After Bechly, 1998-2002, with Recent species from Schorr, Lindeboom & Paulson, 2003
<==o Aeshnidae Leach, 1815 (ukonkorennot) |--o Oplonaeschninae Bechly, 1996 | |-- †Basiaeschna janata (Say, 1839) Selys, 1883 [Aeschna janata Say, 1839] [incl. Telephlebia racleayi Martin, 1909; Aeschna minor Rambur, 1842] | `--o Oplonaeschna Selys, 1883 | |-- O. armata (Hagen, 1861) [Aeschna armata Hagen, 1861] [incl. Aeshna nigripes Navás, 1932, in partim] | `-- O. magna González-Soriano & Novelo-Gutiérrez, 1998 |--o Gynacanthinae Cockerell, 1913 | |== other genera? | `-- Gynacantha Rambur, 1842 [incl. Triacanthagyna Selys, 1883; Acanthagyna Kirby, 1890] [88+7] `--o Aeshninae Leach, 1815 |--o Polycanthagynini Tillyard & Fraser, 1940 sensu Bechly, 1996 | |-- “Aeshna” petalura (Martin, 1908) | `--o Polycanthagyna Fraser, 1933 | |-- P. chaoi Yang & Li, 1994 [incl. Aeshna nigripes Navás, 1932, in partim] | |-- P. erythromelas (McLachlan, 1896) [Aeshna erythromelas McLachlan, 1896] | |-- P. melanictera (Selys, 1883) [Aeschna melanictera Selys, 1883] | `-- P. ornithocephala (McLachlan, 1896) [Aeshna ornithocephala McLachlan, 1896] |--o Aeshnini Leach, 1815 | |== numerous other genera? | `-- Aeshna Fabricius, 1775 [Aeschna Anonymous, 1801] [79] `--o Anacini Cockerell, 1913 emend. Bechly, 1998-2002 [Anactinini Cockerell, 1913; Anactini Davies, 1981; Anactini Bechly, 1996] |-- Amphiaeschnina Cockerell, 1913 sensu Bechly, 1998-2002 `--o Anacina Cockerell, 1913 emend. Bechly, 1998-2002 |-- Anax Leach, 1815 in Leach, 1814-1817 [incl. Hemianax Selys, 1883] [32] `--o Anaciaeschna Selys, 1878 |-- A. donaldi Fraser, 1922 [Anax striatus Kirby, 1899] |-- A. isosceles (Müller, 1767)[Libellula isoceles Müller, 1767 (emended by Lucas)] [incl. Aeshna chrysophthalmus Charpentier, 1825; Aeshna rufescens Vander Linden, 1825] |-- A. jaspidea (Burmeister, 1839) [Aeshna jaspidea Burmeister, 1839] [incl. Aeschna tahitensis Brauer, 1865; Protoaeschna pseudochiri Förster, 1908] |-- A. kashimirensis Singh and Baijal, 1954 |-- A. martini (Selys, 1897) [Aeschna martini Selys, 1897] |-- A. megalopis Martin, 1908 |-- A. melanostoma Lieftinck, 1949 |-- A. moluccana lieftinck, 1930 |-- A. montivagans Lieftinck, 1932 `-- A. triangulifera McLachlan, 1896
Aeshnidae incertae sedis:
[= taxa not listed by Bechly (1998-2002)]Adversaeschna Tillyard, 1916 Adversaeschna brevistyla (Rambur, 1842) [Aeshna brevistyla Rambur, 1842] Agyrtacantha Lieftinck, 1937 Agyrtacantha dirupta (Karsch, 1889) [Triacanthagyna dirupta Karsch, 1889; Aeschna propinqua Scudder, 1866] Agyrtacantha microstigma (Selys, 1878) [Gynacantha microstigma Selys, 1878; Karschia angulata Förster, 1900] Agyrtacantha othello Lieftinck, 1942 Agyrtacantha tumidula Lieftinck, 1937 Andaeschna De Marmels, 1994 Andaeschna andresi (Rácenis, 1958) [Aeshna andresi Rácenis, 1958] [incl. Anax validus Hagen, 1875, nomen nudum] Andaeschna rufipes (Ris, 1918) [Aeshna rufipes