After Chatzimanolis, Engel, Newton & Grimaldi, 2009, with taxa from “Wikispecies”

<==o Scydmaeninae Leach, 1815 sensu Grebennikov & Newton, 2009 [Scydmaenidae Leach, 1815; Anisosphaeridae Tömösváry, 1882]
   |--o †Hapsomelitae Poinar & Brown, 2004 sensu Grebennikov & Newton, 2009 [Haplomelitinae Poinar & Brown, 2004]
   |  |-- †Hapsomela burmitis Poinar & Brown, 2004
   |  |-- †Ektatotricha paradoxa Chatzimanolis, Engel & Newton, 2009
   |  `-- †Electroatopos castaneaus Chatzimanolis, Engel & Newton, 2009
   |--o Mastigitae Fleming, 1821 sensu Grebennikov & Newton, 2009 [Mastiginae Fleming, 1821]
   |  |--o Clidicini
   |  |  |-- †Aenictosoma doenitzi Schaufuss, 1891
   |  |  |--o Clidicus Laporte, 1832
   |  |  |  |-- C. abbotensis
   |  |  |  |-- C. aliquantulus
   |  |  |  |-- †C. balticus
   |  |  |  |-- C. bellator
   |  |  |  |-- C. formicarius
   |  |  |  |-- C. grandis Laporte, 1832
   |  |  |  |-- C. laticeps
   |  |  |  |-- C. loebli
   |  |  |  |-- C. monstrosus
   |  |  |  |-- C. mussardi
   |  |  |  |-- C. mysorensis
   |  |  |  |-- C. omoios
   |  |  |  |-- C. quadricollis
   |  |  |  |-- C. rufescens
   |  |  |  |-- C. taphrocephalus
   |  |  |  |-- C. termitophilus
   |  |  |  `-- C. tonkinensis
   |  |  |--o Leptochromus Motschulsky, 1855
   |  |  |  |-- †L. palaeomexicanus O'Keefe, 2002
   |  |  |  |-- L. agilis
   |  |  |  |-- L. bicolor
   |  |  |  |-- L. fulvescens Motschulsky, 1855
   |  |  |  `-- L. mexicanus
   |  |  |-- †Palaeoleptochromus schaufussi O'Keefe, in O'Keefe, Pike & Poinar, 1997
   |  |  `--o Papusus Casey, 1897
   |  |     |-- P. andrewsi
   |  |     |-- P. esparanza
   |  |     |-- P. guilianii
   |  |     |-- P. loebli
   |  |     |-- P. macer Casey, 1897
   |  |     |-- P. naranjus
   |  |     |-- P. panamint
   |  |     |-- P. piceus
   |  |     `-- P. werneri
   |  |--o Leptomastacini Casey, 1897
   |  |  |-- Ablepton treforti Frivaldszky, 1877
   |  |  |--o Leptomastax Pirazzoli, 1855 [Pyladus Fairmaire, 1856; Leptomastoides Karaman, 1962 (nomen nudum)]
   |  |  |  |-- L. hypogeum Pirazzoli, 1855
   |  |  |  `== 21 sp.
   |  |  `--o Taurablepton  Franz, 1988
   |  |     |-- T. subterraneum Franz, 1988
   |  |     `== 2 sp.
   |  `--o Mastigini Fleming, 1821
   |     |--o Mastigus Latreille, 1802
   |     |  |-- M. spinicornis (Fabricius, 1787) Latreille, 1802 [Ptinus spinicornis Fabricius, 1787]
   |     |  `== 5 sp.
   |     |--o Palaeostigus Newton, in Newton & Franz, 1998
   |     |  |-- P. palpalis Latreille, 1804) Newton, 1998 [Mastigus palpalis Latreille, 1804]
   |     |  `== 13 sp.
   |     `--o Stenomastigus Leleup, 1968
   |        |--o (Acanthostigus) Leleup, 1968
   |        |  |-- S. (A.) basilewskyi Leleup, 1968
   |        |  `== 2 sp.
   |        `--o (Stenomastigus) Leleup, 1968
   |           |-- S. (S.) franzi Leleup, 1968
   |           |-- S. (S.) holmi Franz, 1984
   |           `== 10 sp.
