After Chatzimanolis, Engel, Newton & Grimaldi, 2009, with taxa from “Wikispecies”
<==o Scydmaeninae Leach, 1815 sensu Grebennikov & Newton, 2009 [Scydmaenidae Leach, 1815; Anisosphaeridae Tömösváry, 1882] |--o †Hapsomelitae Poinar & Brown, 2004 sensu Grebennikov & Newton, 2009 [Haplomelitinae Poinar & Brown, 2004] | |-- †Hapsomela burmitis Poinar & Brown, 2004 | |-- †Ektatotricha paradoxa Chatzimanolis, Engel & Newton, 2009 | `-- †Electroatopos castaneaus Chatzimanolis, Engel & Newton, 2009 |--o Mastigitae Fleming, 1821 sensu Grebennikov & Newton, 2009 [Mastiginae Fleming, 1821] | |--o Clidicini | | |-- †Aenictosoma doenitzi Schaufuss, 1891 | | |--o Clidicus Laporte, 1832 | | | |-- C. abbotensis | | | |-- C. aliquantulus | | | |-- †C. balticus | | | |-- C. bellator | | | |-- C. formicarius | | | |-- C. grandis Laporte, 1832 | | | |-- C. laticeps | | | |-- C. loebli | | | |-- C. monstrosus | | | |-- C. mussardi | | | |-- C. mysorensis | | | |-- C. omoios | | | |-- C. quadricollis | | | |-- C. rufescens | | | |-- C. taphrocephalus | | | |-- C. termitophilus | | | `-- C. tonkinensis | | |--o Leptochromus Motschulsky, 1855 | | | |-- †L. palaeomexicanus O'Keefe, 2002 | | | |-- L. agilis | | | |-- L. bicolor | | | |-- L. fulvescens Motschulsky, 1855 | | | `-- L. mexicanus | | |-- †Palaeoleptochromus schaufussi O'Keefe, in O'Keefe, Pike & Poinar, 1997 | | `--o Papusus Casey, 1897 | | |-- P. andrewsi | | |-- P. esparanza | | |-- P. guilianii | | |-- P. loebli | | |-- P. macer Casey, 1897 | | |-- P. naranjus | | |-- P. panamint | | |-- P. piceus | | `-- P. werneri | |--o Leptomastacini Casey, 1897 | | |-- Ablepton treforti Frivaldszky, 1877 | | |--o Leptomastax Pirazzoli, 1855 [Pyladus Fairmaire, 1856; Leptomastoides Karaman, 1962 (nomen nudum)] | | | |-- L. hypogeum Pirazzoli, 1855 | | | `== 21 sp. | | `--o Taurablepton Franz, 1988 | | |-- T. subterraneum Franz, 1988 | | `== 2 sp. | `--o Mastigini Fleming, 1821 | |--o Mastigus Latreille, 1802 | | |-- M. spinicornis (Fabricius, 1787) Latreille, 1802 [Ptinus spinicornis Fabricius, 1787] | | `== 5 sp. | |--o Palaeostigus Newton, in Newton & Franz, 1998 | | |-- P. palpalis Latreille, 1804) Newton, 1998 [Mastigus palpalis Latreille, 1804] | | `== 13 sp. | `--o Stenomastigus Leleup, 1968 | |--o (Acanthostigus) Leleup, 1968 | | |-- S. (A.) basilewskyi Leleup, 1968 | | `== 2 sp. | `--o (Stenomastigus) Leleup, 1968 | |-- S. (S.) franzi Leleup, 1968 | |-- S. (S.) holmi Franz, 1984 | `== 10 sp. `--o Scydmaenitae Leach, 1815 sensu Grebennikov & Newton, 2009 |-- †Kachinus antennatus Chatzimanolis, Engel & Newton, 2009 |--o Eutheniini | |--o Eutheia | | `-- E. puetzi | |--o Eutheimorphus Franz & Loebl, 1990 | | `-- E. paradoxus | |-- Euthiconus | |-- Paeneutheia pallida Jaloszynski, 2003 | |--o Paraneseuthia Franz, 1980 | | |-- P. devia | | |-- P. guineana | | `-- P. inexpectata | |-- Protoeuthia | `--o Veraphis Casey, 1897 | |-- V. hisamatsui | |-- V. horianus | |-- V. kurbatovi | |-- V. modestus | |-- V. odaigaharensis | |-- V. sawadai | |-- V. spinosus | |-- V. striga | `-- V. tottoriensis |--o Cephenniini | |--o Cephennium | | |-- C. anophthalmicum | | |-- C. aridum | | |-- C. canestroi | | |-- C. celsifrons | | |-- C. gilberti | | |-- C. grandarboreum | | |-- C. mariposae | | |-- C. urbanum | | `== other Recent species | |--o Cephennodes Reitter, 1883 | | |--o (Cephennodes) Reitter, 1883 [Chelonoides Croissandeau, 1894 [nec Hitchcock, 1858]; Coatesia Lea, 1915; Chelonoidum Strand, 1935 [nomen novum pro Chelonoides Croissandeau, 1894]] | | | |-- C. (C.) simonis Reitter, 1883 | | | |-- C. (C.) abdominalis | | | |-- C. (C.) araiorum | | | |-- C. (C.) baculifer | | | |-- C. (C.) bicolor | | | |-- C. (C.) brinchangensis | | | |-- C. (C.) carinifrons | | | |-- C. (C.) caudatus | | | |-- C. (C.) cooteri | | | |-- C. (C.) elytratus | | | |-- C. (C.) excavatus | | | |-- C. (C.) guadunanus | | | |-- C. (C.) hamatus | | | |-- C. (C.) hongkongensis | | | |-- C. (C.) impressifrons | | | |-- C. (C.) inflatipes | | | |-- C. (C.) kopeipes | | | |-- C. (C.) latus | | | |-- C. (C.) leai | | | |-- C. (C.) longipes | | | |-- C. (C.) masuanus | | | |-- C. (C.) moderatus | | | |-- C. (C.) nematocerus | | | |-- C. (C.) nigricollis | | | |-- C. (C.) nitens | | | |-- C. (C.) papuanus | | | |-- C. (C.) parabdominalis | | | |-- C. (C.) paramerus | | | |-- C. (C.) penicillatus | | | |-- C. (C.) pteroscapus | | | |-- C. (C.) pullatus | | | |-- C. (C.) scolytoides | | | |-- C. (C.) sichuanus | | | |-- C. (C.) simplicipes | | | |-- C. (C.) spatulipes | | | |-- C. (C.) spinosus | | | |-- C. (C.) subcaudatus | | | |-- C. (C.) superlatus | | | |-- C. (C.) tauroides | | | |-- C. (C.) taurus | | | |-- C. (C.) torosus | | | |-- C. (C.) triangulifrons | | | |-- C. (C.) tuberculifrons | | | |-- C. (C.) uenoi | | | `-- C. (C.) yunnanensis | | `--o (Fusionodes) | | |-- C. (F.) ascipenis | | |-- C. (F.) brachylinguis | | |-- C. (F.) graeseri | | |-- C. (F.) hippopotamus | | |-- C. (F.) loebli | | |-- C. (F.) longilinguis | | |-- C. (F.) lustrifrons | | |-- C. (F.) magnus | | |-- C. (F.) malleiphallus | | |-- C. (F.) microphthalmus | | |-- C. (F.) ovatus | | |-- C. (F.) rectangulicollis | | |-- C. (F.) schuelkei | | |-- C. (F.) serratus | | |-- C. (F.) testudo | | |-- C. (F.) transversicollis | | |-- C. (F.) ussuricus | | `-- C. (F.) wrasei | |--o Cephennomicrus Reitter, 1907 | | |-- C. affouchensis | | |-- C. afiamalui | | |-- C. anderssoni | | |-- C. asperus | | |-- C. cactiformis | | |-- C. caledonicus | | |-- C. cordithorax | | |-- C. cornutus | | |-- C. crenatus | | |-- C. dugdalei | | |-- C. fijii | | |-- C. fossulatus | | |-- C. fujianus | | |-- C. glaber | | |-- C. gomyi | | |-- C. hobbiti | | |-- C. impressus | | |-- C. inconspicuus | | |-- C. insularis | | |-- C. japonigenus | | |-- C. jucundus | | |-- C. latipennis | | |-- C. lauensis | | |-- C. longicornis | | |-- C. lordhowei | | |-- C. mauritiensis | | |-- C. minimus | | |-- C. minor | | |-- C. nomurai | | |-- C. okinawanus | | |-- C. pauliani | | |-- C. perexiguus | | |-- C. perpunctillus Reitter, 1907 | | |-- C. perthi | | |-- C. politus | | |-- C. pusillus | | |-- C. raffrayi | | |-- C. rugosicollis | | |-- C. samoanus | | |-- C. sumatranus | | |-- C. suturalis | | |-- C. taiwanensis | | |-- C. typicus | | |-- C. upoluensis | | |-- C. vadoni | | |-- C. validus | | |-- C. venuensis | | |-- C. vitiensis | | |-- C. zimmermani | | `-- C. zoiai | |-- Etelea | |--o Hlavaciellus Jaloszynski, 2006 | | |-- H. sabahensis | | `-- H. vampirus Jaloszynski, 2006 | |-- Nanophthalmus Motschulsky | |-- Paracephennium O'Keefe, 1999 | |-- Pseudocephennium | `-- Trurlia insana Jaloszynski, 2009 |-- Chevrolatia Jacquelin du Val [Chevrolatiini] |-- Leptoscydmus [Leptoscydmini] |-- Plaumanniola sanctaecatharinae Costa Lima, 1962 [Plaumanniolini] |--o Cyrtoscydmini | |-- Allohoraeomorphus calcarifer Franz, 1986 | |--o Alloraphes Franz, 1980 | | |-- A. bolivarensis | | |-- A. chiapasensis | | |-- A. jamaicae | | |-- A. lenkoi | | |-- A. magnus | | |-- A. myrmecophilus | | |-- A. peruanus | | `-- A. yucatani | |--o Anhoraeomorphus Franz, 1986 | | |-- A. assimilis | | `-- A. obscurus | |--o Archiconnus Franz, 1980 | | |-- A. huallaganus | | `-- A. mexicanus | |-- Atropidus | |-- Brachycepsis | |-- Catalinus | |-- Chathamaenus chathamensis Franz, 1980 [Chathamneus Emberson, 1998 (lapsus calami)] | |--o Elacatophora Schaufuss, 1884 | | |-- E. apion | | |-- E. dissimilis | | |-- E. javanica | | |-- E. longipes | | |-- E. malaysiae (Franz, 1984) [Euconnus malaysiae Franz, 1984; Borneosabahia mirifica Franz, 1992] | | |-- E. maxima | | `-- E. robusta Schaufuss, 1884 | |-- Euconnomorphus pilosissimus Franz, 1980 | |--o Euconnus Thomson, 1859 | | |-- (Alloconophron) Franz, 1986 | | |-- (Allonapochus) Franz, 1986 | | |-- (Androconnus) Franz, 1986 | | |--o (Anthicimorphus) Franz, 1975 | | | |-- E. (A.) anthicoides (Lea, 1914) Franz, 1975 [Phagonophana anthicoides Lea, 1914] | | | `-- E. (A.) aequalis | | |-- (Borneoconnus) Franz, 1992 | | |-- (Cephaloconnus) | | |-- (Cerviconnus) | | |-- (Cladoconnus) | | |--o (Dimorphoconnus) Franz, 1975 | | | `-- E. (D.) insigniventris (Lea, 1913) Franz, 