After Bechly, 1998-2002, with Recent species from Schorr, Lindeboom & Paulson, 2003
<==o Neolamellida Bechly, 1996 [Firmonervata Lohmann, 1996] |--o Gomphomacromiidae Tillyard & Fraser, 1940 sensu Bechly, 1996 [paraphyletic Corduliidae] | |--o Archaeophya Fraser, 1959 | | |-- A. adamsi Fraser, 1959 [Aeschnosoma peruviana Cowley, 1934] | | `-- A. magnifica Theischinger & Watson, 1978 | `--o Gomphomacromia Brauer, 1864 | |-- G. chilensis Martin, 1921 [Epophthalmia cyanocephala Hagen, 1867] | |-- G. fallax McLachlan, 1881 | |-- G. mexicana Needham, 1933 | |-- G. nodisticta Ris, 1928 | `-- G. paradoxa Brauer, 1864 [Cordulia chilensis Hagen, 1861 (nomen nudum); Gomphomacromia effusa Navás, 1918; Gomphomacromia etcheverryi Fraser, 1957; Gomphomacromia tincta Navás, 1918] `--o Valvulida Lohmann, 1996 |--o Pseudocordulia Tillyard, 1909 [Pseudocorduliidae Lohmann, 1996] [paraphyletic Corduliidae] | |-- P. circularis Tillyard, 1909 [Cordulia novae-zeelandiae Brauer, 1865] | `-- P. elliptica Tillyard, 1913 `--o Trichodopalpida Bechly, 1996 |--o Macromiidae Needham, 1903 sensu Davies, 1981 [paraphyletic Corduliidae] | |== other genera [Bechly, 1998-2002c] | `-- Macromia Rambur, 1842 `--o Laterocarinida Bechly, 1996 |-- Austrocorduliidae Bechly, 1996 [paraphyletic Corduliidae] `--o Mediocostida Bechly, 1996 |-- Idionychidae Tillyard & Fraser, 1940 [paraphyletic Corduliidae] `--o Longiansida Bechly, 1996 |--o Idomacromiidae Tillyard & Fraser, 1940 [paraphyletic Corduliidae] | |--o Nesocordulia McLachlan, 1882 [Nesocorduliinae Bechly, 1996] | | |-- N. flavicauda McLachlan, 1882 [Cordulia valga Hagen, 1861 (nomen nudum)] | | |-- N. malgassica Fraser, 1956 | | |-- N. mascarenica Fraser, 1948 | | |-- N. rubricauda Martin, 1900 | | |-- N. spinicauda Martin, 1903 | | `-- N. villiersi Legrand, 1984 | `--o Idomacromia Karsch, 1896 [Idomacromiinae Tillyard & Fraser, 1940] | |-- I. lieftincki Legrand, 1984 [Idionyx dohrni Krüger, 1899] | `-- I. proavita Karsch, 1896 [Idomacromia couturieri Legrand, 1985] `--o Liberaponsida Bechly, 1996 |--o Cordulephyidae Tillyard, 1917 [paraphyletic Corduliidae] | |-- Neophya rutherfordi Selys, 1881 [Cordulia valga Hagen, 1861 (nomen nudum)] [Neophyinae Tillyard & Fraser, 1940] | `--o Cordulephya Selys, 1870 [Cordulephyinae Tillyard, 1917 sensu Tillyard & Fraser, 1940] | |-- C. bidens Sjöstedt, 1917 [Austrocordulia jurzitzai Theischinger, 1999] | |-- C. divergens Tillyard, 1917 | |-- C. montana Tillyard, 1911 | `-- C. pygmaea Selys, 1870 `--o Haplomulida Bechly, 1996 |--o Oxygastridae Bechly, 1996 [paraphyletic Corduliidae] | |-- Hesperocordulia berthoudi Tillyard, 1911 [Hemicordulia delicata Martin, 1896] | |--o Neocordulia Selys, 1882 | | |-- N. androgynis (Selys, 1871) [Gomphomacromia androgynis Selys, 1871; Anacordulia maccullochi Tillyard, 1926] | | |-- N. batesi (Selys, 1871) [Gomphomacromia batesi Selys, 1871; Neocordulia longipollex Calvert, 1909] | | |-- N. biancoi Rácenis, 1970 | | |-- N. campana May & Knopf, 1988 | | |-- N. carlochagasi Santos, 1967 | | |-- N. griphus May, 1992 | | |-- N. mambucabensis Costa & Santos, 2000 | | |-- N. setifera (Hagen, in Selys, 1871) [Gomphomacromia setifera