   `--o Scydmaenitae Leach, 1815 sensu Grebennikov & Newton, 2009
      |-- †Kachinus antennatus Chatzimanolis, Engel & Newton, 2009
      |--o Eutheniini
      |  |--o Eutheia
      |  |  `-- E. puetzi
      |  |--o Eutheimorphus Franz & Loebl, 1990
      |  |  `-- E. paradoxus
      |  |-- Euthiconus
      |  |-- Paeneutheia pallida Jaloszynski, 2003
      |  |--o Paraneseuthia Franz, 1980
      |  |  |-- P. devia
      |  |  |-- P. guineana
      |  |  `-- P. inexpectata
      |  |-- Protoeuthia
      |  `--o Veraphis Casey, 1897
      |     |-- V. hisamatsui
      |     |-- V. horianus
      |     |-- V. kurbatovi
      |     |-- V. modestus
      |     |-- V. odaigaharensis
      |     |-- V. sawadai
      |     |-- V. spinosus
      |     |-- V. striga
      |     `-- V. tottoriensis
      |--o Cephenniini
      |  |--o Cephennium
      |  |  |-- C. anophthalmicum
      |  |  |-- C. aridum
      |  |  |-- C. canestroi
      |  |  |-- C. celsifrons
      |  |  |-- C. gilberti
      |  |  |-- C. grandarboreum
      |  |  |-- C. mariposae
      |  |  |-- C. urbanum
      |  |  `== other Recent species
      |  |--o Cephennodes Reitter, 1883
      |  |  |--o (Cephennodes) Reitter, 1883 [Chelonoides Croissandeau, 1894 [nec Hitchcock, 1858]; Coatesia Lea, 1915; Chelonoidum Strand, 1935 [nomen novum pro Chelonoides Croissandeau, 1894]]
      |  |  |  |-- C. (C.) simonis Reitter, 1883
      |  |  |  |-- C. (C.) abdominalis
      |  |  |  |-- C. (C.) araiorum
      |  |  |  |-- C. (C.) baculifer
      |  |  |  |-- C. (C.) bicolor
      |  |  |  |-- C. (C.) brinchangensis
      |  |  |  |-- C. (C.) carinifrons
      |  |  |  |-- C. (C.) caudatus
      |  |  |  |-- C. (C.) cooteri
      |  |  |  |-- C. (C.) elytratus
      |  |  |  |-- C. (C.) excavatus
      |  |  |  |-- C. (C.) guadunanus
      |  |  |  |-- C. (C.) hamatus
      |  |  |  |-- C. (C.) hongkongensis
      |  |  |  |-- C. (C.) impressifrons
      |  |  |  |-- C. (C.) inflatipes
      |  |  |  |-- C. (C.) kopeipes
      |  |  |  |-- C. (C.) latus
      |  |  |  |-- C. (C.) leai
      |  |  |  |-- C. (C.) longipes
      |  |  |  |-- C. (C.) masuanus
      |  |  |  |-- C. (C.) moderatus
      |  |  |  |-- C. (C.) nematocerus
      |  |  |  |-- C. (C.) nigricollis
      |  |  |  |-- C. (C.) nitens
      |  |  |  |-- C. (C.) papuanus
      |  |  |  |-- C. (C.) parabdominalis
      |  |  |  |-- C. (C.) paramerus
      |  |  |  |-- C. (C.) penicillatus
      |  |  |  |-- C. (C.) pteroscapus
      |  |  |  |-- C. (C.) pullatus
      |  |  |  |-- C. (C.) scolytoides
      |  |  |  |-- C. (C.) sichuanus
      |  |  |  |-- C. (C.) simplicipes
      |  |  |  |-- C. (C.) spatulipes
      |  |  |  |-- C. (C.) spinosus
      |  |  |  |-- C. (C.) subcaudatus
      |  |  |  |-- C. (C.) superlatus
      |  |  |  |-- C. (C.) tauroides
      |  |  |  |-- C. (C.) taurus
