After McKenna & Bell, 1997, de Muizon, Cifelli & Céspedes, 1997, and Szalay & Trofimov, 1996, with species from Alroy, 2002
Family -level taxonomy of non-australian metatherians.
Phylogeny of Metatheria after Rougier, Davis & Novacek, 2015
Phylogeny of Metatheria after Vullo et al., 2009
Phylogeny of Metatheria after Case, Goin & Woodburne, 2005
Phylogeny of Metatheria after Szaley & Trofimov, 1996
<==o Metatheria Huxley, 1880 [Marsupialia sensu lato] (marsupials and relatives; pussieläimet) |-- †Sinodelphys szalayi Luo, Ji, Wible & Yuan, 2003 |-- †Holoclemensia texana (Slaughter, 1968) [†Clemensia Slaughter, 1968 (non Clemensia Packard, 1864, a butterfly)] L.Cret. NA. [Holoclemensiidae Aplin & Archer, 1987; Protodelphia Archer, 1984] `--+--o †Deltatheroida Kielan-Jaworowska, 1982 sensu Ni, Li, Stidham, Li, Lu & Meng, 2016 | |-- †Pappotherium pattersoni Slaughter, 1965 sensu Bi, Jin, Li & Du, 2015 | `-- †Deltatheroida Kielan-Jaworowska, 1982 [taxonomic history] `--o Marsupialiformes Vullo, Gheerbrant, de Muizon & Néraudeau, 2009 [“Neometatheria”] |?- †Arcantiodelphys marchandi Vullo, Gheerbrant, de Muizon & Néraudeau, 2009 |?- †Aenigmadelphys archeri Cifelli & Johanson, 1994, U.Cret. NA. |?-o †Anchistodelphys Cifelli, 1990b; U.Cret. NA. | |?- ?†A. delicatus Cifelli, 1990c | `-- †A. archibaldi Cifelli, 1990b |?- †Neometatheria genera et species indet. [UA8699] L. Cret. Mad. `==o †Boreometatheria (Asiatic and North American metatheres; aasialaiset ja pohjois-amerikkalaiset pussieläimet) [a paraphyletic assemblage of non-Notometatherian metatheres] |?- †Camptomus amplus Marsh, 1889 |?- †Adinodon pattersoni Hershkovitz, 1995 [Adinodontinae Hershkovitz, 1995] `--+-- †Iqualadelphis lactea Fox, 1987 `--+--o †Asiadelphia Trofinov & Szalay, 1994 | |?- †Kokopellia juddi Cifelli, 1993; L.Cret. NA. [Kopellia lapsus calami?] | `--+-- †Asiatherium reshetovi Trofinov & Szalay, 1994 [Asiatheriidae Trofinov & Szalay, 1994] | `?-o †Marsasia Nesov, 1997 [?†Asiatherium; jr. synon. Sulestes (Deltatheroida)] | |-- †M. aenigma Nesov, 1997 | `-- †M. sp. [Averianov & Kielan-Jaworowska, 1999] `==o ?†Ameridelphia Szalay, in Archer, ed., 1982 sensu stricto? [] |?- †Dakotadens morrowi Eaton, 1993b |?-o †Iugomortiferum Cifelli, 1990b; U.Cret. NA. | |-- †I. sp. [Cifelli, 1990b] [undescribed] | `-- †I. thoringtoni Cifelli, 1990b |?-o †Protalphadon Cifelli, 1990 | |-- †P. lulli (Clemens, 1966) Cifelli, 1990 [Alphadon lulli Clemens, 1966] | `-- †P. foxi Johanson, 1996b |-- †Varalphadon cerber (Fox, 1971) Johanson, 1996 [Protalphadon cerber (Fox, 1971); Alphadon cerber Fox, 1971] |-- †Varalphadon waheapensis (Cifelli, 1990a) Johanson, 1996 [Protalphadon waheapensis Cifelli, 1990a] |-- †Varalphadon crebreforme (Cifelli, 1990b) Johanson, 1996 [Protalphadon crebreforme Cifelli, 1990b] |--o †Stagodontidae Marsh, 1889 sensu McKenna & Bell, 1997 [taxonomic history] [†Anchimetatheria Szalay, 1993, in partim] | |?- †Boreodon matutinus Lambe, 1902 (nomen dubium, Fox & Naylor, 2006) | |?