After McKenna & Bell, 1997, with Recent species from Nowak, 1991
<==o Pteropodini Gray, 1821 |--o Rousettina Andersen, 1912 | |--o Eidolon | | |-- E. helvum (keltahedelmälepakko) | | `-- ?E. dupreanum | `--+--o Rousettus | | |-- R. aegyptiacus (Geoffroy, 1810) | | |-- R. obliviosus | | |-- R. amplexicaudatus | | |-- R. spinalatus | | |-- R. leschenaulti | | |-- R. seminudus | | |-- R. lanosus | | |-- R. celebensis (celebesinkoirankuonolepakko) | | |-- R. angolensis | | `-- ?R. madagascariensis | `--+-- Boneia bidens | `--o Myonycteris | |-- M. torquata | |-- M. relicta | `-- M. brachycephala `--+--o Dobsoniina Andersen, 1912 | |-- Aproteles bulmerae | `--o Dobsonia (paljasselkälepakot) | |-- D. peronii | |-- D. viridis | |-- D. crenulata[ssp.=>D. viridis?] | |-- D. beauforti | |-- D. praedatrix | |-- D. inermis | |-- D. moluccensis (isopaljasselkälepakko) | |-- D. magna [ssp.=>D. moluccensis?] | |-- D. emersa | |-- D. anderseni [ssp.=>D. moluccensis?] | |-- D. pannietensis [ssp.=>D. moluccensis?] | |-- D. remota [ssp.=>D. moluccensis?] | |-- †D. chapmani ?(filippiinienpaljasselkälepakko) | |-- D. exoleta (celebesinpaljasselkälepakko) | `-- D. minor `--o Pteropodina Gray, 1821 |-- Pteropus `--+--o Acerodon | |-- A. celebensis (celebesinhedelmälepakko) | |-- A. mackloti | |-- A. humilis | |-- A. jubatus | |-- A. leucotis | `-- †A. lucifer `--+--o Pteralopex | |-- P. anceps | |-- P. atrata | `-- P. acrodonta `--+-- Styloctenium wallacei (viirunaamahedelmälepakko) `-- Neopteryx frosti (pienihammashedelmälepakko)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629