      |  |  |  |-- C. (C.) torosus
      |  |  |  |-- C. (C.) triangulifrons
      |  |  |  |-- C. (C.) tuberculifrons
      |  |  |  |-- C. (C.) uenoi
      |  |  |  `-- C. (C.) yunnanensis
      |  |  `--o (Fusionodes)
      |  |     |-- C. (F.) ascipenis
      |  |     |-- C. (F.) brachylinguis
      |  |     |-- C. (F.) graeseri
      |  |     |-- C. (F.) hippopotamus
      |  |     |-- C. (F.) loebli
      |  |     |-- C. (F.) longilinguis
      |  |     |-- C. (F.) lustrifrons
      |  |     |-- C. (F.) magnus
      |  |     |-- C. (F.) malleiphallus
      |  |     |-- C. (F.) microphthalmus
      |  |     |-- C. (F.) ovatus
      |  |     |-- C. (F.) rectangulicollis
      |  |     |-- C. (F.) schuelkei
      |  |     |-- C. (F.) serratus
      |  |     |-- C. (F.) testudo
      |  |     |-- C. (F.) transversicollis
      |  |     |-- C. (F.) ussuricus
      |  |     `-- C. (F.) wrasei
      |  |--o Cephennomicrus Reitter, 1907
      |  |  |-- C. affouchensis
      |  |  |-- C. afiamalui
      |  |  |-- C. anderssoni
      |  |  |-- C. asperus
      |  |  |-- C. cactiformis
      |  |  |-- C. caledonicus
      |  |  |-- C. cordithorax
      |  |  |-- C. cornutus
      |  |  |-- C. crenatus
      |  |  |-- C. dugdalei
      |  |  |-- C. fijii
      |  |  |-- C. fossulatus
      |  |  |-- C. fujianus
      |  |  |-- C. glaber
      |  |  |-- C. gomyi
      |  |  |-- C. hobbiti
      |  |  |-- C. impressus
      |  |  |-- C. inconspicuus
      |  |  |-- C. insularis
      |  |  |-- C. japonigenus
      |  |  |-- C. jucundus
      |  |  |-- C. latipennis
      |  |  |-- C. lauensis
      |  |  |-- C. longicornis
      |  |  |-- C. lordhowei
      |  |  |-- C. mauritiensis
      |  |  |-- C. minimus
      |  |  |-- C. minor
      |  |  |-- C. nomurai
      |  |  |-- C. okinawanus
      |  |  |-- C. pauliani
      |  |  |-- C. perexiguus
      |  |  |-- C. perpunctillus Reitter, 1907
      |  |  |-- C. perthi
      |  |  |-- C. politus
      |  |  |-- C. pusillus
      |  |  |-- C. raffrayi
      |  |  |-- C. rugosicollis
      |  |  |-- C. samoanus
      |  |  |-- C. sumatranus
      |  |  |-- C. suturalis
      |  |  |-- C. taiwanensis
      |  |  |-- C. typicus
      |  |  |-- C. upoluensis
      |  |  |-- C. vadoni
      |  |  |-- C. validus
      |  |  |-- C. venuensis
      |  |  |-- C. vitiensis
      |  |  |-- C. zimmermani
      |  |  `-- C. zoiai
      |  |-- Etelea
      |  |--o Hlavaciellus Jaloszynski, 2006
      |  |  |-- H. sabahensis
      |  |  `-- H. vampirus Jaloszynski, 2006
      |  |-- Nanophthalmus Motschulsky
      |  |-- Paracephennium O'Keefe, 1999
      |  |-- Pseudocephennium
      |  `-- Trurlia insana Jaloszynski, 2009
      |-- Chevrolatia Jacquelin du Val [Chevrolatiini]
      |-- Leptoscydmus [Leptoscydmini]
      |-- Plaumanniola sanctaecatharinae Costa Lima, 1962 [Plaumanniolini]
