After Bechly, 1998-2002, with Recent species from Schorr, Lindeboom & Paulson, 2003
<==o Aeshnidae Leach, 1815 (ukonkorennot) |--o Oplonaeschninae Bechly, 1996 | |-- †Basiaeschna janata (Say, 1839) Selys, 1883 [Aeschna janata Say, 1839] [incl. Telephlebia racleayi Martin, 1909; Aeschna minor Rambur, 1842] | `--o Oplonaeschna Selys, 1883 | |-- O. armata (Hagen, 1861) [Aeschna armata Hagen, 1861] [incl. Aeshna nigripes Navás, 1932, in partim] | `-- O. magna González-Soriano & Novelo-Gutiérrez, 1998 |--o Gynacanthinae Cockerell, 1913 | |== other genera? | `-- Gynacantha Rambur, 1842 [incl. Triacanthagyna Selys, 1883; Acanthagyna Kirby, 1890] [88+7] `--o Aeshninae Leach, 1815 |--o Polycanthagynini Tillyard & Fraser, 1940 sensu Bechly, 1996 | |-- “Aeshna” petalura (Martin, 1908) | `--o Polycanthagyna Fraser, 1933 | |-- P. chaoi Yang & Li, 1994 [incl. Aeshna nigripes Navás, 1932, in partim] | |-- P. erythromelas (McLachlan, 1896) [Aeshna erythromelas McLachlan, 1896] | |-- P. melanictera (Selys, 1883) [Aeschna melanictera Selys, 1883] | `-- P. ornithocephala (McLachlan, 1896) [Aeshna ornithocephala McLachlan, 1896] |--o Aeshnini Leach, 1815 | |== numerous other genera? | `-- Aeshna Fabricius, 1775 [Aeschna Anonymous, 1801] [79] `--o Anacini Cockerell, 1913 emend. Bechly, 1998-2002 [Anactinini Cockerell, 1913; Anactini Davies, 1981; Anactini Bechly, 1996] |-- Amphiaeschnina Cockerell, 1913 sensu Bechly, 1998-2002 `--o Anacina Cockerell, 1913 emend. Bechly, 1998-2002 |-- Anax Leach, 1815 in Leach, 1814-1817 [incl. Hemianax Selys, 1883] [32] `--o Anaciaeschna Selys, 1878 |-- A. donaldi Fraser, 1922 [Anax striatus Kirby, 1899] |-- A. isosceles (Müller, 1767)[Libellula isoceles Müller, 1767 (emended by Lucas)] [incl. Aeshna chrysophthalmus Charpentier, 1825; Aeshna rufescens Vander Linden, 1825] |-- A. jaspidea (Burmeister, 1839) [Aeshna jaspidea Burmeister, 1839] [incl. Aeschna tahitensis Brauer, 1865; Protoaeschna pseudochiri Förster, 1908] |-- A. kashimirensis Singh and Baijal, 1954 |-- A. martini (Selys, 1897) [Aeschna martini Selys, 1897] |-- A. megalopis Martin, 1908 |-- A. melanostoma Lieftinck, 1949 |-- A. moluccana lieftinck, 1930 |-- A. montivagans Lieftinck, 1932 `-- A. triangulifera McLachlan, 1896
Aeshnidae incertae sedis:
[= taxa not listed by Bechly (1998-2002)]
Adversaeschna Tillyard, 1916
Adversaeschna brevistyla (Rambur, 1842) [Aeshna brevistyla Rambur, 1842]
Agyrtacantha Lieftinck, 1937
Agyrtacantha dirupta (Karsch, 1889) [Triacanthagyna dirupta Karsch, 1889; Aeschna propinqua Scudder, 1866]
Agyrtacantha microstigma (Selys, 1878) [Gynacantha microstigma Selys, 1878; Karschia angulata Förster, 1900]
Agyrtacantha othello Lieftinck, 1942
Agyrtacantha tumidula Lieftinck, 1937
Andaeschna De Marmels, 1994
Andaeschna andresi (Rácenis, 1958) [Aeshna andresi Rácenis, 1958] [incl. Anax validus Hagen, 1875, nomen nudum]
