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<==o Hypolestidae Tillyard & Fraser, 1938 sensu Bechly, 1996 |--o Heteragrioninae Racenis, 1959 sensu Bechly, 1996 | |== other Heteragrionines? [Bechly, 1998-2002] | `-- Heteragrion Selys, 1862 ()? [41] |--o Philogeniinae | |== other Philogeniines? [Bechly, 1998-2002] | `-- Philogenia Selys, 1862 ()? [34] `--o Hypolestinae Tillyard & Fraser, 1938 |--o Philosini Kennedy, 1925 sensu Bechly, 1996 | |-- †Eolestes synthetica Cockrell, 1940 [non Eolestes Bown & Schankler, 1982, an Eocene hedgehog, jr. homonym] | |-- †Prohypolestes dauphinensis Nel & Paicheler, 1994 | |--o Philosina Ris, 1917 ()? [2] | | |-- P. alba Wilson, 1999 | | `-- P. buchi Ris, 1917 | `--+-- Lestomima flavostigma May, 1933 | `--o Rhipidolestes Ris, 1912 ()? [17] | |-- R. aculeatus Ris, 1912 | |-- R. alleni Wilson, 2000 | |-- R. apicatus Navás, 1934 | |-- R. asatoi Asahina, 1994 | |-- R. bastiaani Zhu and Yang, 1998 | |-- R. bidens Schmidt, 1931 | |-- R. cyanoflavus Wilson, 2000 | |-- R. hiraoi Yamamoto, 1955 | |-- R. janetae Wilson, 1997 | |-- R. jucundus Lieftinck, 1948 | |-- R. malaisei Lieftinck, 1948 | |-- R. nectans (Needham, 1929) [Taolestes nectans Needham, 1929] | |-- R. okinawanus Asahina, 1951 [Argia apicalis Matsumura (nec Say, 1839), 1913 (tentative, would have priority)] | |-- R. owadai Asahina, 1997 | |-- R. rubripes (Navás, 1936) [Taolestes rubripes Navás, 1936] | |-- R. truncatidens Schmidt, 1931 | `-- R. yangbingi Davies, 1998 |--o Hypolestes Gundlach, 1888 ()? [2] [Hypolestini Tillyard & Fraser, 1938 sensu Bechly, 1996] | |-- H. clara (Calvert, 1891) [Ortholestes clara Calvert, 1891] | `-- H. trinitatis (Gundlach, 1888) [Lestes trinitatis Gundlach, 1888; Ortholestes abbotti Calvert, 1894] `--o Lestoidea Tillyard, 1913 ()? [4] [Lestoideini Munz, 1919 sensu Bechly, 1996] |-- L. barbarae Watson, 1967 |-- L. brevicauda Theischinger, 1996 |-- L. conjuncta Tillyard, 1913 `-- L. lewisiana Theischinger, 1996
Note: Schorr, Lindeboom & Paulson, 2003 don't regognize family Hypolestidae and they list all these genera to family Megapodagrionidae.
Because I have decided to follow Bechly (1998-2002) in this presentation, I have thus inclueded those generas from Schorr, et al. that Bechly lists to Hypolestidae.
Bechly, G., 1998-2002a: Phylogenetic classification of fossil and extant odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm
Bechly, G., 1998-2002b: Phylogenetic Systematics of Zygoptera / Euzygoptera.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/zygopt2.htm
Schorr, M., Lindeboom, M. & Paulson, D., 2003: World Species List - Odonata.
–iNet: International Dragonfly Fund: http://www.calopteryx.de/idf/
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