomons, Vanuatu Remarks: unverified name, carinata agg. = Lycopodium vanikorense Copel.; Journal of the Arnold Arboretum 12: 49 (1931) = Urostachys vanikorensis (Copel.) Herter; Index Lyc. 87 (1949) Huperzia varia (R. Br.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874) Distribution [complete]: Tasmania, Queensland, SE-New South Wales, S-Victoria, New Zealand (North Isl., South Isl., Three Kings Isl., Stewart Isl.), Kermadec Isl., Chatham Isl., Antipodes Isl., Auckland Isl., Campbell Isl., Lord Howe Isl., New Ireland Remarks: varia agg. = Huperzia billardieri (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 47. 1874 = Huperzia novae-zelandica (Colenso) Holub; Folia Geobot. Phytotax. 20: 75 985) = Lycopodium billardierei Spring; Mon. Lycop. 1, 56 (1843) = Lycopodium flagellaria A. Rich.; Essai Fl. N. Z., 60 [non Bory (1832) = Lycopodium novae-zelandicum Colenso; Trans. & Proc. New Zealand st. 19: 275 (1887) = Lycopodium phlegmaria A. Cunn.; Compan. bot. Mag. 2, 361 [non L. 837) = Lycopodium varium R. Br.; Prodr. 65 (1810) = Lycopodium varium var. alpinum R. Br. (1810) = Lycopodium varium var. billardieri (Spring) T. Kirk; T. N. Z. I. 16, 377 884) = Lycopodium varium var. umbrosum R. Br. (1810) = Urostachys billardieri (Spring) Herter ex Nessel; auml;rlappgewächse 194 (1939) = Urostachys varius (R. Br.) Herter ex Nessel; Bärlappgewächse 2 (1939) Huperzia venezuelanica (Herter) Holub; Folia Geobot. Phytotax. 20: 77 985) Distribution [complete]: Venezuela (Sa. Nevada de Merida) Remarks: saururus agg. = Lycopodium venezuelanicum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 909) = Urostachys venezuelanicus (Herter) Herter ex Nessel; auml;rlappgewächse 74 (1939) = Huperzia schneei (Vareschi) Holub; Folia Geobot. Phytotax. 20: 76 985) = Lycopodium schneei Vareschi; Acta Bot. Venez. 1: 83, f. 1 (1966) Huperzia vernicosa (Grev. & Hook.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 8 (1874) Distribution: India (Tamil Nadu, Korala), Sri Lanka Remarks: selago agg. = Lycopodium vernicosum Hook. & Grev. ex Grev. & Hook.; Bot. Misc. 364 (1831) = Phlegmariurus vernicosus (Grev. & Hook.) Löve & Löve; xon 26: 324 (1977) = Urostachys vernicosus (Grev. & Hook.) Herter ex Nessel; auml;rlappgewächse 61 (1939) Huperzia verticillata (L. f.) Trevisan; Atti della Società Crittogamologica Italiana 17: 248 (1875) Distribution: Puerto Rico, Jamaica, Cuba, Hispaniola, Mexico, Venezuela, Colombia, Brazil, Bolivia, Trinidad, Saba Isl., Guadeloupe, Dominica, Martinique, St. Lucia, Transvaal, Natal, Cape Prov., Swaziland, Zimbabwe, Malawi, Mozambique, Réunion, Mauritius, Kenya, Tanzania, Transkei, Lesotho, NE-Oranje Free State, Madagascar, Comores, Bioko Isl. (Fernando Poo), Sao Tomé, Cameroun, Equatorial Guinea, Zaire, Rwanda Remarks: verticillata agg. = Lycopodium setaceum Lam.; Encycl. 3: 653 (1789 [1792]) = Plananthus setaceus (Lam.) P. Beauv.; Prod. aethéogam. 112 (1805) = Lycopodium squarrulosum Desv. (1823) = Lycopodium portoricense L.M. Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 108 (1906) = Lycopodium verticillatum L. f.; Supplementum Plantarum 448 (1782) = Plananthus verticillatus (L. f.) P. Beauv.; Prod. aethéogam. 112 (1805) = Stachygynandrum verticillatum (L. f.) P. Beauv.; Prodr.aethéogam. 113 (1805) = Urostachys portoricensis (Underwood & Lloyd) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 163 (1923) = Urostachys verticillatus (L. f.) Herter; Beihefte zum Botanischen Centralblatt 39: 249 (1922) Huperzia vorwerkii (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985) Distribution: India (Himalaya) Remarks: unverified name, selago agg. = Urostachys vorwerkii Nessel; Repert. Spec. Nov. Regni Veg. 36: 178, t. 170 (1934) Huperzia vrieseana (Spring) comb. ined. Distribution: Java Remarks: unverified name, phlegmaria agg. = Lycopodium vrieseanum Spring; Mem. