Mikko's Phylogeny Archive

Gomphidae

Main Groups of Insects

After Bechly, 1998-2002, with Recent species from Schorr, Lindeboom & Paulson, 2003

<==o Gomphidae Rambur, 1842 sensu Bechly, 1998 (jokikorennot)
   |--o Austrogomphinae Carle, 1986
   |  |== other genera
   |  `--o Austrogomphus Selys, 1854 [20]
   |     |-- A. amphiclitus (Selys, 1873) [Hemigomphus amphiclitus Selys, 1873] [incl. Gomphus excavatus Bartenev, 1930; Austrogomphus risi Martin, 1901]
   |     |-- A. angeli Tillyard, 1913
   |     |-- A. arbustorum Tillyard, 1906
   |     |-- A. australis Dale, in Selys, 1854
   |     |-- A. bifurcatus Tillyard, 1909
   |     |-- A. collaris Hagen, in Selys, 1854
   |     |-- A. cornutus Watson, 1991
   |     |-- A. divaricatus Watson, 1991
   |     |-- A. doddi Tillyard, 1909
   |     |-- A. gordoni Watson, 1962
   |     |-- A. guerini (Rambur, 1842) [Gomphus guerini Rambur, 1842]
   |     |-- A. lateralis (Selys, 1873) [Hemigomphus lateralis Selys, 1873] [incl. Austrogomphus occidentalis Tillyard, 1908]
   |     |-- A. melaleucae Tillyard, 1909
   |     |-- A. mjobergi Sjöstedt, 1917
   |     |-- A. mouldsorum Theischinger, 1999
   |     |-- A. ochraceus (Selys, 1869) [Hemigomphus ochraceus Selys, 1869]
   |     |-- A. praeruptus (Selys, 1858) [Onychogomphus praeruptus Selys, 1858]
   |     |-- A. prasinus Tillyard, 1906
   |     |-- A. pusillus Sjöstedt, 1917
   |     `-- A. turneri Martin, 1901
   |--o Phyllogomphinae Carle, 1986
   |  |== other genera
   |  `--o Phyllogomphus Selys, 1854
   |     |-- P. aethiops Selys, 1854 [Gomphoides perfida Hagen, 1861; Phyllogomphus helenae Lacroix, 1921]
   |     |-- P. bartolozzii Marconi, Terzani & Carletti, 2001
   |     |-- P. brunneus Pinhey, 1976
   |     |-- P. coloratus Kimmins, 1931
   |     |-- P. corbetae Vick, 1999
   |     |-- P. dundomajoricus Fraser, 1957
   |     |-- P. dundominusculus Fraser, 1957
   |     |-- P. latifasciae Pinhey, 1961
   |     |-- P. margaritae Compte Sart, 1967
   |     |-- P. montanus Fraser, 1957
   |     |-- P. hartwigi Buchholz, 1958
   |     |-- P. moundi Fraser, 1960
   |     |-- P. occidentalis Fraser, 1957
   |     |-- P. orientalis Fraser, 1957 [Phyllogomphus edentatus St. Quentin, 1958]
   |     |-- P. perisi (Compte Sart, 1963) [Guineagomphus perisi Compte Sart, 1963]
   |     |-- P. pseudoccidentalis Lindley, 1972
   |     |-- P. schoutedeni Fraser, 1957
   |     |-- P. selysi Schouteden, 1933 [Phyllogomphus annulus Klots, 1944]
   |     `-- P. symoensi Lieftinck, 1969
   |--o Onychogomphinae Chao, 1984
   |  |--o Crenigomphini Carle, 1986
   |  |  |== other genera
   |  |  `--o Crenigomphus Selys, 1892
   |  |     |-- C. abyssinicus (Selys, 1878) [Onychogomphus abyssinicus Selys, 1878] [incl. Gomphidia guyi Pinhey, 1967, in partim]
   |  |     |-- C. cornutus Pinhey, 1956
   |  |     |-- C. denticulatus Selys, 1892 [incl. Crenigomphus occidentalis Martin, 1911]
   |  |     |-- C. hartmanni (Förster, 1898) [Onychogomphus hartmanni Förster, 1898] [incl. Bursigomphus pardus Martin, 1911]
   |  |     `-- C. renei Fraser, 1936 [incl. Dentigomphus rubrithorax Fraser, 1936]
   |  `--o Onychogomphini Chao, 1984
   |     |== other genera
   |     `-- Onychogomphus Selys, 1854
   |--o Gomphinae Rambur, 1842 sensu Davies & Tobin, 1985
   |  |--o Anisogomphini Carle, 1986
   |  |  |== other genera?
   |  |  `--o Anisogomphus Selys, 1857 [19]
   |  |     |-- A. anderi Lieftinck, 1948 [incl. Agriogomphus aquicola Fraser, 1943]
   |  |     |-- A. bivittatus Selys, 1854 [incl. Acrogomphus mohani Sahni, 1965]
   |  |     |-- A. caudalis Frasser, 1926
   |  |     |-- A. chaoi Liu, 1991
   |  |     |-- A. flavifacies Klots, 1947
   |  |     |-- A. forresti (Morton, 1928) [Temnogomphus forresti Morton, 1928]
   |  |     |-- A. fujianensis Zhou & Wu, 1992
   |  |     |-- A. jinggangshanus Liu, 1991
   |  |     |-- A. koxingai Chao, 1954
   |  |     |-- A. maacki (Selys, 1872) [Gomphus maacki Selys, 1872] [incl. Gomphus emarginatus Okumura, 1937; Gomphus m-flavum Selys, 1878; Temnogomphus amurensis Bartenev, 1930]
   |  |     |-- A. nitidus Yang & Davies, 1993
   |  |     |-- A. occipitalis (Selys, 1854) [Gomphus occipitalis Selys, 1854]
   |  |     |-- A. orites Laidlaw, 1922