Andaeschna rufipes (Ris, 1918) [Aeshna rufipes Ris, 1918]
Andaeschna timotocuica De Marmels, 1994
Andaeschna unicolor (Martin, 1908) [Aeschna unicolor Martin, 1908]
Antipodophlebia Fraser, 1960
Antipodophlebia asthenes (Tillyard, 1916) [Telephlebia asthenes Tillyard, 1916] [incl. Anax validus Hagen, 1875, nomen nudum]
Austrogynacantha Tillyard, 1908
Austrogynacantha heterogena Tillyard, 1908 [incl. Planaeschna longissima Martin, 1901]
Austrophlebia Tillyard, 1916
Austrophlebia costalis (Tillyard, 1907) [Planaeschna costalis Tillyard, 1907] [incl. Planaeschna longissima Martin, 1901; Telephlebia racleayi Martin, 1909]
Austrophlebia subcostalis Theischinger, 1996
Cephalaeschna Selys, 1883
Cephalaeschna acutifrons (Martin, 1909) [Caliaeschna acutifrons Martin, 1909] [incl. Libellula hafniensis Müller, 1764; Indophlebia asiatica Fraser, 1935]
Cephalaeschna chaoi Asahina, 1982 [incl. Indophlebia asiatica Fraser, 1935]
Cephalaeschna klapperichi Schmidt, 1961
Cephalaeschna klotsi Asahina, 1982
Cephalaeschna masoni (Martin, 1909) [Caliaeschna masoni Martin, 1909]
Cephalaeschna needhami Asahina, 1981
Cephalaeschna obversa Needham, 1930
Cephalaeschna orbifrons Selys, 1883
Cephalaeschna patronum Needham, 1930
Cephalaeschna risi Asahina, 1981
Cephalaeschna triadica Lieftinck, 1977
Cephalaeschna viridifrons Fraser, 1922 [incl. Cephalaeschna biguttata Fraser, 1935]
Cephalaeschna dinghuensis Wilson, 1999
Dendroaeschna Tillyard, 1916
Dendroaeschna conspersa (Tillyard, 1907) [Caliaeschna conspersa Tillyard, 1907] [incl. Aeschna virens Rambur, 1842 (nec Charpentier, 1840)]
Heliaeschna Selys, 1882
Heliaeschna bartelsi Lieftinck, 1940 [incl. Cephalaeschna lugubris Martin, 1909]
Heliaeschna crassa Krüger, 1899
Heliaeschna cynthiae Fraser, 1939
Heliaeschna filostyla Martin, 1906 [incl. Heliaeschna gladiostyla Martin, 1909]
Heliaeschna fuliginosa Karsch, 1893
Heliaeschna idae (Brauer, 1865) [Gynacantha idae Brauer, 1865]
Heliaeschna lanceolata Le Roi, 1915
Heliaeschna libyana (Fraser, 1928) [Gynacantha libyana Fraser, 1928]
Heliaeschna longfieldae Gambles, 1967
Heliaeschna raymondi Compte Sart, 1967
Heliaeschna sembe Pinhey, 1962
Heliaeschna simplicia (Karsch, 1891) [Amphiaeschna simplicia Karsch, 1891] [incl. Heliaeschna vanderweeli Martin, 1907]
Heliaeschna trinervulata Fraser, 1955
Heliaeschna ugandica McLachlan, 1896
Heliaeschna ukerewensis Pinhey, 1961
Heliaeschna uninervula Martin, 1909
Hemianax Selys, 1883
Hemianax papuensis (Burmeister, 1839) [Aeschna papuensis Burmeister, 1839] [incl. Heliaeschna vanderweeli Martin, 1907; Anax congener Rambur, 1842]
Indaeschna Fraser, 1926
Indaeschna baluga Needham and Gyger, 1937 [incl. Anax congener Rambur, 1842, in partim]
Indaeschna grubaueri (Förster, 1904) [Amphiaeschna grubaueri Förster, 1904]
Limnetron Förster, 1907
Limnetron antarcticum Förster, 1907 [incl. Anax congener Rambur, 1842, in partim; Aeshna viridivittata Fraser, 1947]