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 33 (1849) = Urostachys vrieseanus (Spring) Herter ex Nessel; Bärlappgewächse 236 (1939) Huperzia warneckei (Nessel) Pichi-Serm.; Webbia 23: 163 (1968) Distribution [complete]: Cameroun, Guinea, Sierra Leone, Ivory Coast, Uganda, Tanzania, Sao Tomé, Liberia Remarks: phlegmaria agg. = Urostachys warneckei Herter ex Nessel; [Bärlappgewächse 213. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 166, t. 13, f. 64 (1940) = Lycopodium warneckei (Nessel) Alston; Bol. Soc. Brot. ser. 2A, 30: 20 (1956) Huperzia watsoniana (Maxon) B. Øllg.; Annals of the Missouri Botanical Garden 79: 701 (1992) Distribution [complete]: El Salvador, Costa Rica, Panama Remarks: verticillata agg. = Lycopodium watsonianum Maxon; Smithsonian Miscellaneous Collections 56(29): 3, t. 3 (1912) = Urostachys watsonianus (Maxon) Herter ex Nessel; Bärlappgewächse 159 (1939) Huperzia weberbaueri (Hieron. & Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985) Distribution [complete]: Peru, Ecuador Remarks: saururus agg. = Huperzia papillata (Rolleri) Holub; Folia Geobot. Phytotax. 20: 75 (1985) = Lycopodium papillatum Rolleri; Amer. Fern Journal 65(1): 3 (1975) = Urostachys weberbaueri Herter ex Nessel; [Bärlappgewächse 99. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 162, t. 10p f. 41 (1940) Huperzia weddellii (Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985) Distribution [complete]: Peru, Ecuador, Bolivia Remarks: brongniartii agg. = Lycopodium weddellii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909) = Urostachys weddellii (Herter) Herter ex Nessel; Bärlappgewächse 91 (1939) Huperzia whangshanensis Ching & P.C. Chiu; Acta Botanica Yunnanica 3(3): 299 (1981) Distribution: S-Anhui Remarks: selago agg. Huperzia whitfordii (Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985) Distribution: Philippines (Luzon) Remarks: unverified name, squarrosa agg. = Urostachys whitfordii Herter; Bot. Arch. 3: 14 (1923) = Lycopodium whitfordii (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931) Huperzia wilsonii (Underw. & F.E. Lloyd) B. Øllg.; Opera Bot. 92: 170 (1987) Distribution [complete]: Puerto Rico, Guatemala, Nicaragua, Costa Rica, Panama, Mexico (Oaxaca), Jamaica, Colombia, Venezuela, Ecuador, Peru, Guadeloupe, Saba Isl., Guadeloupe, Martinique, St. Lucia, St. Vincent, Surinam, Argentina (Misiones, Salta, Tucuman), Brazil, Paraguay, Uruguay Remarks: wilsonii agg. = Huperzia lindaviana (Herter) Holub; Folia Geobot. Phytotax. 33: 97 (1988) = Huperzia mandiocana (Raddi) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248. 1874 = Lycopodium pseudo-mandiocanum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 49 (1909) = Urostachys pseudo-mandiocanus (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 411 (1927) = Lycopodium andinum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 49. 1909 [non Rosenstock (1908) = Lycopodium arcanum Maxon; Field Museum of Natural History, Botanical Series 17(4): 310, t. 3 (1938) = Lycopodium lindavianum Herter; Hedwigia 49(2-3): 90., as Lycopodium "Lindavianum" (1909) = Lycopodium mandiocanum Raddi; Plantarum Brasiliensium Nova Genera 1: 77-78, t. 4 (1825) = Lycopodium stamineum Maxon; Smithsonian Miscellaneous Collections 56(29): 2-3, t. 2 (1912) = Lycopodium trichodendron Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 49 (1909) = Lycopodium wilsonii Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 111 (1906) = Urostachys lindavianus (Herter) Herter ex Nessel; Bärlappgewächse 146 (1939) = Urostachys mandiocanus (Raddi) Herter; Repert. Spec. Nov. Regni Veg. 19: 164 (1923) = Urostachys mandiocanus (Raddi) Herter var. filifolius Herter; Revista