Ris, 1918] Andaeschna timotocuica De Marmels, 1994 Andaeschna unicolor (Martin, 1908) [Aeschna unicolor Martin, 1908] Antipodophlebia Fraser, 1960 Antipodophlebia asthenes (Tillyard, 1916) [Telephlebia asthenes Tillyard, 1916] [incl. Anax validus Hagen, 1875, nomen nudum] Austrogynacantha Tillyard, 1908 Austrogynacantha heterogena Tillyard, 1908 [incl. Planaeschna longissima Martin, 1901] Austrophlebia Tillyard, 1916 Austrophlebia costalis (Tillyard, 1907) [Planaeschna costalis Tillyard, 1907] [incl. Planaeschna longissima Martin, 1901; Telephlebia racleayi Martin, 1909] Austrophlebia subcostalis Theischinger, 1996 Cephalaeschna Selys, 1883 Cephalaeschna acutifrons (Martin, 1909) [Caliaeschna acutifrons Martin, 1909] [incl. Libellula hafniensis Müller, 1764; Indophlebia asiatica Fraser, 1935] Cephalaeschna chaoi Asahina, 1982 [incl. Indophlebia asiatica Fraser, 1935] Cephalaeschna klapperichi Schmidt, 1961 Cephalaeschna klotsi Asahina, 1982 Cephalaeschna masoni (Martin, 1909) [Caliaeschna masoni Martin, 1909] Cephalaeschna needhami Asahina, 1981 Cephalaeschna obversa Needham, 1930 Cephalaeschna orbifrons Selys, 1883 Cephalaeschna patronum Needham, 1930 Cephalaeschna risi Asahina, 1981 Cephalaeschna triadica Lieftinck, 1977 Cephalaeschna viridifrons Fraser, 1922 [incl. Cephalaeschna biguttata Fraser, 1935] Cephalaeschna dinghuensis Wilson, 1999 Dendroaeschna Tillyard, 1916 Dendroaeschna conspersa (Tillyard, 1907) [Caliaeschna conspersa Tillyard, 1907] [incl. Aeschna virens Rambur, 1842 (nec Charpentier, 1840)] Heliaeschna Selys, 1882 Heliaeschna bartelsi Lieftinck, 1940 [incl. Cephalaeschna lugubris Martin, 1909] Heliaeschna crassa Krüger, 1899 Heliaeschna cynthiae Fraser, 1939 Heliaeschna filostyla Martin, 1906 [incl. Heliaeschna gladiostyla Martin, 1909] Heliaeschna fuliginosa Karsch, 1893 Heliaeschna idae (Brauer, 1865) [Gynacantha idae Brauer, 1865] Heliaeschna lanceolata Le Roi, 1915 Heliaeschna libyana (Fraser, 1928) [Gynacantha libyana Fraser, 1928] Heliaeschna longfieldae Gambles, 1967 Heliaeschna raymondi Compte Sart, 1967 Heliaeschna sembe Pinhey, 1962 Heliaeschna simplicia (Karsch, 1891) [Amphiaeschna simplicia Karsch, 1891] [incl. Heliaeschna vanderweeli Martin, 1907] Heliaeschna trinervulata Fraser, 1955 Heliaeschna ugandica McLachlan, 1896 Heliaeschna ukerewensis Pinhey, 1961 Heliaeschna uninervula Martin, 1909 Hemianax Selys, 1883 Hemianax papuensis (Burmeister, 1839) [Aeschna papuensis Burmeister, 1839] [incl. Heliaeschna vanderweeli Martin, 1907; Anax congener Rambur, 1842] Indaeschna Fraser, 1926 Indaeschna baluga Needham and Gyger, 1937 [incl. Anax congener Rambur, 1842, in partim] Indaeschna grubaueri (Förster, 1904) [Amphiaeschna grubaueri Förster, 1904] Limnetron Förster, 1907 Limnetron antarcticum Förster, 1907 [incl. Anax congener Rambur, 1842, in partim; Aeshna viridivittata Fraser, 1947] Limnetron debile (Karsch, 1891) [Epiaeschna debilis