After Prokop, Krzeminski, Krzeminska & Wojciechowski, 2012, Prokop, Krzeminski, Krzeminska, Hörnschemeyer, Ilger, Brauckmann, Grandcolas & Nel, 2013, and Guan, Prokop, Roques, Lapeyrie & Nel, 2015
<==o †Paoliida Handlirsch, 1906 sensu Prokop, Krzeminski, Krzeminska & Wojciechowski, 2012 [Protoptera Sharov, 1966; Paoliida Handlirsch; Rohdendorff, 1977] |?- †Ampeliptera limburgica Pruvost, 1927 |?- †Stygne roemeri Handlirsch, 1906 [type lost in WWII] |?- †Katerinka candida Prokop & Nel, 2007 |--o †Anthracoptilidae Handlirsch, 1922 sensu Guan, Prokop, Roques, Lapeyrie & Nel, 2015 [Strephocladidae Martynov, 1938; Strephoneuridae Martynov, 1940; ?Adiphlebiidae Handlirsch, 1906] | |?- †Adiplebia lacoana Scudder, 1885 [Goldenbergia longitudianalis (Scudder, 1885) Brongniart, 1885] | |-- †Anthracoptilus perrieri (Meunier, 1909) Lameere, 1917 | |--o †Carrizocladus Rasnitsyn, in Rasnitsyn et al., 2004 | | |-- †C. arroyo Rasnitsyn, in Rasnitsyn et al., 2004 | | `?- ?†C. deter Rasnitsyn, in Rasnitsyn et al., 2004 | |-- †Graticladus severus Novokshonov & Aristov, 2004 | |--o †Homocladus Carpenter, 1966 | | |-- †H. grandis Carpenter, 1966 [incl.? Paracladus retardatus Carpenter, 1966] | | `?- †H. ornatus Carpenter, 1966 [=? †H. grandis] | |--o †Jarmilacladus Rasnitsyn & Aristov, 2004 | | |-- †J. variabilis Rasnitsyn & Aristov, 2004 | | `?- ?†J. paties Rasnitsyn & Aristov, 2004 | |--o †Mesoptilus Lameere, 1917 | | |-- †M. dolloi Lameere, 1917 | | `-- †M. carpenteri Guan, Prokop, Roques, Lapeyrie & Nel, 2015 | |?- †Mycteroptila dina Rasnitsyn, 1977 [=? Spargoptilon] | |-- Pruvostia spectabilis Bolton, 1921 | |-- †Pseudomesophilus sellardsi (Lameere, 1917) Guan, Prokop, Roques, Lapeyrie & Nel, 2015 [Mesoptilus sellardsi Lameere, 1917] | |?- †Rhinomaloptila polyneura Rasnitsyn, 1977 | |--o †Spargoptilon Kukalová, 1965 | | |-- †S. latericus Kukalová, 1965 | | `-- †S. confertus Kukalová, 1965 | |--o †Strephocladus Scudder, 1885 | | |-- †S. subtilis (Kliver, 1883) | | `-- †S. permianus Guan, Prokop, Roques, Lapeyrie & Nel, 2015 | |--o †Strephoneura Martynov, 1940 | | `-- †S. tenebrosa Novokshonov & Aristov, 2004 | |--o †Strephophilus Rasnitsyn & Aristov, 2013 | | |-- †S. borealis Rasnitsyn & Aristov, 2013 | | `-- †S. longus Rasnitsyn & Aristov, 2013 | `-- †Westphaloptilus gallicus Guan, Prokop, Roques, Lapeyrie & Nel, 2015 `--+--o †Blattinopsidae Bolton, 1925 sensu Prokop, Krzeminski, Krzeminska, Hörnschemeyer, Ilger, Brauckmann, Grandcolas & Nel, 2013 | `-- Protoblattinopsis stubblefieldi Laurentiaux, 1953 `--o †Paoliidae Handlirsch, 1906 sensu Prokop, Krzeminski, Krzeminska & Wojciechowski, 2012 [incl. Ideliidae