      |--o Cyrtoscydmini
      |  |-- Allohoraeomorphus calcarifer Franz, 1986
      |  |--o Alloraphes Franz, 1980
      |  |  |-- A. bolivarensis
      |  |  |-- A. chiapasensis
      |  |  |-- A. jamaicae
      |  |  |-- A. lenkoi
      |  |  |-- A. magnus
      |  |  |-- A. myrmecophilus
      |  |  |-- A. peruanus
      |  |  `-- A. yucatani
      |  |--o Anhoraeomorphus Franz, 1986
      |  |  |-- A. assimilis
      |  |  `-- A. obscurus
      |  |--o Archiconnus Franz, 1980
      |  |  |-- A. huallaganus
      |  |  `-- A. mexicanus
      |  |-- Atropidus
      |  |-- Brachycepsis
      |  |-- Catalinus
      |  |-- Chathamaenus chathamensis Franz, 1980 [Chathamneus Emberson, 1998 (lapsus calami)]
      |  |--o Elacatophora Schaufuss, 1884
      |  |  |-- E. apion
      |  |  |-- E. dissimilis
      |  |  |-- E. javanica
      |  |  |-- E. longipes
      |  |  |-- E. malaysiae (Franz, 1984) [Euconnus malaysiae Franz, 1984; Borneosabahia mirifica Franz, 1992]
      |  |  |-- E. maxima
      |  |  `-- E. robusta Schaufuss, 1884
      |  |-- Euconnomorphus pilosissimus Franz, 1980
      |  |--o Euconnus Thomson, 1859
      |  |  |-- (Alloconophron) Franz, 1986
      |  |  |-- (Allonapochus) Franz, 1986
      |  |  |-- (Androconnus) Franz, 1986
      |  |  |--o (Anthicimorphus) Franz, 1975
      |  |  |  |-- E. (A.) anthicoides (Lea, 1914) Franz, 1975 [Phagonophana anthicoides Lea, 1914]
      |  |  |  `-- E. (A.) aequalis
      |  |  |-- (Borneoconnus) Franz, 1992
      |  |  |-- (Cephaloconnus)
      |  |  |-- (Cerviconnus)
      |  |  |-- (Cladoconnus)
      |  |  |--o (Dimorphoconnus) Franz, 1975
      |  |  |  `-- E. (D.) insigniventris (Lea, 1913) Franz, 1975 [Scydmaenus insigniventris Lea, 1913]
      |  |  |-- (Drastophus)
      |  |  |--o (Euconnus) Thomson, 1862
      |  |  |  `-- E. (E.) hirticollis Illiger, 1798
      |  |  |--o (Euconophron)
      |  |  |  |-- E. (E.) brevipennis
      |  |  |  `-- E. (E.) howeanus
      |  |  |-- (Eupentarius)
      |  |  |-- (Filonapochus)
      |  |  |-- (Heteroconnus)
      |  |  |--o (Heterotetramelus) Frans, 1971
      |  |  |  |-- E. (H.) baloghianus
      |  |  |  `-- E. (H.) carteri
      |  |  |-- (Himaloconnus)
      |  |  |-- (Microconnus)
      |  |  |-- (Myrmecodytes)
      |  |  |-- (Napochus)
      |  |  |-- (Napoconnus)
      |  |  |-- (Neonapochus)
      |  |  |-- (Nepaloconnus)
      |  |  |-- (Noctophus)
      |  |  |-- (Nodoconnus)
      |  |  |-- (Palaeoconnus)
      |  |  |-- (Paranapochus)
      |  |  |-- (Paratetramelus)
      |  |  |-- (Psomophus)
      |  |  |-- (Pycnophus)
      |  |  |-- (Rhomboconnus) Franz, 1986
      |  |  |-- (Scopophus)
      |  |  |-- (Scydmaenites)
      |  |  |-- (Smicrophus)
      |  |  |--o (Tetramelus)
      |  |  |  |-- E. (T.) castawayensis