Hagen, in Selys, 1871; Cordulia valga Hagen, 1861 (nomen nudum)] | | `-- N. volxemi (Selys, 1874) [Gomphomacromia volxemi Selys, 1874] | `-- Oxygastra curtisii (Dale, 1834) Selys, 1870 [Cordulia curtisii Dale, 1834; Epitheca? jamascarensis Hagen, in Selys, 1878; Libellula nitens Fonscolombe, 1837] `--o Italoansida Bechly, 1996 |-- Corduliidae Selys, in Hagen & Selys, 1850 sensu Bechly, 1996 (kiiltokorennot) `--o Anauriculida Bechly, 1996 |--o Hemicorduliidae Bechly, 1996 [paraphyletic Corduliidae] | |== other genera? [Bechly, 1998-2002c] | `-- Hemicordulia Selys, 1870 `--o Libellulida Leach, 1815 sensu Bechly, 1996 [Libellulidae Leach, 1815 sensu Schorr, Lindeboom & Paulson, 2003] |--o Urothemistidae Lieftinck, 1954 sensu Bechly, 1996 [Macrodiplacidae Fraser, 1957] | |== other genera [Bechly, 1998-2002c] | `--o Urothemis Brauer, 1868 | |-- U. abbotti Laidlaw, 1927 [Pronomaja mimetica Förster, 1909] | |-- U. aliena Selys, 1878 | |-- U. assignata (Selys, 1872) [Libellula assignata Selys, 1872; Libellula designata Selys, 1872; Urothemis madagascariensis Selys, 1897] | |-- U. bisignata Brauer, 1868 | |-- U. consignata Selys, 1897 | |-- U. edwardsii (Selys, 1849) [Libellula edwardsii Selys, 1849; Urothemis iridescens Kirby, 1898; Urothemis rendalli Kirby, 1898] | |-- U. luciana Balinsky, 1961 | |-- U. signata (Rambur, 1842) [Libellula signata Rambur, 1842; Libellula insignata Selys, 1872; Libellula sanguinea Burmeister, 1839 (nec MacLeay, 1826); Urothemis yiei Asahina, 1972] | `-- U. thomasi Longfield, 1932 [Urothemis aethiopica Nielsen, 1957] `--o Eulibellulida Bechly, 1996 `-- Libellulidae Leach, 1815 (varsinaiset sudenkorennot)
Genera not mentioned by Bechly (1998-2002):
“Corduliidae”
Antipodochlora Fraser, 1939
Antipodochlora braueri (Selys, 1871) [Epitheca braueri Selys, 1871; Aeschnosoma peruviana Cowley, 1934]
Apocordulia Watson, 1980
Apocordulia macrops Watson, 1980 [Aeschnosoma peruviana Cowley, 1934]
Austrophya Tillyard, 1909
Austrophya mystica Tillyard, 1909 [Austrocordulia jurzitzai Theischinger, 1999]
Didymops Rambur, 1842
Didymops floridensis Davis, 1921 [Cordulia bifurcata Hagen, 1861 (nomen nudum)]
Didymops transversa (Say, 1839) [Libellula transversa Say, 1839; Didymops servillii Rambur, 1842; Epophthalmia cinnamomea Burmeister, 1839]
Dorocordulia Needham, 1901
Dorocordulia lepida (Hagen, in Selys, 1871) [Cordulia lepida Hagen, in Selys, 1871; Epophthalmia cinnamomea Burmeister, 1839]
Dorocordulia libera (Selys, 1871) [Cordulia libera Selys, 1871]
Epitheca Burmeister, 1839
Epitheca bimaculata (Charpentier, 1825) [Libellula bimaculata Charpentier, 1825; Epophthalmia cinnamomea Burmeister, 1839]
Epitheca canis (McLachlan, 1886) [Tetragoneuria canis McLachlan, 1886]
Epitheca costalis (Selys, 1871) [Cordulia costalis Selys, 1871; Tetragoneuria williamsoni Muttkowski, 1911]
Epitheca cynosura (Say, 1839) [Libellula cynosura Say, 1839; Cordulia basiguttata Selys, 1871; Cordulia complanata Rambur, 1842; Cordulia diffinis Selys, 1871; Epophthalmia lateralis Burmeister, 1839; Tetragoneuria morio Muttkowski, 1911; Tetragoneuria simulans Muttkowski, 1911; ?Libellula tomentosa Fabricius, 1775 (nomen oblitum?)]
