Mikko's Phylogeny Archive

Deltatheroida – deltatherids

After Bi, Jin, Li & Du, 2015, and Rougier, Davis & Novacek, 2015, with genera and species from McKenna & Bell, 1997, Rougier, Wible & Novacek, 1998, Averianov, 2000, Kielan-Jaworowska, Novacek, Trofimov & Dashzeveg, 2000, Kielan-Jaworowska & Cifelli, 2001, and Rougier, Davis & Novacek, 2015

<==o †Deltatheroida Kielan-Jaworowska, 1982 [taxonomic history]
   `--o †Deltatheridiidae Gregory & Simpson, 1926 [taxonomic history]
      |--o †Sulestes Nesov, 1985; U.Cret. Eas. [Deltatherus Nessov, 1997; Marsasia Nesov, 1997] [Sulestinae Nessov, 1985b]
      |  |?- ?†S. sp. [Marsasia sp. Averianov & Kielan-Jaworowska, 1999; = Paranyctoides aralensis Nessov, 1993 (Archibald & Averianov, 2005)]
      |  |-- †S. sp. [Central Asia]
      |  `-- †S. karakshi Nesov, 1985 [Deltatheroides kizylkumensis Nessov, 1993; Deltatherus kizylkumensis (Nessov) Nessov, 1997; Marsasia aenigma Nesov, 1997]
      `--+--o †Oklatheridium Davis, Cifelli & Kielan-Jaworowska, 2008
         |  |-- †O. szalayi Davis, Cifelli & Kielan-Jaworowska, 2008
         |  `-- †O. minax Davis & Cifelli, 2011
         `--+-- †Tsagandelta dashzevegi Rougier, Davis & Novacek, 2015
            `--+-- †Lotheridium mengi Bi, Jin, Li & Du, 2015
               `--+--+-- †Atokatheridium boreni Kielan-Jaworowska & Cifelli, 2001
                  |  `--+-- †Nanocuris improvida Fox, Scott & Bryant, 2007
                  |     `-- †Gurbanodelta kara Ni, Li, Stidham, Li, Lu & Meng, 2016
                  `--+--o †Deltatheridium Gregory & Simpson, 1926; U.Cret. EAs.
                     |  |-- †D. pretrituberculare Gregory & Simpson, 1926 [†D. p. pretrituberculare and †D. p. tardum]
                     |  `-- †D. nessovi Averianov, 1997
                     `--o †Deltatheroides Gregory & Simpson, 1926: U.Cret. EAs. ?NA. [Deltatheroididae Kielan-Jaworowska & Nesov, 1990]
                        |-- †D. cretacicus Gregory & Simpson, 1926
                        `-- †D. sp.

Deltatheroida incertae sedis:

    †Khuduklestes bohlini Nesov, Sigogneau-Russell & Russell, 1994; Cret. As.
    †Beleutinus orlovi Bazhanov, 1972 [is a zalambdalestid?]
    †Falepterus barwini Clemens & Lillegraven, 1986 [see basal Theria]
    †“Deltatheroides” kizylkumensis Nesov, 1993
    †Oxlestes grandis Nesov, 1982; U.Cret. As. [a zhelestid? ]

<==o †Deltatheroida
   |--  †Pappotherium
   `--o †Deltatheridiidae
      |-- †Oklatheridium
      |-- †Sulestes
      `--+-- †Tsagandelta
         `--+-- †Lotheridium
            `--+--+-- †Deltatheridium
               |  `-- †Deltatheroides
               `--+-- †Atokatheridium
                  `--+-- †Nanocuris
                     `-- †Gurbanodelta

<==o †Deltatheroida
   `--o †Deltatheridiidae
      |-- †Sulestes
      `--+-- †Oklatheridium
         `--+-- †Tsagandelta
            `--+-- †Lotheridium
               `--+--+-- †Atokatheridium
                  |  `-- †Nanocuris
                  `--+-- †Deltatheridium
                     `-- †Deltatheroides

<==o †Deltatheroida
   `--o †Deltatheridiidae
      |--+-- †Sulestes
      |  `-- †Oklatheridium
      `--+-- †Tsagandelta
         `--+--+-- †Atokatheridium
            |  `-- †Nanocuris
            `--+-- †Deltatheridium
               `-- †Deltatheroides

