Scarabaeoidea

After Parker, 1982, “The Tree of Life”, and Grimaldi & Engel, 2005

<==o Scarabaeoidea (lehtisarviset, turilasmaiset kovakuoriaiset)
   |-- Glaresidae
   `--+--o Scarabaeidae (scarabs; lehtisarviset)
      |  |?-o †Holcorobeus U.Jur.-Cret.
      |  |  |-- †H. vittatus
      |  |  `-- †H. nigrimontanus
      |  |-- Cetoniinae
      |  |-- Dynastinae
      |  |-- Rutelinae
      |  |-- Melolonthinae
      |  |-- Aphodiinae
      |  `-- Scarabaeinae
      `--+--+-- Geotrupidae
         |  `--+-- Ochodaeidae
         |     `--+-- Hybosoridae
         |        `-- Ceratocanthidae
         `--+-- Glaphyridae
            `--+--+-- Pleocomidae
               |  `-- Bolboceratidae
               `--+-- Trogidae (trogids, hide beetles)
                  `--+-- Passalidae
                     `--+-- Lucanidae (kampasarviset)
                        `-- Diphyllostomatidae

Reference(s):

Grimaldi, D. A. & Engel, M. S., 2005: Evolution of the Insects.
–Cambridge University Press, New York, 2005, xv-755
Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968

–iNet: The Tree of Life Project: http://tolweb.org/

Parker, S. P. (ed.), 1982: Synopsis and classification of living organisms. Vols. 1 & 2
–McGrew-Hill Book Company

Mikko's Phylogeny Archive

Scarabaeoidea – scarabs, dung beetles, grubs, chafers, and relatives

After Parker, 1982, “The Tree of Life”, and Grimaldi & Engel, 2005

Reference(s):