Karsch, 1891] Neuraeschna Hagen, 1867 Neuraeschna calverti Kimmins, 1951 [Aeshna viridivittata Fraser, 1947, in partim] Neuraeschna capillata Machet, 1990 Neuraeschna claviforcipata Martin, 1909 Neuraeschna clavulata Machet, 1990 Neuraeschna cornuta Belle, 1989 Neuraeschna costalis (Burmeister, 1839) [Aeschna costalis Burmeister, 1839] [incl. Gynacantha ferox Erichson, 1848; Neuraeschna costalis var. hyalinata Belle, 1989; Neuraeschna costalis var. marginata Belle, 1989] Neuraeschna dentigera Martin, 1909 [incl. Neuraeschna inarmata Kimmins, 1951] Neuraeschna harpya Martin, 1909 Neuraeschna maxima Belle, 1989 Neuraeschna maya Belle, 1989 Neuraeschna mayoruna Belle, 1989 Neuraeschna mina Williamson and Williamson, 1930 Neuraeschna producta Kimmins, 1935 Neuraeschna titania Belle, 1989 Plattycantha Förster, 1908 Plattycantha acuta Lieftinck, 1937 [incl. Aeshna nigripes Navás, 1932, in partim] Plattycantha cornuta (Förster, 1900) [Karschia cornuta Förster, 1900] Plattycantha venatrix Lieftinck, 1937 Racenaeschna Calvert, 1958 Racenaeschna angustistrigis Calvert, 1958 [incl. Aeshna nigripes Navás, 1932, in partim] Sarasaeschna Karube & Yeh, 2001 Sarasaeschna minuta Karube & Yeh, 2001 Staurophlebia Brauer, 1865 Staurophlebia auca Kennedy, 1937 [Remartinia barbiellina Navás, 1911] Staurophlebia bosqui Navás, 1927 Staurophlebia gigantula Martin, 1909 Staurophlebia reticulata (Burmeister, 1839) [Aeschna reticulata Burmeister, 1839] [incl. Aeschna gigas Rambur, 1842; Staurophlebia magnifica Brauer, 1865; Staurophlebia reticulata guatemalteca Walker, 1915; Staurophlebia reticulata obscura Walker, 1915; ?Staurophlebia platyura Navás, 1920, in partim] Staurophlebia wayana Geijskes, 1959 Subaeschna Martin, 1909 Subaeschna francesca Martin, 1909 [incl.? Staurophlebia platyura Navás, 1920, in partim] Tetracanthagyna Selys, 1883 Tetracanthagyna bakeri Campion in Laidaw, 1928 [incl. Telephlebia mjobergi Sjöstedti, 1917, in partim] Tetracanthagyna brunnea McLachlan, 1898 Tetracanthagyna degorsi Martin, 1895 Tetracanthagyna plagiata (Waterhouse, 1877) [Gynacantha plagiata Waterhouse, 1877] [incl. Tetracanthagyna vittata McLachlan, 1898] Tetracanthagyna waterhousei MacLachlan, 1898 [incl. Tetracanthagyna sumatrana Förster, 1914; Toaeschna fontinalis Förster, 1905]
Reference(s):
- Bechly, G., 1998-2002a: Phylogenetic classification of fossil and extant odonates.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/system.htm - Bechly, G., 1998-2002b: Phylogenetic Systematics of Euanisoptera / Aeshnoptera.
–iNet: Günter Bechly's Odonatology Webpage: Phylogenetic Systematics of Odonata: http://www.bechly.de/anisopt1.htm - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980
Translated from: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968 - Schorr, M., Lindeboom, M. & Paulson, D., 2003: World Species List – Odonata.
–iNet: International Dragonfly Fund: http://www.calopteryx.de/idf/