1975 [Scydmaenus insigniventris Lea, 1913] | | |-- (Drastophus) | | |--o (Euconnus) Thomson, 1862 | | | `-- E. (E.) hirticollis Illiger, 1798 | | |--o (Euconophron) | | | |-- E. (E.) brevipennis | | | `-- E. (E.) howeanus | | |-- (Eupentarius) | | |-- (Filonapochus) | | |-- (Heteroconnus) | | |--o (Heterotetramelus) Frans, 1971 | | | |-- E. (H.) baloghianus | | | `-- E. (H.) carteri | | |-- (Himaloconnus) | | |-- (Microconnus) | | |-- (Myrmecodytes) | | |-- (Napochus) | | |-- (Napoconnus) | | |-- (Neonapochus) | | |-- (Nepaloconnus) | | |-- (Noctophus) | | |-- (Nodoconnus) | | |-- (Palaeoconnus) | | |-- (Paranapochus) | | |-- (Paratetramelus) | | |-- (Psomophus) | | |-- (Pycnophus) | | |-- (Rhomboconnus) Franz, 1986 | | |-- (Scopophus) | | |-- (Scydmaenites) | | |-- (Smicrophus) | | |--o (Tetramelus) | | | |-- E. (T.) castawayensis | | | |-- E. (T.) pseudoramsayi | | | |-- E. (T.) ramsayi | | | |-- E. (T.) threekingensis | | | |?- E. (T.) curvicrus | | | |?- E. (T.) northlandensis | | | |?- E. (T.) picicollis | | | |?- E. (T.) stewartensis | | | |?- E. (T.) eremita | | | |?- E. (T.) ohenae | | | `?- E. (T.) wairauensis | | |-- (Ursoconnus) Franz, 1963 | | |--o (Valdivioconnus) Frans, 1980 | | | `-- E. (V.) auctor | | `-- (Xestophus) | |--o Heteroscydmus Frans, 1980 | | `-- H. yapacariensis | |-- Homoconnus | |--o Horaeomorphus Schaufuss | | |-- (Horaeomorphus) Schaufuss | | `-- (Pseudosyndicus) | |-- Leptocharis | |--o Loeblites Frans, 1986 | | |-- L. mastigicornis | | |-- L. minor | | `-- L. sabahensis | |-- Lophioderus | |-- Madagassoconnus Franz, 1986 | |-- Magellanoconnus (Franz, 1967) Kuschel, 1990 [Euconnus (Magellanoconnus) Franz, 1967] | |-- Maorinus (Franz, 1980) Kuschel, 1990 [nomen novum pro Maoria Franz, 1975] [Euconnus (Maoria) Franz, 1975] | |--o Microraphes Franz, 1980 | | `-- M. pygmaeus | |--o Microscydmus Saulcy & Croissandeau, 1893 | | |-- (Delius) | | |--o (Microscydmus) | | | |-- M. (M.) australiensis | | | |-- M. (M.) bicavatus | | | |-- M. (M.) lynfieldi | | | |-- M. (M.) nasicornis | | | |-- M. (M.) nasutus | | | `-- M. (M.) perpusillus | | |-- (Neladius) | | |-- (Neoscydmus) | | `--o (Parastenichnus) Frans, 1970 | | |-- M. (P.) singaporensis | | `-- M. (P.) unicus | |--o Mimoscydmus Frans, 1980 | | |-- M. baruerii | | `-- M. brasiliensis | |--o Napochomorphus Frans, 1980 | | `-- N. extraneus | |-- Neuraphanax | |--o Neuraphes | | |-- (Neuraphes) | | |-- (Pararaphes) | | `-- (Rhynchoraphes) | |-- Neuraphoconnus | |-- Neuraphomorphus | |-- Oneila | |--o Parapseudoconnus Franz, 1980 | | |-- (Neuraphomimus) | | `-- (Parapseudoconnus) | |-- Parascydmus | |--o Parastenichnaphes Franz, 1984 | | |-- P. ceylonensis | | |?