-o †Pariadens Cifelli & Eaton, 1987 [treated as stagodonts by Cifelli & Eaton, 1987 and Cifelli, 2004] | | |-- †P. kirklandi Cifelli & Eaton, 1987 | | `-- †P. mckennai Cifelli, 2004 | `-- †Stagodontidae Marsh, 1889 sensu stricto [taxonomic history] [†Anchimetatheria Szalay, 1993, in partim] |--+--+-- †Bistius bondi Clemens & Lillegraven, 1986 | | `--o †Turgidodon Cifelli, 1990 [Alphadon Simpson, 1927] | | |-- †T. praesagus (Russell, 1952) Cifelli, 1990 [Alphadon praesagus Russell, 1952] | | |-- †T. rhaister (Clemens, 1966) Cifelli, 1990 [Alphadon rhaister Clemens, 1966] | | |-- †T. russelli (Fox, 1979) Cifelli, 1990 [Alphadon russelli Fox, 1979] | | |-- †T. parapraesagus (Rigby & Wolberg, 1987) Cifelli, 1990 [Alphadon parapraesagus Rigby & Wolberg, 1987] | | |-- †T. petiminis Storer, 1991 | | |-- †T. lillegraveni Cifelli, 1990 | | `-- †T. madseni Cifelli, 1990 | `--+-- †Alphadontidae Marshall, Case & Woodburne, 1990 [Alphadontia Archer, 1984; Didelphidae: Alphadontinae Marshall, Case & Woodburne, 1990 sensu McKenna & Bell, 1997] | `-- †Pediomyidae Simpson, 1927 [†Anchimetatheria Szalay, 1993, in partim] `--+-- †Jaskhadelphys minutus Marshall & de Muizon, 1988 [Jaskhadelphysidae Marshall & de Muizon, 1988 or de Muizon, 1991] |?-+?- †Thylacodon pusillus Matthew & Granger, 1921 [P. pusillus Matthew & Granger, 1921] | `?- †Simpsonitheria Szalay, in Szalay, Novacek & McKenna, eds., 1993 `--+?- †Herpetotheriidae Trouessant, 1879 [Didelphidae: Herpetotheriinae Trouessant, 1879 sensu McKenna & Bell, 1997] [paraphyletic?] `-- Notometatheria (South American and Australian metatheres, incl. Marsupialia; etelä-amerikkalaiset ja australialaiset pussieläimet)
Metatheria (Marsupialia) incertae sedis:
†Garatherium mahboubii Crochet, 1984; E.Eoc. NAf.
†Kasserinotherium tunisiense Crochet, 1989; E.Eoc. NAf.
†Ghamidtherium dimaiensis Sánches-Villagra et al., 2007; L.Eoc. NEAf.
Alternative phylogenies of Metatheria:
After Rougier, Davis & Novacek, 2015<==o Metatheria |-- †Holoclemensia `--+--+-- †Pappotherium | `-- †Deltatheroida `--+--+-- †Asiatherium | `--+-- †Aenigmadelphys 1 | `--+---+--+-- †Kokopellia | | | `-- †Iugomortiferum | | `--+-- †Turgidodon | | `--+-- †Albertatherium | | `-- †Alphadon | `--+?- †Anchistodelphys | `--+--+-- †Pediomys | | `-- †Glasbius | `--+?- †Aenigmadelphys 2 | `--+-- †Pariadens | `--+-- †Eodelphis | `-- †Didelphodon `--+-- Gurlin Tsav Skull `--+-- †Borhyaenidae `--+-- †Mayulestes `--+-- †Jaskhadelphys `--+-- †Pucadelphys `--+-- †Andinodelphys `--+-- Didelphidae `--+-- Microbiotheria `-- Dasyuromorphia Reference(s):
|
After Vullo, Gheerbrant, de Muizon & Néraudeau, 2009<==o Metatheria |-- †Sinodelphys `--+-- †Deltatheroida `--o Marsupialiformes |-- †Arcantiodelphys |-- †Dacotadens `--+-- †Kokopellia `--+-- †Asiatherium `--+-- Didelphis `--+-- †Iugomortiferum |-- †Sinbadelphys |-- †Adelodelphys `-- †Pariadens |
After Case, Goin & Woodburne, 2005<==o Metatheria |?- †Deltatheroida |?-o †Asiadelphia | |?- †Kokopellia | `--+-- †Asiatherium | `-- †Marsasia `--o Marsupialia |--o †“Alphadelphia” | `--o †Peradectia | |-- †Peradectidae | |-- †Pediomyidae | |-- †Stagodontidae | `-- †Caroloameghiniidae |--o Ameridelphia | |?- †Andinodelphys | |--o Didelphimorphia | | `--o Didelphoidea | | |-- †Pucadelphidae | | |-- Didelphidae | | |-- Caluromyidae | | |-- †Herpetotheriidae | | |-- †Derorhynchidae | | `-- †Protodidelphidae | |-- Paucituberculata | `--o †Sparassodonta | `--o †Borhyaenoidea | |-- †Mayulestidae | |-- †Hondadelphidae | |-- †Hathliacynidae | |-- †Borhyaenidae | |-- †Proborhyaenidae | `-- †Thylacosmilidae `--o Australidelphia |-- Microbiotheria |-- Peramelia |-- Dasyuromorphia |-- Notorycteromorpha `-- Diprotodontia |
