After Palker, 1982, Grimaldi & Engel, 2005, and Engel & Grimaldi, 2007
<==o Neuroptera Linnaeus sensu stricto? [Planipennia] (verkkosiipiset (harsosiipiset); lacewings, antlions, owflies, etc.) |?- †Mesohemerobius jeholensis Ping, 1928; L.Cret. EAs. [Neuroptera incertae sedis (Makarkin et al., 2003)] |-- †“Permithonidae” `--+--o Nevrorthidae Nakahara [Neurorthidae (lapsus calami)] [Nevorthiformia] | |-- †Rophalis relicta (Hagen); M.Eoc. NEu. | |-- Austroneurorthus | |--o Nevorthus | | `-- N. fallax | `-- Nipponeurorthus `--+?- †Grammosmylidae |--o Hemerobiiformia (harsokorentomaiset) | |?- †“Osmylitidae” | `--+?-o †Prohemerobiidae | | `-- †Prohemerobius Handlirsch, 1906 | |?- †“Archaeosmylus” complesus Whalley, 1988 | |?- †“Archaeosmylus” alysius Whalley, 1988 | |--+--o Ithonoidea | | | |-- Polystoechotidae Handlirch 1906 (giant lacewings) | | | `-- Ithonidae Newman, 1853 [incl. Rapismatidae Krüger, 1922] (moth lacewings) | | `--+-- Chrysopidae Schneider (green lacewings; harsokorennot) | | `--o Osmyloidea | | |==o †Archeosmylidae Riek, 1953 [probably paraphyletic basal Osmyloids] | | | |--o †Archaeosmylus Riek, 1953; L.Perm.-E.Jur. Aust. | | | | |-- †A. pectinatus Reik, 1953 | | | | |?- †A. stigmatus Reik, 1955 | | | | |?- †A. costalis Reik, 1955 | | | | `?- †A. sp. (L.Trias. Aust) | | | |?- †Archeosmylidae gen. et sp. indet. (L.Perm. South Africa) | | | |-- †Babykamenia eskovi Ponomarenko & Shcherbakov, 2004 | | | `?-o †Lithosmylidia Reik, 1955; M.-L.Trias. Aust. | | | |-- †L. lineata Reik, 1955 | | | |?- †L. barrone Lambkin, 1988 | | | `?- †L. parvula Lambkin, 1988 | | `--o Osmylidae Leach, 1815 (osmylids) | | |-- †Archaeosmylidia fusca Makarkin, Yang & Ren, 2012; M.Jur. EAs. | | `--+?- †Tetanoptilon Bode, 1953 | | |-- †Osmalochrysa Jepson, Makarkin & Coram, 2012 | | |?-o †Mesosmylininae | | | |-- †Mesosmylina Bode, 1953 | | | |-- †Sogjuta Martynova, 1958 | | | `-- †Petrushevskia Martynova, 1958 | | |?-o †Saucrosmylinae Ren & Yin, 2003 | | | |-- †Saucrosmylus sambneurus Ren & Yin, 2003; M.Jur. EAs. | | | |-- †Rudiosmylus ningchengensis Ren & Yin, 2003; M.Jur. EAs. | | | |-- †Laccosmylus calophlebius Ren & Yin, 2003; M.Jur. EAs. | | | `--o †Bellinympha Y. Wang, Ren, Liu, Shih & Engel, 2010 in Y. Wang, Liu, X. Wang, Shih, Zhao, Engel & Ren, 2010; M.Jur. EAs. | | | |-- †B. filicifolia Y. Wang, Ren, Liu, Shih & Engel, 2010 in Y. Wang, Liu, X. Wang, Shih, Zhao, Engel & Ren, 2010 | | | `-- †B. dancei Y. Wang, Ren, Liu, Shih & Engel, 2010 in Y. Wang, Liu, X. Wang, Shih, Zhao, Engel & Ren, 2010 | | |--o Protosmylinae | | | |-- Gryposmylus | | | |-- Heterosmylus | | | |-- Paryphosmylus | | | |-- †Juraheterosmylus | | | |-- †Mesosmylus | | | |-- †Osmylidia | | | |-- †Protosmylina | | | `--o †Protosmylus | | | `-- †P. pictus (Hagen); M.Eoc. NEu. | | |--o Gumillinae | | |--o Spilosmylinae | | | `-- Thyridosmylus | | |-- Kempyninae | | |-- Porismus strigatus (Burmeister, 1839) [Porisminae] | | |-- Stenosmylinae | | |-- Eidoporisminae | | `--o Osmylinae | | `--o Osmylus | | `-- O. fulvicephalus | `--+-- Hemerobiidae Latreille [Hemerobioidea] (brown lacewings; kirvakorennot) | `--+--+--o Sisyridae (spongillaflies; rantakorennot) | | | |?- †Cratosisyrops L.Cret. ESA. (incertae sedis) | | | |?- †Sisyra? amissa Hagen; M.Eoc. NEu. | | | |== numerous Eocene fossil genera | | | |?- Climacia | | | |?- Sisyra | | | `== numerous Recent genera | | `-- Coniopterygidae Burmeister [Coniopterygoidea] (dustywings; vahakorennot) | `--o Mantispoidea | |--o Dilaridae Newman (pleasing lacewings) | | |--o Nallachiinae | | | `--o Nallachius | | | `-- N. americanus | | `--o Dilarinae | | |-- †Cascadilar eocenicus Engel, 1999; M.Eoc. NEu. | | `--o Dilar | | `-- D. septentrionalis | `--+-- Mantispidae Leach (mantispids; rukoilijakorennot) | `--+-- †“Mesithonidae” | `--+--o Rhachiberothidae Tjeder (thorny lacewings) | | |-- †Paraberotha acra Whalley; L.Cret. SWAs. | | |-- †Retinoberotha stuermeri Schlüter; U.Cret. WEu. | | |-- †Eorhachiberotha burmitica Engel, 2004; U.Cret. SEAs. | | |-- †Rhachibermissa splendida Grimaldi; U.Cret. ENAm. | | |-- †Whalfera venatrix (Whalley; 1983) M.Eoc. NEu. | | |-- †Alloberotha petrulevicii Nel et al., 2005; U.Cret. WEu. | | |-- †Chimerhachiberotha acrasarii Nel et al., 2005; L.Cret. SWAs. | | |-- †Spinoberotha mickaelacrai Nel et al., 2005; L.Cret. SWAs. | | |-- †Oisea celinea (Nel et al., 2005) Eoc. WEu. | | `== Recent genera | `--o Berothidae Handlirsch (beaded lacewings) | |?- †Banoberotha enigmatica Whalley; U.Cret. SWAs.. | |?-o †Jersiberotha Grimaldi; U.Cret. NAm. | | |-- †J. luzzii Grimaldi | | `-- †J. similis Grimaldi | |?- †Nascimberotha picta Grimaldi; U.Cret. NAm. | |?- †Microberotha macculloughi Archibald & Makarkin; Eoc. | |?- †Plesiorobius? anadensis Klimaszewski & Kevan; U.Cret. NAm. | |?- †Proberotha prisca Krüger, 1923; M.Eoc. NEu. | |?- Lomamyia | |-- Cyrenoberothinae | |-- Berothinae | |-- Nosybinae | |-- Nyrminae | |-- Protobiellinae | `-- Trichomatinae `--o Myrmeleontiformia (muurahaiskorentomaiset) |?