Sudamericana de Botánica 10: 124 (1954) = Urostachys stamineus (Maxon) Herter ex Nessel; Die Bärlappgewächse 148 (1939) = Urostachys trichodendron (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 163 (1923) = Urostachys wilsonii (Underw. & Lloyd) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 163 (1923) Huperzia x bartleyi (Cusick) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution [complete]: NE-USA, E-USA, SE-Canada Remarks: selago agg. Remarks: Lycopodium lucidulum x Lycopodium porophilum = Lycopodium x bartleyi Cusick; Amer. Fern Journal 77(3): 100 (1987) Huperzia x buttersii (Abbe) Kartesz & Gandhi Distribution: E-USA, E-Canada, Minnesota Remarks: selago agg. Remarks: Lycopodium lucidulum x Lycopodium selago = Lycopodium x buttersii Abbe; Rhodora 55(651): 91-92 (1953) Huperzia x calquistii Distribution [complete]: Hawaii (Maui) Remarks: Huperzia erubescens x Huperzia subintegra Huperzia x gillettii Beitel & W.H. Wagner Distribution [complete]: Hawaii (Maui, Kauai, Oahu, Molokai, Lanai, Hawaii Isl.) Remarks: Huperzia serrata x Huperzia subintegra Huperzia x medeirosii Beitel & W.H. Wagner Distribution [complete]: Hawaii (Maui) Remarks: Huperzia haleakalae x Huperzia subintegra Huperzia x sulcinervia (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution: Hawaii (Oahu, Lanai, Hawaii Isl.), E-Asia Remarks: Huperzia erubescens x Huperzia serrata = Huperzia sulcinervia (Spring) Rothm.; Feddes Repert. 54: 59 (1951) = Lycopodium x helleri Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909) = Urostachys helleri (Herter) Herter ex Nessel; Bärlappgewächse 53 (1939) = Lycopodium sulcinervium Spring; Bull. Acad. Roy. Sci. Belg. 8: 514 (1841) = Urostachys sulcinervius (Spring) Herter ex Nessel; Bärlappgewächse 54 (1939) Huperzia xiphophylla (Baker) Tardieu; Adansonia ser. 2 10: 20 (1970) Distribution [complete]: Madagascar Remarks: squarrosa agg. = Lycopodium xiphophyllum Baker; Journal of the Linnean Society, Botany 22: 535 (1887) = Urostachys xiphophyllus (Baker) Herter ex Nessel; Die Bärlappgewächse 72, t. 11, f. 9 (1939) Huperzia yakusimensis (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985) Distribution: Japan Remarks: selago agg. = Lycopodium yakusimensis Herter ex Nessel; [Bärlappgewächse 65. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 160, t. 8, f. 27 (1940) Huperzia yandongensis (Ching & Zhang) comb. ined. Distribution: Zhejiang Remarks: hamiltonii agg. = Phlegmariurus yandongensis Ching & Zhang; Bull. Bot. Res. North-East. Forest. Inst. 3 (3): 2 (1983) Huperzia yunnanensis (Ching) Holub; Folia Geobot. Phytotax. 20: 78 (1985) Distribution: Yunnan Remarks: cancellata agg. = Phlegmariurus yunnanensis Ching; Acta Bot. Yunnanica 4: 121 (1982) Huperzia zollingeri (Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985) Distribution: Java Remarks: unverified name, selago agg. = Lycopodium zollingeri Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909) = Urostachys zollingeri (Herter) Herter ex Nessel; Bärlappgewächse 120 (1939) `-- H. selago (ketunlieko)
Reference(s):
- Hassler, M. & Swale, B., 2001: Checklist of ferns and fern allies. (Updated: 2001-07-10)
–iNet: http://homepages.caverock.net.nz/~bj/fern/; [visited: 2005-12-14] - Hämet-Ahti, L., Suominen, J., Ulvinen, T., Uotila, P. & Vuokko, S., 1984: Retkeilykasvio.
–Suomen Luonnonsuojelun Tuki Oy & Forssan Kirjapaino Oy, Forssa, 1984, 1-544 - Judd, W. A., Campbell, C. S., Kellogg, E. A. & Stevens, P. F., 1999: Plant Systematics: A Phylogenetic Approach.
–Sinauer Associates, Inc., Sunderland, Massachusetts, 1999, xvi-464 - –iNet: The Tree of Life: http://tolweb.org/tree/
- Steward, W. N. & Rothwell, G. W., 1993: Paleobotany and the evolution of plants, 2nd Edition.
–Cambridge University Press, Cambridge, U.S.A., xii-521