Epitheca marginata (Selys, 1883) [Somatochlora marginata Selys, 1883]
Epitheca petechialis (Muttkowski, 1911) [Tetragoneuria petechialis Muttkowski, 1911]
Epitheca princeps Hagen, 1861 [Cordulia regina Hagen, in Selys, 1871]
Epitheca semiaquea (Burmeister, 1839) [Libellula semiaquea Burmeister, 1839; Tetragoneura calverti Muttkowski, 1915; Tetragoneuria diffinis Hagen, 1861 (nomen nudum)]
Epitheca sepia (Gloyd, 1933) [Tetragoneuria sepia Gloyd, 1933]
Epitheca spinigera (Selys, 1871) [Cordulia spinigera Selys, 1871; Tetragoneuria indistincta Morse, 1895]
Epitheca spinosa (Hagen, in Selys, 1878) [Cordulia spinosa Hagen, in Selys, 1878]
Epitheca stella (Williamson, in Muttkowski, 1911) [Tetragoneuria stella Williamson, in Muttkowski, 1911]
Epophthalmia Burmeister, 1839
Epophthalmia australis Hagen, 1867 [Tetragoneuria indistincta Morse, 1895]
Epophthalmia elegans (Brauer, 1865) [Macromia elegans Brauer, 1865]
Epophthalmia frontalis Selys, 1871
Epophthalmia kuani Jiang, 1998
Epophthalmia vittata Burmeister, 1839 [Epophthalmia cyanocephala Hagen, 1867]
Epophthalmia vittigera (Rambur, 1842) [Macromia vittigera Rambur, 1842]
Guadalca Kimmins, 1957
Guadalca insularis Kimmins, 1957 [Gomphomacromia tincta Navás, 1918]
Helocordulia Needham, 1901
Helocordulia selysii (Hagen, in Selys, 1878) [Cordulia? selysii Hagen, in Selys, 1878; Gomphomacromia tincta Navás, 1918]
Helocordulia uhleri (Selys, 1871) [Cordulia uhleri Selys, 1871]
Lathrocordulia Tillyard, 1911
Lathrocordulia garrisoni Theischinger & Watson, 1991 [Idomacromia couturieri Legrand, 1985]
Lathrocordulia metallica Tillyard, 1911
Macromidia Martin, 1907
Macromidia asahinai Lieftinck, 1971 [Macromia flavipennis Walsh, 1862]
Macromidia atrovirens Lieftinck, 1935
Macromidia donaldi (Fraser, 1924) [Indomacromia donaldi Fraser, 1924]
Macromidia ellenae Wilson, 1996
Macromidia fulva Laidlaw, 1915
Macromidia genialis Laidlaw, 1923 [Macromidia erratica Lieftinck, 1948; Macromidia shanensis Fraser, 1927]
Macromidia hanzhouensis Zhou & Wei, 1979
Macromidia ishidai Asahina, 1964
Macromidia kelloggi Asahina, 1978
Macromidia rapida Martin, 1907 [Macromia cantonensis Tinkham, 1936]
Macromidia samal Needham & Gyger, 1937
Metaphya Laidlaw, 1912
Metaphya elongata Campion, 1921 [Macromia cantonensis Tinkham, 1936]
Metaphya micans Laidlaw, 1912
Metaphya stueberi (Lieftinck, 1938) [Anacordulia stueberi Lieftinck, 1938]
Metaphya tillyardi Ris, 1913 [Anacordulia maccullochi Tillyard, 1926]
Micromidia Fraser, 1959
Micromidia atrifrons (McLachlan, 1883) [Syncordulia atrifrons McLachlan, 1883; Anacordulia maccullochi Tillyard, 1926]
Micromidia convergens Theischinger & Watson, 1978
Micromidia rodericki Fraser, 1959
Navicordulia Machado & Costa, 1995
Navicordulia amazonica Machado & Costa, 1995 [Anacordulia maccullochi Tillyard, 1926]
Navicordulia atlantica Machado & Costa, 1995
Navicordulia errans (Calvert, 1909) [Dorocordulia errans Calvert, 1909]
Navicordulia kiautai Machado & Costa, 1995
Navicordulia leptostyla Machado & Costa, 1995
Navicordulia longistyla Machado & Costa, 1995
Navicordulia mielkei Machado & Costa, 1995
