After Bechly, 1998-2002, Nel, Petrulevicius & Martínez-Delclós, 2005, and Nel, Bechly, Prokop, Béthoux & Fleck, 2011
<==o †Protozygoptera Tillyard, 1925 [Archisozygoptera Handlirsch 1906 sensu Grimaldi & Engel, 2005] |--o †Permagrionoidea Tillyard, 1928 sensu Nel, Bechly, Prokop, Béthoux & Fleck, 2011 | `--o †Permagrionidae Tillyard, 1928 sensu Nel, Bechly, Prokop, Béthoux & Fleck, 2011 [incl. Permolestidae Martynov, 1932; Solikamptilonidae Zalessky, 1948] | |-- †Permagrion falklandicum Tillyard, 1928 [Permagrionidae Tillyard, 1928 sensu Pritykina, 1980] | `--o †Permolestidae Martynov, 1932 | |--o †Permolestes Martynov, 1932 | | |-- †P. gracilis Martynov, 1932 | | |-- †P. sheimogorai Nel, Bechly, Prokop, Béthoux & Fleck, 2011 | | `-- †P. soyanaiensis Nel, Bechly, Prokop, Béthoux & Fleck, 2011 | |--o †Epilestes Martynov, 1937 | | |-- †E. kargalensis Martynov, 1937 | | |-- †E. gallica Nel et al., 1999 | | `-- †E. angustapterix Nel, Bechly, Prokop, Béthoux & Fleck, 2011; 87. [E. angustus Nel et al., 2011; 86. (lapsus calami)] | |-- †Scytolestes stigmalis Martynov, 1937 | `--o †Solikamptilon Zalessky, 1948 [Solikamptilonidae Zalessky, 1948] | |-- †S. remuliforme Zalessky, 1948 | `-- †S. pectinatus Nel, Bechly, Prokop, Béthoux & Fleck, 2011 `--+-- Panodonata `--o †Archizygoptera Handlirsch, 1906 sensu Bechly, 1996 |-- †Permepallage angustissima Martynov, 1938 [Permepallagidae Martynov, 1938] |-- †Bechlya ericrobinsoni Jarzembowski & Nel, 2002 [Bechlyidae Jarzembowski & Nel, 2002] |-- †Lodevia longialata Nel, Gand, Garric & Lapeyrie, 1999 [Lodeviidae Nel, Bechly, Prokop, Béthoux & Fleck, 2011] [originally classified as a Permepallagid (Bechly, 1998-2002)] |-- †Luiseia breviata Nel, Bechly, Prokop, Béthoux & Fleck, 2011 [Luiseiidae Nel, Bechly, Prokop, Béthoux & Fleck, 2011] |--o †Kennedyidae Tillyard, 1925 [Kennedyoidea (Tillyard, 1925) Pritykina, 1981] | |?-o †Sushkinia Martynov, 1930 [sensu Nel, Gand, Garric & Lapeyrie, 1999] | | |-- S. parvula Martynov, 1930 | | `-- S. elongata Martynov, 1930 | |--o †Kennedya Tillyard, 1925 | | |-- †K. mirabilis Tillyard, 1925 | | |-- †K. reducta Carpenter, 1939 | | |-- †K. fraseri Carpenter, 1947 | | |-- †K. carpenteri Pritykina, 1981 | | |-- †K. gracilis Pritykina, 1981 | | |-- †K. sp. (Tasch & Zimmerman, 1959, 1962; Nel, Bechly, Prokop, Béthoux & Fleck, 2011) | | |-- †K. azari Nel, Bechly, Prokop, Béthoux & Fleck, 2011 | | |-- †K. pritykinae Nel, Bechly, Prokop, Béthoux & Fleck, 2011 | | `-- †K. ivensis Nel, Bechly, Prokop, Béthoux & Fleck, 2011 | `--o †Progoneura Carpenter, 1931 | |-- †P. minuta Carpenter, 1931 | |-- †P. nobilis Carpenter, 1947 | |-- †P. venula Carpenter, 1947 | |-- †P. sp. Carpenter, 1947 | `-- †P. grimaldii Nel, Bechly, Prokop, Béthoux & Fleck, 2011 |-- †Kaltanoneura bartenevi Rohdendorf, 1961 [Kaltanoneuridae Rohdendorf, 