After Bergmans, 1997 [from Giannini & Simmons, 2003]
<==o Pteropodinae Gray, 1821 ()? |--o Pteropodina Gray, 1821 ()? | |-- Pteropus ()? | |--o Acerodon ()? | | |-- A. celebensis (celebesinhedelmälepakko) | | |-- A. mackloti ()? | | |-- A. humilis ()? | | |-- A. jubatus ()? | | |-- A. leucotis ()? | | `-- †A. lucifer ()? | |--o Pteralopex ()? | | |-- P. anceps ()? | | |-- P. atrata ()? | | `-- P. acrodonta ()? | |-- Styloctenium wallacei (viirunaamahedelmälepakko) | `-- Neopteryx frosti (pienihammashedelmälepakko) |--o Macroglossini Gray, 1821 | |--o Macroglossus (kielilepakot) | | |-- M. sobrinus (isokielilepakko) | | |-- M. minimus [incl. M. lagochilus] (pikkukielilepakko) | | `-- M. fructivorus ()? | `--o Syconycteris (kukkalepakot) | |-- S. australis (etelänkukkalepakko) | |-- S. hobbit ()? | `-- S. carolinae ()? |--o Notopterini Andersen, 1912 ()? | |-- Nesonycteris [jr. synon. to Melonycteris??] | |-- Notopteris macdonaldi (pitkähäntähedelmälepakko) | `--o Melonycteris ()? | |-- M. melanops (mustavatsahedelmälepakko) | |-- M. aurantius ()? | `-- M. woodfordi ()? |--o Rousettini ()? | |--o Eidolon ()? | | |-- E. helvum (keltahedelmälepakko) | | `?- E. dupreanum ()? [ssp. => E. helvum] [Madagascar] | |--o Eonycteris (aamulepakot) | | |-- E. spelaea (aamulepakko) | | |-- E. robusta ()? | | |-- E. major ()? | | `-- E. rosenbergi ()? | `--o Rousettus [Boneia; Stenonycteris] ()? | |-- R. aegyptiacus (Geoffroy, 1810) ()? | |-- R. bidens [Boneia bidens] ()? | |-- R. obliviosus ()? | |-- R. amplexicaudatus ()? | |-- R. spinalatus ()? | |-- R. leschenaulti ()? | |-- R. seminudus ()? | |-- R. lanosus ()? | |-- R. celebensis (celebesinkoirankuonolepakko) | |-- R. angolensis ()? | `-- ?R. madagascariensis ()? `--o Dobsoniini Andersen, 1912 ()? |-- Aproteles bulmerae ()? `--o Dobsonia (paljasselkälepakot) |-- D. peronii ()? |-- D. viridis ()? |-- D. crenulata ()?[ssp.=>D. viridis?] |-- D. beauforti ()? |-- D. praedatrix ()? |-- D. inermis ()? |-- D. moluccensis (isopaljasselkälepakko) |-- D. magna [ssp.=>D. moluccensis?] ()? |-- D. emersa ()? |-- D. anderseni [ssp.=>D. moluccensis?] ()? |-- D. pannietensis [ssp.=>D. moluccensis?] ()? |-- D. remota [ssp.=>D. moluccensis?] ()? |-- †D. chapmani ?(filippiinienpaljasselkälepakko) |-- D. exoleta (celebesinpaljasselkälepakko) `-- D. minor ()?
Reference(s):
- Giannini, N. P. & Simmons, N. B., 2003: A phylogeny of megachiropteran bats (Mammalia: Chiroptera: Pteropodidae) based on direct optimization analysis of one nuclear and four mitochondrial genes.
–Cladistics: Vol. 19, pp. 496-511 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629