After Bergman, 1997 [from Giannini & Simmons, 2003]
<==o Nyctimeninae Miller, 1907 [Pteropodidae: Pteropodinae: Cynopterini: Nyctimenina sensu McKenna & Bell, 1997]
|-- Paranyctimene raptor ()?
`--o Nyctimene (putkikuonolepakot)
|-- N. minutus (pikkuputkikuonolepakko)
|-- N. albiventer ()?
|-- N. draconilla ()?
|-- N. robinsoni ()?
|-- N. cephalotes ()?
|-- N. vizcaccia ()?
|-- N. masalai ()?
|-- N. rabori ()?
|-- N. malaitensis ()?
|-- N. major (isoputkikuonolepakko)
|-- N. sanctacrucis ()?
|-- N. cyclotis ()?
|-- N. aello ()?
|-- N. celaeno ()?
`-- ?N. certans ()?
Reference(s):
- Giannini, N. P. & Simmons, N. B., 2003: A phylogeny of megachiropteran bats (Mammalia: Chiroptera: Pteropodidae) based on direct optimization analysis of one nuclear and four mitochondrial genes.
–Cladistics: Vol. 19, pp. 496-511
- Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii
- Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629