From Kazlev, 2000, and Tollerton, 1989
Traditional taxonomy of Eurypterida
Alternative taxonomy after Tollerton, 1989
<==o †Eurypterida Burmeister, 1843 [Gigantostraca Haeckel, 1866; Cyrtoctenida Størmer & Waterston, 1968] (meriskorpionit; seascorpions) |--o †Stylonuroidea Diener, 1924 [Stylonauracea Diener, 1924; Drepanopteroidea Kjellesvig-Waering, 1966] | |--o †Stylonuridae Diener, 1924 [Pageidae Kjellesvig-Waering, 1966, in Waterston, 1979] | | |--o †Stylonurus Page, 1856 | | | |-- †S. powriei Page, 1856 | | | |-- †S. dolichopteroides Størmer, | | | |-- †S. logani Woodward, | | | `-- †S. macrophthalmus Laurie | | |-- †Stylonurella Kjellesvig-Waering, 1966 | | |-- †Kiaeropterus Waterston, 1979 | | |-- †Tarsopterella scoticus (Woodward, 1864) Størmer, 1951 [pro Tarsopterus Clarke & Ruedemann, 1912 (non Reinhardt & Luetken, 1862); Stylonurus scoticus Woodward, 1864] [incl.? Erieopterus brewsteri (Woodward, 1864) (juveline)] | | |-- †Melbournopterus crossotus Caster & Kjellesvig-Waering, 1953 | | `--o †Campulocephalus Eichw., 1860 | | |-- †C. oculatus (Kutorga, 1852) [Limulus oculatus Kutorga, 1852; Eidothea Scouler, 1831 (non Risso, 1826; nec Chambers, 1873] | | `-- †C. scouleri (Hibbert) | |--o †Drepanopteridae Kjellesvig-Waering 1966 | | |-- †Alkenopterus Størmer, 1974 | | |-- †Moselopterus Størmer, 1974 | | `--o †Drepanopterus Laurie, 1892 | | |-- †D. abonensis Simpson, 1951 | | |-- †D. struvei Størmer, 1976 | | |-- †D. pentlandicus Laurie, 1892 | | `-- †D. longicaudatus Clarke & Ruedemann | |?-o †Parastylonuridae Waterston, 1979 | | |?- ?†Stylonuroides Kjellesvig-Waering, 1966 | | |?-o ?†Brachyopterus Størmer, 1951 | | | |-- †B. stubblefieldi Størmer, 1951 | | | `-- †B. pentagonalis (Størmer) | | `-- †Parastylonurus Kjellesvig-Waering, 1966 | |--o †Laurieipteridae Kjellesvig-Waering, 1966 | | |-- †Laurieipterus Kjellesvig-Waering, 1966 | | |--o †Ctenopterus Clark & Ruedemann, 1912 | | | |-- †C. cestrotus (Clarke, 1907) [Eurypterus cestrotus Clarke, 1907; = ?Dolichocephala Claypole, 1883 (non MacQuart, 1823)] | | | `-- †C.? lacoana (Claypole) | | |-- †Hallipterus Kjellesvig-Waering, 1963a | | |-- †Pagea Waterston, 1962 | | `?- ?†Mazonipterus Kjellesvig-Waering, 1963b | |--o †Dolichopteridae Kjellesvig-Waering & Størmer, 1952 | | |-- †Dolichopterus macrocheirus Hall, 1859 [Eurypterus (Dolichopterus) macrocheirus Hall, 1859] | | |-- †Strobilopterus princetoni Ruedemann, 1935 | | |-- †Ruedemannipterus Kjellesvig-Waering, 1966 | | `-- †Syntomopterus Kjellesvig-Waering, 1961a | |--o †Kokomopteroidea Kjellesvig-Waering, 1966 [nom. trans. Tollerton, 1989] | | |--o †Kokomopteridae Kjellesvig-Waering, 1966 | | | |--o †Kokomopterus Kjellesvig-Waering, 1966 | | | | `-- †K. longicaudatus (Clarke & Ruedemann, 1912) | | | `-- †Lamontopterus Waterston, 1979 | | `--o †Hardieopteridae Tollerton, 1989 | | `-- †Hardieopterus Waterston, 1979 | |--o †Brachyopterelloidea Tollerton, 1989 | | `--o †Brachyopterellidae Tollerton, 1989 | | `-- †Brachypterella Kjellesvig-Waering, 1966 | |--o †Rhenopteroidea Størmer, 1951 | | `--o †Rhenopteridae Størmer, 1951 | | `-- †Rhenopterus diensti Størmer, 1936 | `--o †Mycteropoidea Cope, 1886 | |--o †Mycteropidae Cope, 1886 | | `--o †Mycterops Cope, 1886 [Glaucodes Pruvost, 1923] | | |-- †M. scabrosus (Woodward, H.) | | `-- †M. mathieui (Pruvost, 1923) [Glaucodes mathieui Pruvost, 1923] | `--o †Woodwardopteridae Kjellesvig-Waering, 1959 | |-- †Woodwardopterus Kjellesvig-Waering, 1959 | `-- †Vernopterus Waterson, 1968 `--o †Eurypteroidea Burmeister, 1843 [Eurypteracea Burmeister, 1843] |--o †Eurypteridae Burmeister, 1843 | |--o †Eurypterus De Kay, 1825 [non Mabille, 1877; nec Sharpe, 1896] [incl. Baltoeurypterus Størmer, 1973 sensu Tetlie, 2006] | | |-- †E. minor Laurie, 1898 | | |-- †E. remipes | | |-- †E. pittsfordensis | | |-- †E. fischeri Eichw. | | |-- †E. hankeni Tetlie, 2006 | | |-- †E. leopoldi Tetlie, 2006 | | |-- †E. tetragonophthalmus (Fischer, 1839) [Baltoeurypterus tetragonophthalmus] | | |-- †E. latus [Baltoeurypterus latus] | | `-- †E. remipes [Baltoeurypterus remipes] | `-- †Tylopterella boyli (Whiteaves, 1884) Størmer, 1951 [pro Tylopterus Clarke & Ruedemann, 1912 (non Capiomont, 1868; nec Leconte, 1876); Eurypterus boyli Whiteaves, 1884] |--o †Erieopteridae Tollerton, 1989 | |--o †Buffalopterus Kjellesvig-Waering & Heubusch, 1962 | | `-- †B. pustulosus [Erieopterus pustulosus] | |--o †Onychopterella Størmer, 1951 [pro Onychopterus Clarke & Ruedemann, 1912 (non Reichenbach, 1850)] | | `-- †O. kokomoensis (Miller & Gurley, 1896) [Eurypterus kokomoensis Miller & Gurley, 1896] | `--o †Erieopterus Kjellesvig-Waering, 1958 | |--o †E. microphthalmus (Hall, 1859) | | |-- †E. m. microphthalmus (Hall, 1859) | | |-- †E. m. eriensis (Whitfield, 1882) | | `-- †E. m. turgidus Stumm & Kjellesvig-Waering, 1962 | |-- †E. laticeps (Schmidt, 1883) | |-- †E. chadwicki (Clarke & Ruedemann, 1912) | `-- †E. hudsonicus (Ruedemann, 1934) `--+--o †Adelophthalmidae Tollerton, 1989 | |-- Adelophthalmus imhofi (Jordan & Meyer, 1856) Jordan & Meyer, 1854 [=? Lepidoderma imhofi (Jordan & Meyer, 1856) Reuss, 1855] | |-- †Bassipterus Kjellesvig-Waering & Leutze, 1966 | `-- †Unionopterus Chernyshev, 1948 `--+--+--o †Slimonioidea Novojilov, 1962 | | `--o Slimoniidae Novojilov, 1962 | | |-- †Slimonia acuminatus (Salter, 1856) Page, 1856 [Himantopterus acuminata Salter, 1856] | | `-- †Salteropterus abbreviatus (Salter, 1859) Kjellesvig-Waering, 1951 [Eurypterus abbreviatus Salter, 1859] | `--o †Pterygotoidea Clarke & Ruedemann, 1912 [Pterygotina Caster & Kjellesvig-Waering, 1964] | |--o †Jaekelopteridae Størmer, 1974 | | `-- †Jaekelopterus Waterston, 1964 | `--o †Pterygotidae Clarke & Ruedemann, 1912 | |-- †Truncatiramus Kjellesvig-Waering, 1961b | `--o †Pterygotus Agassiz, 1839 | |--o †(Pterygotus) Agassiz, 1839 [= Curviramus Ruedemann, 