Zalessky, 1928; Herbstialidae Schmidt, 1953] |-- †Acropermula Aristov &: Storozhenko, 2011; Perm. CEu. [Ideliidae] |-- †Aenigmidelia Sharov, 1961; Perm. NCAs. [Ideliidae] |-- †Archidelia Sharov, 1961; Perm. NCAs. [Ideliidae] |-- †Darekia sanquinea Prokop, Krzeminski, Krzeminska & Wojciechowski, 2012 |-- †Herbstiala herbsti Schmidt, 1953; Dev. CEu. [Herbstialidae] |--o †Holasicia Kukalová, 1958 | |-- †H. vetula Kukalová, 1958 | `-- †H. rasnitsyni Brauckmann, 1984 |-- †Kemperala hagensis Brauckmann, 1984 |-- †Kochopteron hoffmannorum Brauckmann, 1984 |-- †Mertovia sustai (Kukalová, 1958) Prokop & Nel, 2000 |-- †Micaidelia Aristov, 2004; Perm. EEu. [Ideliidae] |-- †Mongoloidelia Storozhenko, 1992; Perm. EAs. [Ideliidae] |-- †Olinka modica Kukalová, 1958 |-- †Paolia vetusta Smith, 1871 [type] |-- †Paoliola gurleyi Handlirsch, 1919 |-- †Protoblattina bouvieri Meunier, 1909 |-- †Pseudofouquea cambrensis (Allen, 1901) Handlirsch, 1906 |-- †Silesiapteron jarmilae Prokop, Krzeminski, Krzeminska, Hörnschemeyer, Ilger, Brauckmann, Grandcolas & Nel, 2013 |-- †Sojanidelia Storozhenko, 1992; Perm. EEu. [Ideliidae] |-- †Stenaropodites Martynov, 1928; Perm. EEu. [Ideliidae] |-- †Sustaia impar Kukalová, 1958 [Sustainia (lapus calami)] |-- †Sylvidelia Martynov, 1940; Perm. EEu. [Ideliidae] `--o †Zdenekia Kukalová, 1958 |-- †Z. grandis Kukalová, 1958 |-- †Z. occidentalis Laurentiaux-Vieira & Laurentinaux, 1986 `-- †Z. silensiensis Prokop, Krzeminski, Krzeminska & Wojciechowski, 2012
Reference(s):
- Guan, Z., Prokop, J., Roques, P., Lapeyrie, J. & Nel, A., 2015: Revision of the Enigmatic Insect Family Anthracoptilidae Enlightens the Evolution of Palaeozoic Stem-Dictyopterans.
–Acta Palaeontologica Polonica: Vol. 61, #1, pp. 71-87 [doi: 10.4202/app.00051.2014] - Prokop, J., Krzeminski, W., Krzeminska, E., Hörnschemeyer, T., Ilger, J.-M., Brauckmann, C., Grandcolas, P. & Nel, A., 2013: Late Palaeozoic Paoliida is the sister group of Dictyoptera (Insecta: Neoptera).
–Journal of Systematic Palaeontology: Vol. 12, #5, pp. 601-622 [doi: 10.1080/14772019.2013.823468] - Prokop, J., Roques, P. & Nel, A., 2014: New non-holometabolous insects from the Pennsylvanian of the Avion locality in Pas-de-Calais, France (Insecta: ‘Exopterygota’).
–Alcheringa: An Australasian Journal of Palaeontology: Vol. 38, #2, pp. 155-169 [doi: 10.1080/03115518.2014.848620] - Prokop, J., Krzeminski, W., Krzeminska, E. & Wojciechowski, D., 2012: Paoliida, a Putative Stem-Group of Winged Insects: Morphology of New Taxa from the Upper Carboniferous of Poland.
–Acta Palaeontologica Polonica: Vol. 57, #1, pp. 161-173 [doi: 10.4202/app.2010.0064]