After Bechly, 1998-2002, and Grimaldi & Engel, 2005
<==o †Meganisoptera Martynov, 1932 [†Protodonata in partim sensu Grimaldi & Engel, 2005] (griffenflies; griippikorennot) |-- †Namurotypus sippeli Brauckmann & Zessin, 1989 [Namurotypidae Bechly, 1996] `--o †Meganeuromorpha Pritykina, 1980 sensu Bechly, 1996 (Meganeuramaiset sudenkorennot) |--o †Paralogidae Handlirsch, 1906 | |--o †Paralogus Scudder, 1893 | | |-- †P. aeschnoides | | `-- †P. hispanicus | |-- †Truemania multiplicata (Bolton, 1922) Bolton, 1934 | `--o †Oligotypus Carpenter, 1931 [Sinomeganeura Prokop, Nel & Ren, 2008] | |-- †O. huangheensis (Prokop, Nel & Ren, 2008) [Sinomeganeura huangheensis Prokop, Nel & Ren, 2008] | |-- †O. makowskii Carpenter & Richardson, 1971 | |-- †O. tillyardi Carpenter, 1931 | `-- †O. tuscaloosae Beckemeyer & Engel, 2011 |-- †Kargalotypus kargalensis (Martynov, 1932) Rohdendorf, 1962 [Megatypus kargalensis Martynov, 1932] [Kargalotypidae Zessin, 1983] |--o †Kohlwaldiidae Guthörl, 1962 sensu Bechly, 1996 | |-- †Solutotherates analis (Carpenter, 1981) | |-- †Kohlwaldia kuehni Guthörl, 1962 | `-- †Boltonites radstockensis (Bolton, 1914) Handlirsch, 1919 `--o †Meganeuridae Handlirsch, 1906 |-- †Carpentertypus durhami Zessin, 1983 [Carpentertypinae Zessin, 1983] |--o †Tupinae Handlirsch, 1919 sensu Bechly, 1996 [Typinae Handlirsch, 1919] | |--o †Arctotypus Martynov, 1932 | | |-- †A. carbonicus | | |-- †A. diluculum | | |-- †A. fortis | | |-- †A. gallicus | | |-- †A. giganteus | | |-- †A. intermedius | | |-- †A. merifonsensis | | |-- †A. sinuatus | | `-- †A. sylvaensis | |-- †Bohemiatupus elegans Prokop & Nel, 2010 | |--o †Curvitupus Nel, Fleck, Garrouste, Gand, Lapeyrie, Bybee & Prokop, 2009 | | |-- †C. ariegensis | | |-- †C. elongatus | | |-- †C. magnificus | | `-- †C. verneti | |-- †Ephemerites rueckerti Geinitz, 1865 | |-- †Gilsonia titana Meunier, 1909 [Meganeurula titana] | |-- †Meganeurina confusa Handlirsch, 1919 | |--o †Megatypus Tillyard, 1925 | | |-- †M. ingentissimus | | |-- †M. parvus | | |-- †M. schucherti | | `-- †M. vetustus | |-- †Nannotupus pumilio Nel, Fleck, Garrouste, Gand, Lapeyrie, Bybee & Prokop, 2009 | |-- †Palaeotherates pennsylvanicus Handlirsch, 1906 | |--o †Permotupus Nel, Fleck, Garrouste, Gand, Lapeyrie, Bybee & Prokop, 2009 | | |-- †P. minor | | `-- †P. ollieorum | |-- †Petrotypus multivenosus Zalessky, 1950 | |-- †Stephanotypus schneideri Zessin, 1983 | `--o †Tupus Sellards, 1906 [Typus Sellards, 1909; Meganeurela Handlirsch 1906] | |-- †T. gallicus | |-- †T. gilmorei | |-- †T. gracilis Carpenter, 1947 | |-- †T. orientalis | |-- †T. permianus | `-- †T. whitei `--o †Meganeurinae Handlirsch, 1906sensu Bechly, 1996 |-- †Meganeurula selysii (Brongniart, 1894) Handlirsch, 1906 |-- †Meganeuropsis permiana Carpenter 1939 [M. americana Carpenter 1947] `-- †Meganeura monyi Brongniart, 1885 [Meganura monyi Brongniart, 1885]
Reference(s):
- Bechly, G., 1998-2002: Phylogenetic classification of fossil and extant odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm - Bechly, G., 1998-2002b: Phylogenetic Systematics of basal Pterygota and Stem-Group Odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/odonato1.htm - Beckemeyer, R. J. 2006-: Checklist of Midco, Oklahoma Permian insects. [visited: 2006-08-04]
–iNet: www.windofkansas.com: http://www.windsofkansas.com/midco_taxa.html - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968 - Nel, A., Fleck, G., Garrouste, R., Gand, G., Lapeyrie, J., Bybee, S. M. & Prokop, J., 2009: Revision of Permo-Carboniferous griffenflies (Insecta: Odonatoptera: Meganisoptera) based upon new species and redescription of selected poorly known taxa from Eurasia.
–Palaeontographica Abteilung A: Vol. 289, #4-6, pp. 89-121