   |  |     |-- A. pinratani Hämäläinen, 1991
   |  |     |-- A. resortus Yang & Davies, 1996
   |  |     |-- A. solitaris Lieftinck, 1971
   |  |     |-- A. vulvalis Yousuf & Yunes, 1977
   |  |     |-- A. wuzhishanus Chao, 1982
   |  |     `-- A. yunnanensis Zhou & Wu, 1992
   |  |--o Cyclogomphini Carle, 1986
   |  |  |== other genera
   |  |  `--o Cyclogomphus Selys, 1854
   |  |     |-- C. gynostylus Fraser, 1926 [incl. Ebegomphus strumens Needham, 1944]
   |  |     |-- C. heterostylus Selys, 1854
   |  |     |-- C. vesiculosus Selys, 1873
   |  |     |-- C. wilkinsi Fraser, 1926
   |  |     `-- C. ypsilon Selys, 1854
   |  |--o Gomphini Rambur, 1842 sensu Tillyard, 1917
   |  |  |== other genera
   |  |  `-- Gomphus Leach, 1815
   |  `--o Neurogomphini
   |     |== other genera
   |     `--o Neurogomphus Karsch, 1890
   |        |-- N. agilis (Martin, 1908) [Notogomphus agilis Martin, 1908; Nepogomphoides pinheyi Fraser, 1952]
   |        |-- N. angustisigna Pinhey, 1971
   |        |-- N. chapini (Klots, 1944) [Oxygomphus chapini Klots, 1944]
   |        |-- N. featheri Pinhey, 1967
   |        |-- N. fuscifrons Karsch, 1890
   |        |-- N. martininus (Lacroix, 1921) [Oxygomphus martininus Lacroix, 1921; Karschiogomphus ghesquierei Schouteden, 1934]
   |        |-- N. pallidus Cammaerts, 1967
   |        |-- N. pinheyi Cammaerts, 1968
   |        |-- N. uelensis Schouteden, 1934 [Neurogomphus vicinus Schouteden, 1934]
   |        `-- N. wittei Schouteden, 1934
   `--o Octogomphinae Carle & Cook, 1984
      |--o Hemigomphini Carle, 1986
      |  |== other genera
      |  `--o Hemigomphus Selys, 1854
      |     |-- H. atratus Watson, 1991 [Gomphus consobrinus Laidlaw, 1902]
      |     |-- H. comitatus (Tillyard, 1909) [Austrogomphus comitatus Tillyard, 1909]
      |     |-- H. cooloola Watson, 1991
      |     |-- H. gouldii (Selysi, 1854) [Austrogomphus gouldii Selys, 1854]
      |     |-- H. heteroclytus Selys, 1854
      |     |-- H. magela Watson, 1991
      |     `-- H. theischingeri Watson, 1991
      |--o Anomalophlebiini Belle, 1996
      |  |== other genera
      |  `-- Anomalophlebia nitida Belle, 1995 [incl. Temnogomphus amurensis Bartenev, 1930, in partim]
      |--o Octogomphini Carle & Cook, 1984
      |  |== other genera
      |  `-- Octogomphus specularis (Hagen inSelys, 1859 Selys, 1873 [Neogomphus? specularis Hagen inSelys, 1859; Notogomphus longus Martin, 1915]
      `--o Trigomphini Carle, 1986
         |== other genera
         `--o Trigomphus Bartenev, 1911
            |-- T. agricola (Ris, 1916) [Gomphus agricola Ris, 1916; Gomphus elongatus Selys, 1854]
            |-- T. beatus Chao, 1954
            |-- T. carus Chao, 1954
            |-- T. citimus (Needham, 1931) [Gomphus citimus Needham, 1931]
            |-- T. interruptus (Selys, 1854) [Hemigomphus interruptus Selys, 1854; Gomphus chichibui Fraser, 1936]
            |-- T. lautus (Needham, 1931) [Gomphus lautus Needham, 1931]
            |-- T. melampus (Selys, 1869) [Gomphus melampus Selys, 1869; Gomphus unifasciatus Oguma, 1926]
            |-- T. nigripes (Selys, 1887) [Gomphus nigripes Selys, 1887; Trigomphus anormolobatus Bartenev, 1912]
            |-- T. ogumai Asahina, 1949
            |-- T. succumbens (Needham, 1930) [Gomphus succumbens Needham, 1930]
            |-- T. svenhedini (Sjöstedt, 1933) [Gomphus svenhedini Sjöstedt, 1933]
            `-- T. yunnanensis Zhou & Wu, 1992

Reference(s):

• Bechly, G., 1998-2002a: Phylogenetic classification of fossil and extant odonates.
  –iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm
• Bechly, G., 1998-2002b: Phylogenetic Systematics of Euanisoptera / Exophytica – Gomphides.
  –iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/anisopt2.htm
• Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
  –Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
        Translated from: Beaut, Du Monde Animal XI & XII Invertébrés.
  –Rizzoli Editore, Milano, 1968
• Schorr, M., Lindeboom, M. & Paulson, D., 2003: World Species List – Odonata.
  –iNet: International Dragonfly Fund: http://www.calopteryx.de/idf/

 

© Mikko Haaramo, | http://www.mv.helsinki.fi/home/mhaaramo | Content updated 2007-10-29 | Page modified 2019-02-27