Limnetron debile (Karsch, 1891) [Epiaeschna debilis Karsch, 1891]
Neuraeschna Hagen, 1867
Neuraeschna calverti Kimmins, 1951 [Aeshna viridivittata Fraser, 1947, in partim]
Neuraeschna capillata Machet, 1990
Neuraeschna claviforcipata Martin, 1909
Neuraeschna clavulata Machet, 1990
Neuraeschna cornuta Belle, 1989
Neuraeschna costalis (Burmeister, 1839) [Aeschna costalis Burmeister, 1839] [incl. Gynacantha ferox Erichson, 1848; Neuraeschna costalis var. hyalinata Belle, 1989; Neuraeschna costalis var. marginata Belle, 1989]
Neuraeschna dentigera Martin, 1909 [incl. Neuraeschna inarmata Kimmins, 1951]
Neuraeschna harpya Martin, 1909
Neuraeschna maxima Belle, 1989
Neuraeschna maya Belle, 1989
Neuraeschna mayoruna Belle, 1989
Neuraeschna mina Williamson and Williamson, 1930
Neuraeschna producta Kimmins, 1935
Neuraeschna titania Belle, 1989
Plattycantha Förster, 1908
Plattycantha acuta Lieftinck, 1937 [incl. Aeshna nigripes Navás, 1932, in partim]
Plattycantha cornuta (Förster, 1900) [Karschia cornuta Förster, 1900]
Plattycantha venatrix Lieftinck, 1937
Racenaeschna Calvert, 1958
Racenaeschna angustistrigis Calvert, 1958 [incl. Aeshna nigripes Navás, 1932, in partim]
Sarasaeschna Karube & Yeh, 2001
Sarasaeschna minuta Karube & Yeh, 2001
Staurophlebia Brauer, 1865
Staurophlebia auca Kennedy, 1937 [Remartinia barbiellina Navás, 1911]
Staurophlebia bosqui Navás, 1927
Staurophlebia gigantula Martin, 1909
Staurophlebia reticulata (Burmeister, 1839) [Aeschna reticulata Burmeister, 1839] [incl. Aeschna gigas Rambur, 1842; Staurophlebia magnifica Brauer, 1865; Staurophlebia reticulata guatemalteca Walker, 1915; Staurophlebia reticulata obscura Walker, 1915; ?Staurophlebia platyura Navás, 1920, in partim]
Staurophlebia wayana Geijskes, 1959
Subaeschna Martin, 1909
Subaeschna francesca Martin, 1909 [incl.? Staurophlebia platyura Navás, 1920, in partim]
Tetracanthagyna Selys, 1883
Tetracanthagyna bakeri Campion in Laidaw, 1928 [incl. Telephlebia mjobergi Sjöstedti, 1917, in partim]
Tetracanthagyna brunnea McLachlan, 1898
Tetracanthagyna degorsi Martin, 1895
Tetracanthagyna plagiata (Waterhouse, 1877) [Gynacantha plagiata Waterhouse, 1877] [incl. Tetracanthagyna vittata McLachlan, 1898]
Tetracanthagyna waterhousei MacLachlan, 1898 [incl. Tetracanthagyna sumatrana Förster, 1914; Toaeschna fontinalis Förster, 1905]
Reference(s):
- Bechly, G., 1998-2002a: Phylogenetic classification of fossil and extant odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm - Bechly, G., 1998-2002b: Phylogenetic Systematics of Euanisoptera / Aeshnoptera.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/anisopt1.htm - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980
Translated from: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968 - Schorr, M., Lindeboom, M. & Paulson, D., 2003: World Species List – Odonata.
–iNet: International Dragonfly Fund: http://www.calopteryx.de/idf/