- ?P. myrmecophilus | | `-- P. sumatrensis | |--o Protandroconnus Franz, 1989 | | `-- P. peruensis | |-- Protoconnus | |-- Protoscydmus | |-- Psepharobius | |-- Pseudoraphes Franz, 1980 | |--o Sciacharis Broun, 1893 | | |-- (Anthicimimus) | | |-- (Sciacharis) | | `-- (Valdivioconnus) | |-- Scydmoraphes | |-- Siamites | |--o Stenichnaphes Franz, 1980 | | |-- S. madagassicus | | |-- S. newtoni | | |-- S. sakarahanus | | |-- S. similis | | `-- S. urbanus Franz, 1980 | |-- Stenichnoconnus Franz, 1985 | |-- Stenichnodes Franz, 1966 | |-- Stenichnoteras | |--o Stenichnus Thomson, 1859 | | |-- (Austrostenichnus) | | |-- (Scydmaenichnus) | | |-- (Scydmaenilla) | | `-- (Stenichnus) | |--o Syndicus [Syndicomorphus Franz, 1992] | | |-- (Semisyndicus) | | `-- (Syndicus) | |-- Taphroscydmus | |--o Venezolanoconnus Franz, 1987 | | |-- V. andinus | | |-- V. caracasensis | | |-- V. minor | | `-- V. ranchoi | `-- Vinsoniana `--o Scydmaenini |--o Adrastia Broun, 1881 | |-- A. angulifrons | |-- A. angustissima | |-- A. anophthalma | |-- A. anthicoides | |-- A. brookesi | |-- A. brouni | |-- A. clarkei | |-- A. clavatella | |-- A. curticornis | |-- A. decipiens | |-- A. dentipes | |-- A. edwardsi | |-- A. gourlayi | |-- A. greymouthi | |-- A. haastensis | |-- A. laetans | |-- A. nelsoni | |-- A. novaezeelandiae | |-- A. paravillosipennis | |-- A. peckorum | |-- A. stokesi | |-- A. subinermis | |-- A. townsendi | |-- A. wilmotensis | `-- A. villosipennis |-- Ceramphis |-- Clavigeroscydmus |-- Eudesis |-- Palaeoscydmaenus australiensis Franz, 1975 |--o Pseudoeudesis | |-- (Afroeudesis) | |-- (Oreoeudesis) | `-- (Pseudoeudesis) `--o Scydmaenus Latreille, 1802 |-- (Afroscydmaenus) |-- (Agathelor) |-- (Alaudula) |-- (Androscydmaenus) |-- (Androscydmus) |-- (Archiscydmaenus) |-- (Armatoscydmaenus) |-- (Breviclavatoscydmaenus) |-- (Choleropsis) |-- (Cholerus) |-- (Conoscydmaenus) |-- (Corbulifer) |-- (Eustemmoides) |-- (Eustemmus) |-- (Geoscydmaenus) |-- (Heteroeustemmus) |-- (Heteromicrus) |-- (Kingius) |-- (Mascarensia) |-- (Mimoscydmaenus) |-- (Nepaloscydmaenus) |-- (Paraeustemmus) |-- (Parageoscydmaenus) |-- (Parallomicrus) |-- (Paracydmaenus) |-- (Pseudeumicrus) |-- (Scottiscydmaenus) Franz, 1975 |-- (Scydmaenus) |-- (Seychellinus) Franz, 1986 |-- (Termitoscydmaenus) Franz, 1980 `-- (Trapezoscydmaenus) Franz, 1986
Reference(s):
- Anonymous, 2009: Scydmaeninae –iNet: Wikispecies, http://species.wikimedia.org/wiki/Scydmaeninae
- Chatzimanolis, S., Engel, M. S., Newton, A. F. & Grimaldi, D. A., 2009: New ant-like stone beetles in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Scydmaeninae).
–Cretaceous Research: Vol. 31, #1, pp. 77-84 [doi: 10.1016/j.cretres.2009.09.009] - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968