Reference(s):

• Averianov, A. O., 2000: Mammals from the Mesozoic of Kirgizstan, Uzbekistan, Kazakhstan and Tadzhikistan. 627-652
  in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
  –Cambridge University Press, Cambridge, 2000, xxxix-696
• Bi, S., Jin, X., Li, S. & Du, T., 2015: A new Cretaceous Metatherian mammal from Henan, China.
  –PeerJ: Vol. 3, pp. e896 [doi: 10.7717/peerj.896]
• Carroll, R. L., 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698
• Carroll, R. L., 1988: Appendix. 594-648.
  in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698
• Cifelli, R. L., 1990: Cretaceous mammals of Southern Utah. II. Marsupials and marsupial-like mammals from the Wahweap Formation (Early Campanian).
  –Journal of Vertebrate Paleontology: Vol. 10, #3, pp. 320-331
• Cifelli, R. L., 1990: Cretaceous mammals of Southern Utah. III. Therian mammals from the Turonian (Early Late Cretaceous).
  –Journal of Vertebrate Paleontology: Vol. 10, #3, pp. 332-345
• Cifelli, R. L. & Johanson, Z., 1994: New marsupial from the Upper Cretaceous of Utah.
  –Journal of Vertebrate Paleontology: Vol. 14, #2, pp. 292-295
• Fox, R. C., Scott, C. S. & Bryant, H. N., 2007: A new, unusual therian mammal from the Upper Cretaceous of Saskatchewan, Canada.
  –Cretaceous Research: Vol. 28, #5, pp. 821-829 [doi: 10.1016/j.cretres.2006.12.005]
• Johanson, Z., 1993: A revision of the Late Cretaceous (Campanian) marsupial Iqualadelphis lactea Fox, 1987.
  –Journal of Vertebrate Paleontology: Vol. 13, #3, pp. 373-377
• Kielan-Jaworowska, Z., Novacek, M. J., Trofimov, B. A. & Dashzeveg, D., 2000: Mammals from the Mesozoic of Mongolia. 573-626
  in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
  –Cambridge University Press, Cambridge, 2000, xxxix-696
• Kielan-Jaworowska, Z. & Cifelli, R.L. 2001: Primitive boreosphenidan mammal (?Deltatheroida) from the Early Cretaceous of Oklahoma.
  –Acta Palaeontologica Polonica 46, 3, 377-391.
• Krause, D. W., 2001: Fossil molar from a Madagascan marsupial
  –Nature: Vol. 412, 2 August, pp. 487-498
• McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
  –Columbia University Press, New York, 1997, xii-631
• de Muizon, C., Cifelli, R. L. & Céspedes Paz, R., 1997: The origin of the dog-like borhyaenoid marsupials of South America.
  –Nature: Vol. 389, 2 October, pp. 486-489
• Ni, X., Li, Q., Stidham, T. A., Li, L., Lu, X. & Meng, J., 2016: A late Paleocene probable metatherian (?deltatheroidan) survivor of the Cretaceous mass extinction.
  –Scientific Reports: Vol. 6, art. 38547 [doi: 10.1038/srep38547]
• Rougier, G. W., Davis, B. M. & Novacek, M. J., 2015: A deltatheroidan mammal from the Upper Cretaceous Baynshiree Formation, eastern Mongolia.
  –Cretaceous Research: Vol. 52, pp. 167-177 [doi: 10.1016/j.cretres.2014.09.009]
• Rougier, G. W., Wible, J. R. & Novacek, M. J., 1998: Implications of Deltatheridium speciments for early marsupial history.
  –Nature: Vol. 396, 3 December, pp. 459-463
• Wilson, G. P. & Riedel, J. A., 2010: New Specimen Reveals Deltatheroidan Affinities of the North American Late Cretaceous Mammal Nanocuris.
  –Journal of Vertebrate Paleontology: Vol. 30, #3, pp. 872-884 [doi: 10.1080/02724631003762948]

Special thanks to:

Jos H. M. Dols, for gathering additional information and species.