After Bechly, 1998-2002 & Schorr, Lindeboom & Paulson, 2003
<==o Megapodagrionidae Tillyard, 1917 ()?
|?- Agnophilogenia monotis Kennedy, 1941
|?- Agriomorpha fusca May, 1933
|?- Allolestes maclachlani Selys, 1869
|?-o Allopodagrion Förster, 1910 ()? [11]
| |-- A. contortum (Hagen in Selys, 1862) [Podagrion contortum Hagen in Selys, 1862]
| |-- A. curtum Selys, 1886
| |-- A. erinys Ris, 1913
| |-- A. lepidum Rácenis, 1959
| |-- A. macropus (Selys, 1862) [Podagrion macropus Selys, 1862]
| |-- A. mercenarium (Hagen, 1869) [Podagrion mercenarium Hagen, 1869; Podagrion nebulosum Selys, 1886]
| |-- A. muzana (Navás, 1934) [Argia muzana Navás, 1934]
| |-- A. oscillans (Selys, 1862) [Podagrion oscillans Selys, 1862]
| |-- A. setigerum Selys, 1886
| |-- A. temporale (Selys, 1862) [Podagrion temporale Selys, 1862]
| `-- A. venale (Selys, 1862) [Podagrion venale Selys, 1862]
|?- Amanipodagrion gilliesi Pinhey, 1962
|?-o Archaeopodagrion Kennedy, 1939 2
| |-- A. bicorne Kennedy, 1939
| `-- A. bilobatum Kennedy, 1946
|?-o Archiargiolestes Kennedy, 1925 ()? [3]
| |-- A. parvulus (Watson, 1977) [Argiolestes parvulus Watson, 1977]
| |-- A. pusillissimus Kennedy, 1925
| `-- A. pusillus (Tillyard, 1908) [Argiolestes pusillus Tillyard, 1908]
|?-o Arrhenocnemis Lieftinck, 1933 ()? [2]
| |-- A. amphidactylis Lieftinck, 1949
| `-- A. sinuatipennis Lieftinck, 1933
|?-o Austroargiolestes Kennedy, 1925 ()? [10]
| |-- A. chrysoides (Tillyard, 1913) [Argiolestes chrysoides Tillyard, 1913]
| |-- A. alpinus (Tillyard, 1913) [Argiolestes alpinus Tillyard, 1913]
| |-- A. amabilis (Förster, 1899) [Argiolestes amabilis Förster, 1899]
| |-- A. aureus (Tillyard, 1906) [Argiolestes aureus Tillyard, 1906]
| |-- A. brookhousei Theischinger & O'Farrell, 1986
| |-- A. calcaris (Fraser, 1958) [Argiolestes calcaris Fraser, 1958]
| |-- A. christine Theischinger & O'Farrell, 1986
| |-- A. elke Theischinger & O'Farrell, 1986
| |-- A. icteromelas (Selys, 1862) [Argiolestes icteromelas Selys, 1862; Argiolestes calcaris tenuis Fraser, 1959 (nec tenuis Tillyard, 1913); Austroargiolestes icteromelas nigrolabiatus Theischinger & O'Farrell, 1986]
| `-- A. isabellae Theischinger & O'Farrell, 1986
|?- Bornargiolestes niger Kimmins, 1936
|?-o Burmargiolestes Kennedy, 1925 ()? [3]
| |-- B. laidlawi Lieftinck, 1960
| |-- B. melanothorax (Selys, 1891) [Argiolestes melanothorax Selys, 1891; B. flaviceps Fraser, 1933]
| `-- B. xinglongensis Wilson & Reels, 2001
|?- Caledargiolestes Kennedy, 1925 ()? [2]
| |-- C. janiceae Lieftinck, 1975
| `-- C. uniseries (Ris, 1915) [Argiolestes uniseries Ris, 1915]
|?-o Caledopteryx Kennedy, 1925 ()? [2]
| |-- C. maculata Winstanley & Davies, 1982
| `-- C. sarasini (Ris, 1915) [Argiolestes sarasini Ris, 1915]
