After Solodovnikov, Yue, Tarasov & Ren, 2012, with genera from “The Tree of Life” and “Wikispecies”
<==o Staphylininae Latreille, 1802 (aitolyhytsiipiset) |-- †Cretoquedius oculatus Ryvkin, 1988; L.Cret. EAs. |?- †Laasbium |--o †Laostaphylinus Zhang, 1988; E.Cret. EAs. | |-- †L. fuscus Zhang, 1988 | `-- †L. nigritellus Zhang, 1988 |--o †Mesostaphylinus Zhang, 1988; E.Cret. EAs. | |-- †M. fraternus Zhang, Wang & Xu, 1992 | `-- †M. laiyangensis Zhang, 1988 |?-o †Cretoquedius | `-- †C. oculatus Ryvkin, 1988; L.Cret. EAs. |?-o Maorothiini | `-- Maorothius |?-o Othiini | |-- Atrecus | |-- Othius | `-- Parothius |--+--o Platyprosopini | | `--o Platyprosopus | | |-- (Megaprosopoda) | | `-- (Platyprosopus) | |--o Diochini | | |-- Antarctothius | | |-- Coomania | | `-- Diochus | `--o Xantholinini | |-- Achemia | |-- Achmonia | |-- Adhavara | |-- Agerodes | |-- Allolinus | |-- Allotrichus | |-- Andamania | |-- Andelis | |-- Archaites | |-- Atopolinus | |-- Australinus | |-- Caecolinus | |-- Calontholinus | |-- Cibyra | |-- Crinolinus | |-- Cylindrinus | |-- Dactylaptatus | |-- Daolus | |-- Denon | |-- Dibothroglyptus | |-- Domea | |-- Eachamia | |-- Elgonia | |-- Enervia | |-- Erymus | |-- Eulissus | |-- Faxilla | |-- Gauropterus | |-- Grevillia | |-- Gyrohypnus | |-- Habrolinus | |-- Hesperolinus | |-- Heterocinus | |-- Heterolinus | |-- Himmala | |-- Holocorynus | |-- Homalolinus | |-- Homorocerus | |-- Hypnogyra | |-- Indolinus | |-- Indomorphus | |-- Lemiganus | |-- Lepidophallus | |-- Lepitacnus | |-- Lepta | |-- Leptacinellus | |-- Leptacinus | |-- Leptomicrus | |-- Leptophius | |-- Leurocorynus | |-- Linohesperus | |-- Linosomus | |-- Liotesba | |-- Lissohypnus | |-- Lithocharodes | |-- Maharadja | |-- Mahavana | |-- Manilla | |-- Medhiama | |-- Megalinus | |-- Metocinus | |-- Metolinus | |-- Metosina | |-- Microleptus | |-- Microlinus | |-- Mitomorphus | |-- Neohypnus | |-- Neoleptacinus | |-- Neoxantholinus | |-- Nepalinus | |-- Nilla | |-- Notolinus | |-- Nudobius | |-- Oculolabrus | |-- Otagonia | |-- Oxybleptes | |-- Pachycorynus | |-- Pahanghella | |-- Paracorynus | |-- Paratesba | |-- Paulianella | |-- Phacophallus | |-- Platydromus | |-- Plochionocerus | |-- Pseudocorynus | |-- Pseudoxantholinus | |-- Queenslandina | |-- Renda | |-- Sagarmatha | |-- Scytalinus | |-- Someira | |-- Somoleptus | |-- Spaniolinus | |-- Stenistoderus | |-- Stenolinus | |-- Stictolinus | |-- Sulawesina | |-- Sumatera | |-- Sungaria | |-- Sylea | |-- Symilla | |-- Talliella | |-- Tamilla | |-- Tesba | |-- Tetraulacus | |-- Thyreocephalus | |-- Tralichia | |-- Ulisseus | |-- Vulda | |-- Waitatia | |-- Walesia | |-- Whangareiella | |-- Xanthocorynus | |-- Xantholinus | |-- Xanthophius | |-- Xestolinus | |-- Yunna | |-- Yunnella | |-- Zenon | `-- Zeteotomus `--+--o †Thayeralinini | `--o †Thayeralinus | |-- †T. fraternus (Zhang, Wang et Xu, 1992) | |-- †T. giganteus | |-- †T. glandulifer | `--+-- †T. fieldi | `-- †T. longelytratus `--+--o Arrowinini Solodovnikov & Newton, 2005 | `-- Arrowinus Bernhauer 1935 `--o Staphylinini |--o Amblyopinina | |-- Amblyopinus | |-- Amblyopinodes | |-- Chilamblyopinus piceus | |-- Edrabius | |-- Megamblyopinus | `-- Myotyphlus jansoni |-- Anisolinina | |-- Amelinus | |-- Amichrotus | |-- Anisolinus | |-- Barygnathus | |-- Bombylodes | |-- Diatrechus | |-- Hesperoschema | |-- Hesperosoma | |-- Misantlius | |-- Pammegus | |-- Paratympanophorus | |-- Philomyceta | |-- Tolmerinus | |-- Turgiditarsus | `-- Tympanophorus |--o Hyptiomina Casey, 1906 | `-- Holisus |--o Philonthina | |-- Actinomorphus | |-- Actinus | |-- Afrorabigus | |-- Agacerus | |-- Atopocentrum | |-- Belonuchus | |-- Bisnius | |-- Cafius | |-- Chroaptomus | |-- Craspedomerus | |-- Diplostictus | |-- Dorcophilonthus | |-- Ecitophytes | |-- Endeius | |-- Erichsonius | |-- Flohria | |-- Gabriopalpus | |-- Gabrius | |-- Gabronthus | |-- Glyphesthus | |-- Hesperomimus | |-- Hesperopalpus | |-- Hesperus | |-- Hybridolinus | |-- Jurecekia | |-- Laetulonthus | |-- Leptopeltus | |-- Leucitus | |-- Linoderus | |-- Mentophilonthus | |-- Moeocerus | |-- Mysolius | |-- Neobisnius | |-- Neoleucitus | |-- Odontolinus | |-- Ophionthus | |-- Orthidus | |-- Pachypelmopus | |-- Paederallus | |-- Paederomimus | |-- Paracraspedomerus | |-- Pescolinus | |-- Peucoglyphus | |-- Phileciton | |-- Philonthus | |-- Phucobius | |-- Platyschema | |-- Proxenobius | |-- Pseudocraspedomerus | |-- Pseudomoeocerus | |-- Pterygolaetus | |-- Rabigus | |-- Remus | |-- Scelotrichus | |-- Shaverdolena | |-- Sphaeriolinus | |-- Sternotoxus | |-- Stevensia | |-- Taxiplagus | |-- Thinocafius | |-- Trachyphilonthus | |-- Tropiopterius | |-- Xanthodermus | `-- Xenobius |--o Quediina | |-- Acylohsellus | |-- Acylophorus | |-- Anaquedius | |-- Anchocerus | |-- Astrapaeus | |-- Beeria | |-- Bolitogyrus | |-- Cafioquedus | |-- Cheilocolpus | |-- Ctenandropus | |-- Euristus | |-- Euryporus | |-- Heinzia | |-- Hemiquedius | |-- Heterothops | |-- Indoquedius | |-- Ioma | |-- Loncovilius | |-- Lonia | |-- Mimosticus | |-- Natalignathus | |-- Paratolmerus | |-- Parisanopus | |-- Philonthellus | |-- Pseudorientis | |-- Quediocafus | |-- Quediomacrus | |-- Quediomimus | |-- Quediopsis | |-- Quedius | |-- Quelaestrygon | |-- Quetarsius | |-- Quwatanabius | |-- Rolla | |-- Sphingoquedius | |-- Strouhalium | |-- Termitoquedius | |-- Valdiviodes | |-- Velleiopsis | `-- Velleius |--o Staphylinina | |-- Abemus | |-- Agelosus | |-- Apecholinus | |-- Apostenolinus | |-- Aulacocypus | |-- Bafutella | |-- Collocypus | |-- Creophilus | |-- Dinothenarus | |-- Emus | |-- Eucibdelus | |-- Guillaumius | |-- Hadropinus | |-- Hadrotes | |-- Leistotrophus | |-- Liusus | |-- Menoedius | |-- Miobdelus | |-- Naddia | |-- Ocychinus | |-- Ocypus | |-- Ontholestes | |-- Palaestrinus | |-- Pancarpius | |-- Paragastrisus | |-- Parapalaestrinus | |-- Paraphytolinus | |-- Philetaerius | |-- Physetops | |-- Phytolinus | |-- Platydracus | |-- Protocypus | |-- Protogoerius | |-- Rhynchocheilus | |-- Rhyncocheilus | |-- Saniderus | |-- Sphaerobulbus | |-- Sphaeromacrops | |-- Staphylinus | |-- Tasgius | |-- Thinopinus | |-- Thoracostrongylus | |-- Trichocosmetes | `-- Wasmannellus |--o Tanygnathinina Reitter, 1909 | |-- Atanygnathus | `-- Natalignathus `--o Xanthopygina |-- Algon |-- Antimerus |-- Dysanellus |-- Elecatopselaphus |-- Elmas |-- Gastrisus |-- Glenus |-- Hasumius |-- Isanopus |-- Nausicotus |-- Nordus |-- Ocyolinus |-- Oligotergus |-- Paraxenopygus |-- Phanolinopsis |-- Phanolinus |-- Philothalpus |-- Plociopterus |-- Prianophthalmus |-- Prionopedinus |-- Prionophilonthus |-- Rientis |-- Scaponopselaphus |-- Styngetus |-- Torobus |-- Triacrus |-- Tricholinus |-- Trigonopalpus |-- Trigonopselaphus |-- Xanthopygoides |-- Xanthopygus `-- Xenopygus
Reference(s):
- Chatzimanolis, S., Grimaldi, D. A., Engel, M. S. & Fraser, N. C., 2012: Leehermania prorova, the earliest staphyliniform beetle, from the Late Triassic of Virginia (Coleoptera: Staphylinidae).
–American Museum Novitates: #3761, pp. 1-28 [doi: 10.1206/3761.2] - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968 - –iNet: The Tree of Life Project: http://tolweb.org/
- Solodovnikov, A., Yue, Y., Tarasov, S. & Ren, D., 2012: Extinct and extant rove beetles meet in the matrix: Early Cretaceous fossils shed light on the evolution of a hyperdiverse insect lineage (Coleoptera: Staphylinidae: Staphylininae).
–Cladistics: in press [doi: 10.1111/j.1096-0031.2012.00433.x] - Yue, Y., Zhao, Y. & Ren, D., 2009: Three new Mesozoic Staphylinids (Coleoptera) from Liaoning, China.
–Cretaceous Research: Vol. 31 (2010), #1, pp. 61-70 [doi: 10.1016/j.cretres.2009.09.005]