Navicordulia miersi Machado & Costa, 1995
Navicordulia nitens (De Marmels, 1991) [Dorocordulia nitens De Marmels, 1991]
Navicordulia vagans (De Marmels, 1989) [Dorocordulia vagans De Marmels, 1989]
Paracordulia Martin, 1906
Paracordulia sericea (Selys, 1871) [Cordulia sericea Selys, 1871; Libellula nitens Fonscolombe, 1837]
Pentathemis Karsch, 1890
Pentathemis membranulata Karsch, 1890 [Libellula nitens Fonscolombe, 1837]
Phyllomacromia Selys, 1878
Phyllomacromia aeneothorax (Nunney, 1895) [Ceratogomphus aeneothorax Nunney, 1895; Libellula nitens Fonscolombe, 1837; Macromia camerunica Pinhey, 1974; Macromia lieftincki Fraser, 1954; Macromia selysi Kirby, 1900]
Phyllomacromia aequatorialis Martin, 1906
Phyllomacromia amicorum (Gambles, 1979) [Macromia amicorum Gambles, 1979]
Phyllomacromia aureozona (Pinhey, 1966) [Macromia aureozona Pinhey, 1966]
Phyllomacromia bicristulata (Legrand, 1975) [Macromia bicristulata Legrand, 1975]
Phyllomacromia bifasciata (Martin, 1912) [Macromia bifasciata Martin, 1912; Phyllomacromia leoni Fraser, 1928]
Phyllomacromia bispina (Fraser, 1954) [Macromia bispina Fraser, 1954]
Phyllomacromia caneri (Gauthier, 1987) [Macromia caneri Gauthier, 1987]
Phyllomacromia congolica (Fraser, 1955) [Macromia congolica Fraser, 1955]
Phyllomacromia flavimitella (Pinhey, 1966) [Macromia flavimitella Pinhey, 1966]
Phyllomacromia funicularia (Martin, 1906) [Macromia funicularia Martin, 1906; Macromia bredoi Schouteden, 1934; Macromia martorelli Compte Sart, 1964]
Phyllomacromia funicularioides (Legrand, 1983) [Macromia funicularioides Legrand, 1983]
Phyllomacromia gamblesi (Lindley, 1980) [Macromia gamblesi Lindley, 1980]
Phyllomacromia girardi (Legrand, 1991) [Macromia girardi Legrand, 1991]
Phyllomacromia hervei (Legrand, 1980) [Macromia hervei Legrand, 1980]
Phyllomacromia kimminsi (Fraser, 1954) [Macromia kimminsi Fraser, 1954; Macromia junior Pinhey, 1961]
Phyllomacromia lamottei (Legrand, 1993) [Macromia lamottei Legrand, 1993]
Phyllomacromia legrandi (Gauthier, 1987) [Macromia legrandi Gauthier, 1987]
Phyllomacromia maesi (Schouteden, 1917) [Macromia maesi Schouteden, 1917]
Phyllomacromia melania (Selys, 1871) [Macromia melania Selys, 1871; Hylaeschna paludis Sjöstedt, 1899; Phyllomacromia biflava Martin, 1907; Phyllomacromia contumax Selys, 1879]
Phyllomacromia monoceros (Förster, 1906) [Macromia monoceros Förster, 1906; Macromia thetis Ris, 1921]
Phyllomacromia nigeriensis (Gambles, 1971) [Macromia nigeriensis Gambles, 1971]
Phyllomacromia nyanzana (Grünberg, 1911) [Macromia nyanzana Grünberg, 1911; Macromia halei Fraser, 1928; Macromia reginae Le Roi, 1915]
Phyllomacromia occidentalis (Fraser, 1954) [Macromia occidentalis Fraser, 1954]
Phyllomacromia onerata (Martin, 1907) [Macromia onerata Martin, 1907; Macromia clymene Ris, 1921]
Phyllomacromia overlaeti (Schouteden, 1934) [Macromia overlaeti Schouteden, 1934; Macromia schoutedeni Fraser, 1954]
Phyllomacromia pallidinervis (Förster, 1906) [Macromia pallidinervis Förster, 1906]
Phyllomacromia paludosa (Pinhey, 1976) [Macromia paludosa Pinhey, 1976]