|?- Calilestes pallidistigma Fraser, 1926
|?- Celebargiolestes cinctus (Selys, 1886) Kennedy, 1925 [Argiolestes cincta Selys, 1886]
|?-o Dimeragrion Calvert, 1913 ()? [4]
| |-- D. mesembrium De Marmels, 1989
| |-- D. percubitale Calvert, 1913
| |-- D. secundum Needham, 1933
| `-- D. unturanense De Marmels, 1992
|?-o Griseargiolestes Theischinger, 1998 ()? [7]
| |-- G. albescens (Tillyard, 1913) [Argiolestes albescens Tillyard, 1913]
| |-- G. bucki Theischinger, 1998
| |-- G. eboracus (Tillyard, 1913) [Argiolestes eboracus Tillyard, 1913]
| |-- G. fontanus (Tillyard, 1913) [Argiolestes fontanus Tillyard, 1913]
| |-- G. griseus (Hagen in Selys, 1862) [Argiolestes griseus Hagen in Selys, 1862; Argiolestes subgriseus Fraser, 1959; Argiolestes tenuis Tillyard, 1913]
| |-- G. intermedius (Tillyard, 1913) [Argiolestes intermedius Tillyard, 1913]
| `-- G. metallicus (Sjöstedt, 1917) [Argiolestes metallicus Sjöstedt, 1917]
|?-o Heteropodagrion Selys, 1885 ()? [2]
| |-- H. sanguinipes Selys, 1895
| `-- H. superbum Ris, 1918
|?- Mesagrion leucorhinum Selys, 1885
|?- Mesopodagrion tibetanum McLachlan, 1896 [M. yacohwensis Chao, 1953]
|?- Miniargiolestes minimus (Tillyard, 1908) Theischinger, 1998 [Argiolestes minimus Tillyard, 1908]
|?-o Nesolestes Selys, 1891 ()? [17]
| |-- N. albicaudus Fraser, 1952
| |-- N. albicolor Fraser, 1955
| |-- N. alboterminatus Selys, 1891
| |-- N. angydna Schmidt, 1951
| |-- N. drocera Fraser, 1951
| |-- N. elizabethae Lieftinck, 1965
| |-- N. forficuloides Fraaser, 1955
| |-- N. mariae Aguesse, 1968
| |-- N. martini Schmidt, 1951
| |-- N. nigeriensis Gambles, 1970
| |-- N. pauliani Fraser, 1951
| |-- N. pulverulans Lieftinck, 1965
| |-- N. radama Lieftinck, 1965
| |-- N. ranavalona Schmidt, 1951
| |-- N. robustus Aguesse, 1968
| |-- N. rubristigma Martin, 1903
| `-- N. tuberculicollis Fraser, 1949
|?- Neuragrion mysticum Karsch, 1891
|?- Neurolestes trinervis Selys, 1885
|?-o Oxystigma Selys, 1862 ()? [4]
| |-- O. caerulans De Marmels, 1987
| |-- O. cyanofrons Williamson, 1919
| |-- O. petiolatum (Selys, 1862) [Heteragrion petiolatum Selys, 1862]
| `-- O. williamsoni Geijskes, 1976
|?-o Paraphlebia Selys, 1862 ()? [4]
| |-- P. duodecima Calvert, 1901
| |-- P. hyalina Brauer, 1871
| |-- P. quinta Calvert, 1901 [P. abrogata Calvert, 1907]
| `-- P. zoe Selys, 1862
|?-o Podolestes Selys, 1862 ()? [8]
| |-- P. atomarius Lieftinck, 1950
| |-- P. buwaldai Lieftinck, 1940
| |-- P. chrysopus Selys, 1886
| |-- P. coomansi Lieftinck, 1940
| |-- P. furcifer Lieftinck, 1950
| |-- P. harrissoni Lieftinck, 1953
| |-- P. orientalis Selys, 1862
| `-- P. pandanus Wilson & Reels, 2001
|?-o Podopteryx Selys, 1871 ()? [2]
| |-- P. casuarina Lieftinck, 1949