      |  |  |  |-- E. (T.) pseudoramsayi
      |  |  |  |-- E. (T.) ramsayi
      |  |  |  |-- E. (T.) threekingensis
      |  |  |  |?- E. (T.) curvicrus
      |  |  |  |?- E. (T.) northlandensis
      |  |  |  |?- E. (T.) picicollis
      |  |  |  |?- E. (T.) stewartensis
      |  |  |  |?- E. (T.) eremita
      |  |  |  |?- E. (T.) ohenae
      |  |  |  `?- E. (T.) wairauensis
      |  |  |-- (Ursoconnus) Franz, 1963
      |  |  |--o (Valdivioconnus) Frans, 1980
      |  |  |  `-- E. (V.) auctor
      |  |  `-- (Xestophus)
      |  |--o Heteroscydmus Frans, 1980
      |  |  `-- H. yapacariensis
      |  |-- Homoconnus
      |  |--o Horaeomorphus Schaufuss
      |  |  |-- (Horaeomorphus) Schaufuss
      |  |  `-- (Pseudosyndicus)
      |  |-- Leptocharis
      |  |--o Loeblites Frans, 1986
      |  |  |-- L. mastigicornis
      |  |  |-- L. minor
      |  |  `-- L. sabahensis
      |  |-- Lophioderus
      |  |-- Madagassoconnus Franz, 1986
      |  |-- Magellanoconnus (Franz, 1967) Kuschel, 1990 [Euconnus (Magellanoconnus) Franz, 1967]
      |  |-- Maorinus (Franz, 1980) Kuschel, 1990 [nomen novum pro Maoria Franz, 1975] [Euconnus (Maoria) Franz, 1975]
      |  |--o Microraphes Franz, 1980
      |  |  `-- M. pygmaeus
      |  |--o Microscydmus Saulcy & Croissandeau, 1893
      |  |  |-- (Delius)
      |  |  |--o (Microscydmus)
      |  |  |  |-- M. (M.) australiensis
      |  |  |  |-- M. (M.) bicavatus
      |  |  |  |-- M. (M.) lynfieldi
      |  |  |  |-- M. (M.) nasicornis
      |  |  |  |-- M. (M.) nasutus
      |  |  |  `-- M. (M.) perpusillus
      |  |  |-- (Neladius)
      |  |  |-- (Neoscydmus)
      |  |  `--o (Parastenichnus) Frans, 1970
      |  |     |-- M. (P.) singaporensis
      |  |     `-- M. (P.) unicus
      |  |--o Mimoscydmus Frans, 1980
      |  |  |-- M. baruerii
      |  |  `-- M. brasiliensis
      |  |--o Napochomorphus Frans, 1980
      |  |  `-- N. extraneus
      |  |-- Neuraphanax
      |  |--o Neuraphes
      |  |  |-- (Neuraphes)
      |  |  |-- (Pararaphes)
      |  |  `-- (Rhynchoraphes)
      |  |-- Neuraphoconnus
      |  |-- Neuraphomorphus
      |  |-- Oneila
      |  |--o Parapseudoconnus Franz, 1980
      |  |  |-- (Neuraphomimus)
      |  |  `-- (Parapseudoconnus)
      |  |-- Parascydmus
      |  |--o Parastenichnaphes Franz, 1984
      |  |  |-- P. ceylonensis
      |  |  |?- ?P. myrmecophilus
      |  |  `-- P. sumatrensis
      |  |--o Protandroconnus Franz, 1989
      |  |  `-- P. peruensis
      |  |-- Protoconnus
      |  |-- Protoscydmus
      |  |-- Psepharobius
      |  |-- Pseudoraphes Franz, 1980
      |  |--o Sciacharis Broun, 1893
      |  |  |-- (Anthicimimus)
      |  |  |-- (Sciacharis)
      |  |  `-- (Valdivioconnus)
      |  |-- Scydmoraphes
      |  |-- Siamites
      |  |--o Stenichnaphes Franz, 1980
      |  |  |-- S. madagassicus
      |  |  |-- S. newtoni