Phyllomacromia paula (Karsch, 1892) [Macromia paula Karsch, 1892; Macromia bicornis Förster, 1906]
Phyllomacromia picta (Hagen, in Selys, 1871) [Macromia picta Hagen, in Selys, 1871; Macromia africana Hagen, in Selys, 1871; Macromia flavicans Fraser, 1954; Macromia kochi Grünberg, 1911; Macromia trochi Grünberg, 1911; Macromia tropicalis Selys, 1878; Phyllomacromia flavicincta Kirby, 1898]
Phyllomacromia pseudafricana (Pinhey, 1961) [Macromia pseudafricana Pinhey, 1961]
Phyllomacromia royi (Legrand, 1982) [Macromia royi Legrand, 1982]
Phyllomacromia seydeli (Fraser, 1954) [Macromia seydeli Fraser, 1954]
Phyllomacromia sophia (Selys, 1871) [Macromia sophia Selys, 1871; Pseudogomphus insignis Kirby, 1889]
Phyllomacromia subtropicalis (Fraser, 1954) [Macromia subtropicalis Fraser, 1954]
Phyllomacromia sylvatica (Fraser, 1954) [Macromia sylvatica Fraser, 1954]
Phyllomacromia trifasciata (Rambur, 1842) [Macromia trifasciata Rambur, 1842]
Phyllomacromia unifasciata (Fraser, 1954) [Macromia unifasciata Fraser, 1954]
Phyllomacromia villiersi (Legrand, 1992) [Macromia villiersi Legrand, 1992]
Procordulia Martin, 1907
Procordulia affinis (Selys, 1871) [Cordulia affinis Selys, 1871; Pseudogomphus insignis Kirby, 1889]
Procordulia artemis Lieftinck, 1930
Procordulia astridae Lieftinck, 1935
Procordulia fusiformis Lieftinck, 1977
Procordulia grayi (Selys, 1871) [Epitheca grayi Selys, 1871]
Procordulia irregularis Martin, 1907
Procordulia jacksoniensis (Rambur, 1842) [Cordulia jacksoniensis Rambur, 1842]
Procordulia karnyi Fraser, 1926
Procordulia leopoldi Fraser, 1932
Procordulia lompobatang van Tol, 1997
Procordulia moroensis Lieftinck, 1977
Procordulia papandayanensis van Tol, 1997
Procordulia rantemario van Tol, 1997
Procordulia sambawana (Förster, 1899) [Somatochlora sambawana Förster, 1899]
Procordulia smithii (White, 1846) [Cordulia smithii White, 1846; Cordulia novae-zeelandiae Brauer, 1865]
Procordulia sylvia Lieftinck, 1935
Rialla Navás, 1915
Rialla villosa (Rambur, 1842) [Cordulia villosa Rambur, 1842; Cordulia novae-zeelandiae Brauer, 1865; Rialla membranata Navás, 1915]
Santosia Costa & Santos, 1992
Santosia machadoi Costa & Santos, 2000 [Rialla membranata Navás, 1915]
Santosia marshalli Costa & Santos, 1992
Santosia newtoni Costa & Santos, 2000
Somatochlora Selys, 1871
Somatochlora albicincta (Burmeister, 1839) [Epophthalmia albicincta Burmeister, 1839; Rialla membranata Navás, 1915; Cordulia eremita Scudder, 1866; Somatochlora albicincta form massettensis Whitehouse, 1941]
Somatochlora alpestris (Selys, 1840) [Cordulia alpestris Selys, 1840]
Somatochlora arctica (Zetterstedt, 1840) [Aeshna arctica Zetterstedt, 1840; Cordulia subalpina Selys, 1841]
Somatochlora borisi Marinov, 2001
Somatochlora brevicincta Robert, 1954
Somatochlora calverti Williamson & Gloyd, 1933
Somatochlora cingulata (Selys, 1871) [Epitheca cingulata Selys, 1871]
Somatochlora clavata Oguma, 1922
Somatochlora daviesi Lieftinck, 1977
Somatochlora dido Needham, 1930
Somatochlora elongata (Scudder, 1866) [Cordulia elongata Scudder, 1866; Cordulia saturata Hagen, 1861 (nomen nudum)]