1935] | | |-- †P. (P.) rhenaniae Jaekel | | `-- †P. (P.) anglicus Agassiz | |--o †(Acutiramus) Ruedemann, 1935 | | |-- †P. (A.) buffaloensis Pohlman, 1881 | | `-- †P. (A.) bohemicus (Barrande) | `--o †(Erettopterus) Salter, 1859 [pro Himantopterus Salter, 1856 (non Westmail, 1836)] | |-- †P. (P.) bilobus (Salter, 1856?) | `-- †P. (P.) osiliensis Schmidt `--+-- †Hughmillerioidea Kjellesvig-Waering, 1951 [nom. trans. Størmer, 1974] `--+--o †Mixopteroidea Caster & Kjellesvig-Waering, 1955 | |--o †Mixopteridae Caster & Kjellesvig-Waering, 1955) | | `--o †Mixopterus Ruedemann, 1921 | | |-- †M. multispinosus (Clarke & Ruedemann, 1912) [Stylonurus (Ctenopterus) multispinosus Clarke & Ruedemann, 1912] | | `-- †M. kiaeri Størmer, | `--o †Lanarkopteridae Tollerton, 1989 | `--o †Lanarkopterus Ritchie, 1968 | `-- †L. dolichoschelus (Størmer) `--o †Megalograptoidea Caster & Kjellesvig-Waering, 1955 [nom. trans. Tollerton, 1989] `--o †Megalograptidae Caster & Kjellesvig-Waering, 1955 |-- †Echinognathus clevelandi (Walcott, 1882) Walcott, 1882 [Eurypterus clevelandi Walcott, 1882] `--o †Megalograptus Miller, 1874 |-- †M. welchi Miller, 1874 `-- †M. ohioensis Caster & Kjellesvig-Waering,
Eurypterida incertae sedis:
†Angustidontus
†Dorfopterus Kjellesvig-Waering, 1955
†Pittsfordipterus Kjellesvig-Waering & Leutze, 1966
†Willwerathia Størmer, 1969
†Glyptoscorpius perornatus Peach, 1882
†Belinuropsis wigodensis Matthew, 1909
Reference(s):
- Agnér, I. B., 199?: Eurypterida
–iNet: http://www.ettnet.se/~iba/Fosspict/pict7.htm - Braddy, S. J. & Dunlop, J. A., 2000: Early Devonian eurypterids from the Northwest Territories of Arctic Canada.
–Canadian Journal of Earth Sciences: Vol. 37, pp. 1167-1175 - Kazlev, M. A., 2000: Eurypterida – All about “Sea scorpions”.
–iNet: The Kheper Site: http://www.kheper.auz.com/gaia/biosphere/arthropods/eurypterida/index.html - Kazlev, M. A., pers comm. [originally from http://palaeo.gly.bris.ac.uk/personnel/braddy/Braddy.html]
- Stott, C. A., Tetlie, O. E., Braddy, S. J., Nowlan, G. S., Glasser, P. M. & Devereux, M. G., 2005: A new eurypterid (Chelicerata) from the Upper Ordovician of Manitoulin Island, Ontario, Canada.
–Journal of Paleontology: Vol. 79, #6, pp. 1166-1174 - Størmer, L., Petrunkevitch, A. & Hedgpeth, J. W., 1955: Arthropoda 2 – Chelicerata with sections on Pycnogonida and Palaeoisopus. xvii-181
in Moore, R. C. (ed.): Treatise on Invertebrate Paleontology – Part P
–Geological Society of America and University of Kansas Press, Lawrence, 1955 - Tollerton Jr., V. P., 1989: Morphology, taxonomy, and classification of the order Eurypterida Burmeister, 1843.
–Journal of Paleontology: Vol. 63, #5, pp. 642-657 - Tetlie, O. E., 2006: Two new Silurian species of Eurypterus (Chelicerata: Eurypterida) from Norway and Canada and the phylogeny of the genus.
–Journal of Systematic Paleontology: Vol. 4, #4, pp. 397-412 [doi:10.1017/S1477201906001921] [Abstract only]