<==o Mecistocephalidae Verhoeff, 1901
   |-- Brahmaputrus Verhoeff, 1942
   |-- Formosocephalus Verhoeff, 1937
   |--o Fusichila
   |  `-- F. waipaheenis Chamberlin, 1953
   |-- Megethmus Cook, 1895
   |-- Nodocephalus
   |-- Takashimaia
   |--o Arrupinae
   |  |--o Agnostrup
   |  |  |-- A. striatus (Takakuwa, 1949) [Taiwanella striatus Takakuwa, 1949]
   |  |  |-- A. paucipes (Miyosi, 1955) [Taiwanella paucipes Miyosi, 1955]
   |  |  `-- A. striganova (Titova, 1975) [Krateraspis striganova Titova, 1975]
   |  |--o Anarrup Chamberlin, 1920
   |  |  `-- A. flavipes (Attems, 1930) [Sundarrup flavipes Attems, 1930]
   |  |--o Arrup Chamberlin, 1912
   |  |  |-- A. areolatus (Shinohara, 1957) [Nodocephalus areolatus Shinohara, 1957]
   |  |  |-- A. asiaticus (Titova, 1975) [Nodocephalus asiaticus Titova, 1975]
   |  |  `-- A. mamaevi (Titova, 1975)[Prolamnonyx mamaevi Titova, 1975]
   |  |--o Nannarrup
   |  |  `-- N. hoffmani
   |  |--o Partygarrupius Verhoeff, 1939 [Tygarrup {Partygarrupius} Verhoeff, 1939]
   |  `--o Tygarrup Chamberlin, 1914
   |     |-- T. anepipe Verhoeff, 1939
   |     |-- T. asiaticus Verhoeff, 1930
   |     |-- T. intermedius Chamberlin, 1914
   |     |-- T. javanicus (Attems)
   |     |-- T. nepalensis Shinohara, 1965]
   |     `-- T. singaporiensis Verhoeff, 1937
   |--o Dicellophilinae
   |  `--o Dicellophilus Cook, 1895
   |     |-- D. anomalans (Chamberlin, 1904) [Mescistocephalus anomalansChamberlin, 1904]
   |     `-- D. carniolensis (Koch, 1847) [Mescistocephalus carniolensisKoch, 1847]
   |--o Mecistocephalinae [Oryidae: Mecistocephalinae]
   |  |--o Lamnonyx Cook, 1896
   |  |  |-- L. gracilis Verhoeff, 1925 [Mecistocephalus gracilis (Verhoeff, 1925)]
   |  |  |-- L. mater Verhoeff, 1925 [Mecistocephalus mater Verhoeff, 1925]
   |  |  |--o L. punctifrons ( Newport, 1842) [Mecistocephalus punctifrons Newport , 1842]
   |  |  |  `-- L. p. furculigera Verhoeff, 1925 [Mecistocephalus punctifrons fulculigera (Verhoeff, 1925)]
   |  |  |--o L. tahitiensis (Wood, 1863) [Mecistocephalus tahitiensis Wood, 1863]
   |  |  |  `-- L. t. major Verhoeff, 1925 [Mecistocephalus t. major (Verhoeff, 1925)]
   |  |  `-- L. togoensis Cook, 1896
   |  |--o MecistocephalusNewport, 1842 [Lamnonyx Cook, 1896]
   |  |  |-- M. castaneiceps Haase, 1887
   |  |  |-- M. cephalotes Meinert, 1871
   |  |  |-- M. collinus Verhoeff, 1937
   |  |  |-- M. gigas Haase, 1887
   |  |  |-- M. guildingii Newport, 1844
   |  |  |-- M. heteropus Humbert, 1865
   |  |  |-- M. hungaricus Tömösvary, 1880
   |  |  |-- M. insularis (Lucas, 1863)
   |  |  |-- M. insulomontanus Gressitt
   |  |  |-- M. japonicus Meinert, 1886
   |  |  |-- M. kurandanus Chamberlin, 1920
   |  |  |-- M. mauritanus Verhoeff, 1939
   |  |  |-- M. maxillaris(Gervais, 1837) [Geophilus maxillaris Gervais, 1837]
   |  |  |-- M. meinerti Sseliwanoff, 1881
   |  |  |-- M. microporus Haase, 1887
   |  |  |-- M. millepunctatus (Gervais, 1849)
   |  |  |-- M. modustusSilvestri
   |  |  |-- M. pilosus Wood, 1863
   |  |  |-- M. porteri Silvestri, 1899
   |  |  |-- M. punctilabium Newport, 1842
   |  |  |-- M. rubriceps Wood, 1863
   |  |  |-- M. simplex Chamberlin, 1920
   |  |  |-- M. spissus Wood, 1863 [Lamnonyx spissus]
   |  |  |-- M. sulcicollis Tömösvary, 1885
   |  |  |-- M. takakuwai Verhoeff, 1934
   |  |  |-- M. tenuiculus (Koch, 1878)
   |  |  |-- M. tridensChamberlin, 1922
   |  |  |-- M. waikaneus Chamberlin, 1953
   |  |  |-- M. yanagiharai (Takakuwa, 1936) [Taiwanella yanahiharai Takakuwa, 1936]
   |  |  |--o M. {Brachyptyx} Chamberlin, 1920
   |  |  |--o M. {Dasyptyx} Chamberlin, 1920
   |  |  |--o M. {Ectoptyx} Chamberlin, 1920
   |  |  `--o M. {Pauroptyx} Chamberlin, 1920
   |  `?-o Proterotaiwanella
   |     |-- P. sculptulata (Takakuwa, 1936) [Taiwanella sculptulata Takakuwa, 1936]
   |     `-- P. tanabei
   `--o Lamotteophilinae [Oryidae: Lamotteophilinae]
      `-- Lamotteophilus

Reference(s):

Special thanks to:

Jos Dols, for gathering this information.