      |  |  |-- S. sakarahanus
      |  |  |-- S. similis
      |  |  `-- S. urbanus Franz, 1980
      |  |-- Stenichnoconnus Franz, 1985
      |  |-- Stenichnodes Franz, 1966
      |  |-- Stenichnoteras
      |  |--o Stenichnus Thomson, 1859
      |  |  |-- (Austrostenichnus)
      |  |  |-- (Scydmaenichnus)
      |  |  |-- (Scydmaenilla)
      |  |  `-- (Stenichnus)
      |  |--o Syndicus [Syndicomorphus Franz, 1992]
      |  |  |-- (Semisyndicus)
      |  |  `-- (Syndicus)
      |  |-- Taphroscydmus
      |  |--o Venezolanoconnus Franz, 1987
      |  |  |-- V. andinus
      |  |  |-- V. caracasensis
      |  |  |-- V. minor
      |  |  `-- V. ranchoi
      |  `-- Vinsoniana
      `--o Scydmaenini
         |--o Adrastia Broun, 1881
         |  |-- A. angulifrons
         |  |-- A. angustissima
         |  |-- A. anophthalma
         |  |-- A. anthicoides
         |  |-- A. brookesi
         |  |-- A. brouni
         |  |-- A. clarkei
         |  |-- A. clavatella
         |  |-- A. curticornis
         |  |-- A. decipiens
         |  |-- A. dentipes
         |  |-- A. edwardsi
         |  |-- A. gourlayi
         |  |-- A. greymouthi
         |  |-- A. haastensis
         |  |-- A. laetans
         |  |-- A. nelsoni
         |  |-- A. novaezeelandiae
         |  |-- A. paravillosipennis
         |  |-- A. peckorum
         |  |-- A. stokesi
         |  |-- A. subinermis
         |  |-- A. townsendi
         |  |-- A. wilmotensis
         |  `-- A. villosipennis
         |-- Ceramphis
         |-- Clavigeroscydmus
         |-- Eudesis
         |-- Palaeoscydmaenus australiensis Franz, 1975
         |--o Pseudoeudesis
         |  |-- (Afroeudesis)
         |  |-- (Oreoeudesis)
         |  `-- (Pseudoeudesis)
         `--o Scydmaenus Latreille, 1802
            |-- (Afroscydmaenus)
            |-- (Agathelor)
            |-- (Alaudula)
            |-- (Androscydmaenus)
            |-- (Androscydmus)
            |-- (Archiscydmaenus)
            |-- (Armatoscydmaenus)
            |-- (Breviclavatoscydmaenus)
            |-- (Choleropsis)
            |-- (Cholerus)
            |-- (Conoscydmaenus)
            |-- (Corbulifer)
            |-- (Eustemmoides)
            |-- (Eustemmus)
            |-- (Geoscydmaenus)
            |-- (Heteroeustemmus)
            |-- (Heteromicrus)
            |-- (Kingius)
            |-- (Mascarensia)
            |-- (Mimoscydmaenus)
            |-- (Nepaloscydmaenus)
            |-- (Paraeustemmus)
            |-- (Parageoscydmaenus)
            |-- (Parallomicrus)
            |-- (Paracydmaenus)
            |-- (Pseudeumicrus)
            |-- (Scottiscydmaenus) Franz, 1975
            |-- (Scydmaenus)
            |-- (Seychellinus) Franz, 1986
            |-- (Termitoscydmaenus) Franz, 1980
            `-- (Trapezoscydmaenus) Franz, 1986

Reference(s):