Mikko's Phylogeny Archive Main Groups of Insects   Calopterygidae

After Bechly, 1998-2002 and Schorr, Lindeboom & Paulson, 2003

<==o Calopterygidae Selys, 1850 [Agriidae] (neidonkorennot)
   |-- Hetarina Hagen in Selys, 1853 ()? [incl. Mnesarete Cowley, 1934 sensu Bechly, 1998-2002] [Hetaerininae Tillyard & Fraser, 1939] []
   `--o Calopteryginae Selys, 1850
      |--o Archineura Kirby, 1894 ()? [2]
      |  |-- A. hetaerinoides (Fraser, 1933) [Leucopteryx hetaerinoides Fraser, 1933]
      |  `-- A. incarnata (Karsch, 1891) [Echo incarnata Karsch, 1891; A. basilactea Kirby, 1894]
      |-- Calopteryx Leach, 1815 ()? [29] [for detailed taxonomic history, see Bechly, 1998-2002b]
      |--o Echo Selys, 1853 ()? [4]
      |  |-- E. margarita Selys, 1853
      |  |-- E. maxima Martin, 1904
      |  |-- E. modesta Laidlaw, 1902 [Climacobasis lugens Laidlaw, 1902]
      |  `-- E. uniformis Selys, 1879
      |--o Iridictyon Needham and Fisher, 1940 ()? [2]
      |  |-- I. myersi Needham and Fisher, 1940
      |  `-- I. trebbaui Rácenis, 1968
      |--o Matrona Selys, 1853 ()? [2]
      |  |-- M. basilaris Selys, 1853 [Calopteryx okinawana Matsumura, 1931; M. japonica Förster, 1897; M. kricheldorffi Karsch, 1891; M. nigripectus Selys, 1879]
      |  `-- M. cyanoptera Hämäläinen and Yeh, 2000
      |--o Mnais Selys, 1853 ()? [9]
      |  |-- M. andersoni McLachlan in Selys, 1873 [M. earnshawi Williamson, 1904]
      |  |-- M. gregoryi Fraser, 1924 
      |  |-- M. incolor Martin, 1921 
      |  |-- M. maclachlani Fraser, 1924 
      |  |-- M. mneme Ris, 1916 
      |  |-- M. nawai Yamamoto, 1956 
      |  |-- M. pruinosa Selys, 1853 [M. strigata Hagen in Selys, 1853]
      |  |-- M. semiopaca May, 1935 
      |  `-- M. tenuis Oguma, 1913 [M. auripennis Needham, 1930; M. pieli Navas, 1936]
      |--o Neurobasis Selys, 1853 ()? [11]
      |  |-- N. anderssoni Sjöstedt, 1926 
      |  |-- N. anumariae Hämäläinen, 1989 
      |  |-- N. australis Selys, 1878 
      |  |-- N. chinensis (Linnaeus, 1758) [Libellula chinensis Linnaeus, 1758; Calopteryx disparilis Rambur, 1842]
      |  |-- N. cyaneipennis (Förster, 1897) [Matrona cyaneipennis Förster, 1897]
      |  |-- N. daviesi Hämäläinen, 1993 
      |  |-- N. ianthinipennis Lieftinck, 1949 
      |  |-- N. kaupi Brauer, 1867 
      |  |-- N. kimminsi Lieftinck, 1955 
      |  |-- N. longipes Hagen, 1887 
      |  `-- N. luzoniensis Selys, 1879  
      |--o Phaon Selys, 1853 ()? [2]
      |  |-- P. camerunensis Sjöstedt, 1900 [P. fraseri Pinhey, 1962]
      |  `-- P. iridipennis (Burmeister, 1839) [Calopteryx iridipennis Burmeister, 1839; P. fuliginosus Selys, 1879; P. rasoherinae Fraser, 1949; Sapho occidentalis Förster, 1906]
      |-- Psolodesmus mandarinus McLachlan, 1870 [incl. P. dorothea Williamson, 1904]
      |--o Sapho Selys, 1853 ()? [6]
      |  |-- S. bicolor Selys, 1853 [S. superba Sjöstedt, 1917]
      |  |-- S. ciliata (Fabricius, 1781) [Agrion ciliata Fabricius, 1781]
      |  |-- S. fumosa Longfield, 1932
      |  |-- S. gloriosa McLachlan in Selys, 1873
      |  |-- S. orichalcea McLachlan, 1869 [S. venusta Karsch, 1889]
      |  `-- S. puella (Sjöstedt, 1917) [Umma puella Sjöstedt, 1917]
      |--o Umma Kirby, 1890 ()? [11]
      |  |-- U. cincta (Hagen in Selys, 1853) [Cleis cincta Hagen in Selys, 1853]
      |  |-- U. declivium Förster, 1906 
      |  |-- U. distincta Longfield, 1933 
      |  |-- U. electa Longfield, 1933 
      |  |-- U. femina Longfield, 1947 
      |  |-- U. infumosa Fraser, 1951 
      |  |-- U. longistigma (Selys, 1869) [Sapho longistigma Selys, 1869]
      |  |-- U. mesostigma (Selys, 1879) [Cleis mesostigma Selys, 1879; Umma fuscomarginis Sjöstedt, 1899; Umma splendida Navás, 1922]
      |  |-- U. mesumbei Vick, 1996 
      |  |-- U. purpurea Pinhey, 1961 
      |  `-- U. saphirina Förster, 1916
      `--o Vestalis Selys, 1853 ()? [17]
         |-- V. amabilis Lieftinck, 1965
         |-- V. amaryllis Lieftinck, 1965
         |-- V. amethystina Lieftinck, 1965
         |-- V. amnicola Lieftinck, 1965
         |-- V. amoena Selys, 1853
         |-- V. anacolosa Lieftinck, 1965
         |-- V. anne Hämäläinen, 1985 
         |-- V. apicalis Selys, 1873
         |-- V. atropha Lieftinck, 1965
         |-- V. beryllae Laidlaw, 1915
         |-- V. gracilis (Rambur, 1842) [Calopteryx gracilis Rambur, 1842]
         |-- V. luctuosa (Burmeister, 1839)[Calopteryx luctuosa Burmeister, 1839]
         |-- V. lugens Albarda in Selys, 1879
         |-- V. melania Selys, 1873
         |-- V. miao Wilson & Reels, 2001
         |-- V. smaragdina Selys, 1879
         `-- V. yunosukei Asahina, 1990

Reference(s):

Bechly, G., 1998-2002a: Phylogenetic classification of fossil and extant odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm

Bechly, G., 1998-2002b: Phylogenetic Systematics of Zygoptera / Caloptera.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/zygopt1.htm

Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut‚ Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968.

Schorr, M., Lindeboom, M. & Paulson, D., 2003: World Species List - Odonata.
–iNet: International Dragonfly Fund: http://www.calopteryx.de/idf/