After Szalay & Trofimov, 1996<==o Metatheria |-- †Deltatheroida `--+--o †Asiadelphia | |?- †Kokopellia | `--+-- †Marsasia | `-- †Asiatherium `==o †Archimetatheria |-- †Stagodontidae |-- †Pediomyidae |--+-- †Sudameridelphia | `--+-- †Simpsonitheria | `-- Paucituberculata `--+-- Didelphimorphia `--+--+-- Microbiotheria | `-- Dasyuromorphia `--o Syndactyli |--+?- Peramelia | `?- Notorycteromorpha `--+?-o †Yalkaparidontia | `-- †Yalkaperidon `-- Diprotodontia |
Reference(s):
- Averianov, A. O., 2000: Mammals from the Mesozoic of Kirgizstan, Uzbekistan, Kazakhstan and Tadzhikistan. 627-652
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000, xxxix-696 - Bi, S., Jin, X., Li, S. & Du, T., 2015: A new Cretaceous Metatherian mammal from Henan, China.
–PeerJ: Vol. 3, pp. e896 [doi: 10.7717/peerj.896] - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Cifelli, R. L., 1990: Cretaceous mammals of Southern Utah. II. Marsupials and marsupial-like mammals from the Wahweap Formation (Early Campanian).
–Journal of Vertebrate Paleontology: Vol. 10, #3, pp. 320-331 - Cifelli, R. L., 1990: Cretaceous mammals of Southern Utah. III. Therian mammals from the Turonian (Early Late Cretaceous).
–Journal of Vertebrate Paleontology: Vol. 10, #3, pp. 332-345 - Cifelli, R. L. & Johanson, Z., 1994: New marsupial from the Upper Cretaceous of Utah.
–Journal of Vertebrate Paleontology: Vol. 14, #2, pp. 292-295 - Gunnell, G. F., 2010: 6. Marsupialia (77-87).
in Werdelin, L. & Sanders, W. J., (eds.) 2010: Cenozoic mammals of Africa. –University of California Press, Berkeley, 2010, xxi-986 - Johanson, Z., 1993: A revision of the Late Cretaceous (Campanian) marsupial Iqualadelphis lactea Fox, 1987.
–Journal of Vertebrate Paleontology: Vol. 13, #3, pp. 373-377 - Kielan-Jaworowska, Z., Novacek, M. J., Trofimov, B. A. & Dashzeveg, D., 2000: Mammals from the Mesozoic of Mongolia. 573-626
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000, xxxix-696 - Kielan-Jaworowska, Z. & Cifelli, R.L. 2001: Primitive boreosphenidan mammal (?Deltatheroida) from the Early Cretaceous of Oklahoma.
–Acta Palaeontologica Polonica 46, #3, pp. 377-391.
[from iNet abstract: http://www.paleo.pan.pl/acta/acta46-3.htm] - Krause, D. W., 2001: Fossil molar from a Madagascan marsupial.
–Nature: Vol. 412, 2 August, pp. 487-498 - Luo, Z.-X., Ji, Q., Wible, J. R. & Yuan, C.-X., 2003: An Early Cretaceous Tribosphenic Mammal and Metatherian Evolution.
–Science: Vol. 302, #5652, pp. 1934-1940 [doi: 10.1126/science.1090718] - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - de Muizon, C., Cifelli, R. L. & Céspedes Paz, R., 1997: The origin of the dog-like borhyaenoid marsupials of South America.
–Nature: Vol. 389, 2 October, pp. 486-489 - Rougier, G. W., Wible, J. R. & Novacek, M. J., 1998: Implications of Deltatheridium speciments for early marsupial history.
–Nature: Vol. 396, 3 December, pp. 459-463 - Vullo, R., Gheerbrant, E., de Muizon, C. & Néraudeau, D., 2009: The oldest modern therian mammal from Europe and its bearing on stem marsupial paleobiogeography.
–Proceedings of the National Academy of Science: Vol. 106, #47, pp. 19910-19915 [doi: 10.1073/pnas.0902940106] [http://www.pnas.org/cgi/doi/10.1073/pnas.0902940106]