| `-- P. selysi (Förster, 1899) [Argiolestes selysi Förster, 1899; P. roseonotata Selys, 1871]
|?-o Priscagrion Zhou & Wilson, 2001 ()? [2]
| |-- P. kiautai Zhou & Wilson, 2001
| `-- P. pinheyi Zhou & Wilson, 2001
|?-o Protolestes Fraser, 1899 8
| |-- P. fickei Förster, 1899
| |-- P. furcatus Aguesse, 1967
| |-- P. kerckhoffae Schmidt in Fraser, 1949
| |-- P. leonorae Schmidt, 1951
| |-- P. milloti Fraser, 1949
| |-- P. proselytus Lieftinck, 1965
| |-- P. rufescens Aguesse, 1967
| `-- P. simonei Aguesse, 1967
|?- Pseudolestes mirabilis Kirby, 1900
|?-o Rhinagrion Calvert, 1913 ()? [7]
| |-- R. borneense (Selys, 1886) [Amphilestes borneensis Selys, 1886]
| |-- R. elopurae (McLachlan in Selys, 1886) [Amphilestes eloporae McLachlan in Selys, 1886]
| |-- R. hainanense Wilson & Reels, 2001
| |-- R. macrocephalum (Selys, 1862) [Amphilestes macrocephala Selys, 1862]
| |-- R. mima (Karsch, 1891) [Amphilestes mima Karsch, 1891; R. viridata Fraser, 1938]
| |-- R. philippinum (Selys, 1882) [Amphilestes philippina Selys, 1882]
| `-- R. tricolor (Krüger, 1898) [Amphilestes tricolor Krüger, 1898]
|?-o Sciotropis Rácenis, 1959 ()? [2]
| |-- S. cyclanthorum Rácenis, 1959
| `-- S. lattkei De Marmels, 1994
|?-o Tatocnemis Kirby, 1889 ()? [10]
| |-- T. crenulatipennis Fraser, 1952
| |-- T. denticularis Aguesse, 1968
| |-- T. emarginatipennis Fraser , 1960
| |-- T. malgassica Kirby, 1889
| |-- T. mellisi Schmidt, 1951
| |-- T. micromalgassica Aguesse, 1968
| |-- T. olstieffi Schmidt, 1951
| |-- T. robinsoni Schmidt, 1951
| |-- T. sinuatipennis (Selys, 1891) [Nesocnemis sinuatipennis Selys, 1891]
| `-- T. virginiae Legrand, 1992
|?- Trineuragrion percostale Ris, 1915
|--o Agriolestinae Fraser, 1957 sensu Bechly, 1996 ()?
| |== other Agriolestines [genera not listed in Bechly, 1998-2002, Schorr, Lindeboom & Paulson (2003) don't use subfamilies or other levels between family & genus]
| `-- Argiolestes Selys, 1862 ()? [37]
`--o Megapodagrioninae Tillyard, 1917 sensu Bechly, 1996 ()?
|== other Agriolestines [genera not listed in Bechly, 1998-2002, Schorr, Lindeboom & Paulson (2003) don't use subfamilies or other levels between family & genus]
`-- Megapodagrion megalopus (Selys, 1862) Selys, 1885 [Podagrion megalopus Selys, 1862; M. arachne Rácenis, 1959]
Reference(s):
Bechly, G., 1998-2002a: Phylogenetic classification of fossil & extant odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm
Bechly, G., 1998-2002b: Phylogenetic Systematics of Zygoptera / Euzygoptera.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/zygopt2.htm
Schorr, M., Lindeboom, M. & Paulson, D., 2003: World Species List - Odonata.
–iNet: International Dragonfly Fund: http://www.calopteryx.de/idf/