1961] |-- †Oboraneura kukalovae Zessin, 2008 [Oboraneuridae Zessin, 2008] |--o †Bakteniidae Pritykina, 1981 [Batkeniinae (Pritykina, 1981) Bechly, 1996] | |-- †Batkenia pusilla Pritykina, 1981 | |-- †Paratrissoneura primitiva (Pritykina, 1981) Bechly, 1997 ["Trissoneura" primitiva Pritykina, 1981] | `-- †Engellestes chekardensis Nel, Bechly, Prokop, Béthoux & Fleck, 2011 `--o †Protomyrmeleontoidea (Handlirsch, 1906) Bechly, 1996 |--o Voltzialestidae (Bachly, 1996) Nel, Bechly, Prokop, Béthoux & Fleck, 2011 [Bakteniidae: Voltzialestinae Bechly, 1998] | |-- †Azaroneura permiana Nel, Bechly, Prokop, Béthoux & Fleck, 2011; Perm. | |-- †Voltzialestes triasicus Nel, et al., 1996; Perm. | `--o †Terskeja Pritykina, 1981 | |-- †T. paula Pritykina, 1981 | |-- †T. pumilio Pritykina, 1981 | `-- †T. tenuis Pritykina, 1981 `--o †Protomyrmeleontidae Handlirsch, 1906 |-- †Triassagrion australiense Tillyard, 1922 [Triassagrioninae (Tillyard, 1922) Bechly, 1996; Triassagrionidae Tillyard, 1922] Trias. `--o †Protomyrmeleontinae (Handlirsch, 1906) Bechly, 1996 |-- †Ferganagrion kirghiziensis Nel, Petrulevicius & Martínez-Delclós, 2005; Trias. |-- †Italomyrmeleon Bechly, 1997; Trias. |-- †Tillyardomyrmeleon Henrotay et al., 1997; Trias. |-- †Obotritagrion Zessin, 1991; E.Jur. |-- †Paraobotritagrion tenuiformis (Zessin, 1991) Nel, Petrulevicius & Martínez-Delclós, 2005; E.Jur. |-- †Mongolagrion shartegensis Nel, Petrulevicius & Martínez-Delclós, 2005; L.Jur. |--o †Protomyrmeleon Geinitz, 1887; E.Jur.-E.Cret. | |-- †P. grandis Nel, Petrulevicius & Martínez-Delclós, 2005 | |-- †P. karatauensis Nel, Petrulevicius & Martínez-Delclós, 2005 | |-- †P. kazakhstanensis Nel, Petrulevicius & Martínez-Delclós, 2005 | |-- †P. pumilio Nel, Petrulevicius & Martínez-Delclós, 2005 | `?- †P. sp. indet. (Nel, Petrulevicius & Martínez-Delclós, 2005) |?- †Malmagrion eichstaettense (Hagen, 1862) [sensu Bechly, 1998] |-- †Malmomyrmeleon Martínez-Delclos & Nel, 1996 (Late Jurassic), `-- †Saxomyrmeleon Nel &: Jarzembowski, 1998; E.Cret.
Reference(s):
- Bechly, G., 1998-2002a: Phylogenetic classification of fossil and extant odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm - Bechly, G., 1998-2002b: Phylogenetic Systematics of Stem-Group Odonates / Discoidalia.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/odonato2.htm - Beckemeyer, R. J. 2006-: Checklist of Midco, Oklahoma Permian insects. [visited: 2006-08-04]
–iNet: www.windofkansas.com: http://www.windsofkansas.com/midco_taxa.html - Nel, A., Bechly, G., Prokop, J., Béthoux, O. & Fleck, G., 2011: Systematics and Evolution of Paleozoic And Mesozoic Damselfly-Like Odonatoptera of the 'Protozygopteran' Grade.
–Journal of Paleontology: Vol. 86, #1, pp. 81-104 [doi: 10.1666/11-020.1]
–Journal of Systematic Palaeontology: Vol. 3, #2, pp. 187-201 [doi: 10.1017/S1477201905001549]