Somatochlora ensigera Martin, 1907 [Somatochlora charadraea Williamson, 1907]
Somatochlora filosa (Hagen, 1861) [Cordulia filosa Hagen, 1861]
Somatochlora flavomaculata (Vander Linden, 1825) [Libellula flavomaculata Vander Linden, 1825]
Somatochlora forcipata (Scudder, 1866) [Cordulia forcipata Scudder, 1866; Cordulia chalybea Hagen, 1861 (nomen nudum)]
Somatochlora franklini (Selys, 1878) [Epitheca franklini Selys, 1878; Somatochlora macrotona Williamson 1909]
Somatochlora georgiana Walker, 1925
Somatochlora graeseri Selys, 1887 [Somatochlora borealis Bartenev]
Somatochlora hineana Williamson, 1931
Somatochlora hudsonica (Hagen, in Selys, 1871) [Epitheca hudsonica Hagen, in Selys, 1871]
Somatochlora incurvata Walker, 1918 [Somatochlora borealis Bartenev]
Somatochlora japonica Matsumura, 1911 [Somatochlora borealis Bartenev]
Somatochlora kennedyi Walker, 1918
Somatochlora linearis (Hagen, 1861) [Cordulia linearis Hagen, 1861; Epitheca procera Selys, 1871; Somatochlora lateralis Needham, 1901 (lapsus)]
Somatochlora lingyinensis Zhou & Wa, 1979
Somatochlora margarita Donnelly, 1962
Somatochlora meridionalis Nielsen, 1935
Somatochlora metallica (Vander Linden, 1825) [Libellula metallica Vander Linden, 1825; Somatochlora coreana Doi, 1938; Somatochlora exuberata Bartenev, 1912]
Somatochlora minor Calvert, 1898
Somatochlora nepalensis Asahina, 1982
Somatochlora ozarkensis Bird, 1933
Somatochlora provocans Calvert, 1903
Somatochlora sahlbergi Trybom, 1889 [Somatochlora theeli Trybom, 1889; Somatochlora walkeri Kennedy, 1917]
Somatochlora semicircularis (Selys, 1871) [Epitheca semicircularis Selys, 1871; Epitheca nasalis Selys, 1874]
Somatochlora septentrionalis (Hagen, 1861) [Cordulia septentrionalis Hagen, 1861; Cordulia richardsoni Hagen, 1861 (nomen nudum)]
Somatochlora shanxiensis Zhu & Zhang, 1999
Somatochlora taiwana Inoue & Yokota, 2001
Somatochlora tenebrosa (Say, 1839) [Libellula tenebrosa Say, 1839; Cordulia tenebrica Hagen, 1861 (nomen nudum)]
Somatochlora uchidai Förster, 1909
Somatochlora viridiaenea (Uhler, 1858) [Cordulia viridiaenea Uhler, 1858]
Somatochlora walshii (Scudder, 1866) [Cordulia walshii Scudder, 1866]
Somatochlora whitehousei Walker, 1925
Somatochlora williamsoni Walker, 1907
Williamsonia Davis, 1913
Williamsonia fletcheri Williamson, 1923 [Cordulia tenebrica Hagen, 1861 (nomen nudum); Diplax vacua Hagen, 1867 (nomen nudum)]
Williamsonia lintneri (Hagen, in Selys, 1878) [Cordulia lintneri Hagen in Selys, 1878]
Reference(s):
- Bechly, G., 1998-2002a: Phylogenetic classification of fossil and extant odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm - Bechly, G., 1998-2002b: Phylogenetic Systematics of Euanisoptera / Cavilabiata.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/anisopt3.htm - Bechly, G., 1998-2002c: Phylogenetic Systematics of Euanisoptera / Neolamellida.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/anisopt4.htm - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968. - Schorr, M., Lindeboom, M. & Paulson, D., 2003: World Species List – Odonata.
–iNet: International Dragonfly Fund: http://www.calopteryx.de/idf/