- †Aetheogramma speciosa Ren & Engel, 2008 [Aetheogrammatidae Ren & Engel, 2008] `--o Myrmeleontoidea |--o Nemopteroidea | |--o †Kalligrammatidae Handlirsch, 1906 | | |--o †Kalligrammatinae Handlirsch, 1906 | | | |-- †Huiyingogramma formosum Liu et al., 2014 | | | |--o †Kalligramma Walther, 1904 | | | | `-- †K. paradoxum Liu et al., 2014 | | | |--o †Limnogramma Ren, 2003 | | | | |-- †L. hani | | | | |-– †L. mira | | | | `-- &38224;L. mongolicum | | | |--o †Meioneurites Handlirsch, 1906; L.Jur. CAs. | | | | |-- †M. (Parameioneurites) spectabilis Engel, 2005 | | | | |-- †M. schlosseri Handlirsch, 1906 | | | | `-- †M. villosus Panfilov, 1980 | | | |-- †Protokalligramma bifasciatum Yang, Makarkin & Ren, 2011 | | | `--o †Sophogramma Ren & Guo, 1996 | | | |-- †S. papilionaceum Ren & Guo, 1996 | | | `-- †S. wimbledoni Jepson, Makarkin & Coram, 2012 | | `--o †Kallihemerobiinae Ren & Engel, 2008 | | |-- †Apochrysogramma rotundum Yang, Makarkin & Ren, 2011 | | `-– †Kallihemerobius pleioneurus Ren & Oswald, 2002 | `--+-- Psychopsidae Handlirsch (silky lacewings) | `-- Nemopteridae (spoon-, thread-winged lacewings; lippukorennot) `--+--o †Osmylopsychopidae | `-- †Petropsychops superba Trias. Aust. |-- †Nymphitidae |-- †“Solenoptilidae” |-- †Brogniartiellidae `--+--o Nymphidae Rambur [incl. Myiodactylidae] (split-footed lacewings) | |-- †Pronymphes mengeanus (Hagen); M.Eoc. NEu. | |-- Nymphes | |-- Myiodactylus | |-- Osmylops | `== numerous other Recent genera `--+-- †Babinskaiidae |--o Myrmeleontidae Latreille [incl. †Palaeoleontidae] (antlions; muurahaiskorennot, -leijonat) | |?- †Samsonileon fragmentus Ponomarenko; L.Cret. SWAs. [†Palaeoleontidae] | |?- †Palaeoleon ferrogeneticus Rice; U.Cret. NENAm. [†Palaeoleontidae] | |?- †Dendroleon septemmontanus Statz, 1936 | |?-o Porrelus | | |-- †P. dominicanus Poinar & Stange, 1996; Mioc. CAm. | | `== recent Porrelus -species | |?- Dimarinae | |?- Stilbopteryginae | |?- Pseudimarinae | |?- Echthromyrmicinae | |?- Palparinae [incl. †Araripeneurinae] | |?- Myrmecaelurinae | |?- Acanthaclisinae | |?- Myrmeleontinae | |?- Brachynemurinae | |?- Dendroleontinae | |?- Glenurinae | `?- Nemoleontinae `--o Ascalaphidae Rambur (owlflies; pöllökorennot) |?- †Ascaloptynx oligocenicus Nel, 1990; Olig. WEu. |?- †Borgia proavus (Hagen, 1858) Navás, 1913; Olig. CEu. |?- †Mesascalaphus yangi Ren et al., 1995; U.Jur. EAs. |?- †Neadelphus protae MacLeod, 1970; M.Eoc. NEu. [larva] |?- †Prosuhpalacsa biamoensis Makarkin, 1998; ?U.Olig.-?L.Mioc. EAs. |?- †Ricartus edwardsi (Oustalet, 1870) Navás, 1913; Olig. WEu. |?- Cordulecerus |?- Albardiinae |--o Ascalaphinae | |--o Ululodes | | `-- †U. paleonesia Engel & Grimaldi, 2007; Mioc. CAm. | `== Other Ascalaphines `--o Haplogleniinae |--o Amoea | `-- †A. electrodominicana Engel & Grimaldi, 2007; Mioc. CAm. `== Other Haplogleniines
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