After Giannini & Simmons, 2003, and Giannini, Cunha Almeida, Simmons & DeSalle, 2006
<==o Megachiroptera (suurlepakot)
`--o Pteropodidae (hedelmälepakot)
|-- †Archaeopteropus transiens Meschinelli, 1903 [Archaeopteropodinae]
`--+-- †Propotto leakeyi Simpson, 1967 [Propottinae]
`--o Pteropodidae sensu Giannini & Simmons, 2003 [recent Pteropodids]
|?- Plerotes [Plerotini Bergmans, 1997] [not treated by Giannini & Simmons, 2003]
|?-o Scotonycterini Bergmans, 1997 [not treated by Giannini & Simmons, 2003]
| |-- Scotonycteris
| `-- Casinycteris
|-- Melanonycteris
`--+--o Nyctimeninae Miller, 1907
| |-- Paranyctimene
| `-- Nyctimene
`--+--+-- Eiolodon
| `-- Pteralopex
`--+--+-- Acerodon
| `-- Pteropus
`--+--+-- Notopteris
| `--+-- Harpyionycterinae Miller, 1907
| `--+--+-- Macroglossus
| | `-- Syconycteris
| `--o Cynopterinae Andersen, 1912
| |?- Ptenochirus [not treated by Giannini & Simmons, 2003]
| |?- Dyacopterus [not treated by Giannini & Simmons, 2003]
| |?- Sphaerias [not treated by Giannini & Simmons, 2003]
| |?- Penthetor [not treated by Giannini & Simmons, 2003]
| |?- Latidens [not treated by Giannini & Simmons, 2003]
| |?- Alionycteris [not treated by Giannini & Simmons, 2003]
| |?- Otopteropus [not treated by Giannini & Simmons, 2003]
| |?- Haplonycteris [not treated by Giannini & Simmons, 2003]
| |--+-- Cynopterus
| | `--+-- Megaerops
| | `-- Ptenochirus
| `--+-- Thoopterus
| |-- Chironax
| |-- Aethalops
| `-- Balionycteris
`--+--+-- Eonycteris
| `--+-- Stenonycteris
| `-- Rousettus
`--+--o Epomophorini Gray, 1821
| |?- Hypsignathus [not treated by Giannini & Simmons, 2003]
| |?- Nanonycteris [not treated by Giannini & Simmons, 2003]
| |-- Epomops
| `--+-- Epomophorus
| `-- Micropteropus
`--o Myonycterini Lawrence & Novick, 1963
|-- Megaloglossus
`--+-- Lissonycteris
`-- Myonycteris
Reference(s):
- Butler, P. C., 1978: Insectivora and Chiroptera. 56-68
in Maglio, V. J. & Cooke, H. B. S., (eds.) 1978: Evolution of African Mammals, 1.
–Harvard University Press, Cambridge, Massachusetts, London, England, 1978, xiv-641
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698
- Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698
- Giannini, N. P., Cunha Almeida, F., Simmons, N. B. & DeSalle, R., 2006: Phylogenetic relationships of the enigmatic harpy fruit bat, Harpyionycteris (Mammalia: Chiroptera: Pteropodidae).
–American Museum Novitates: #3533, pp. 1-12
- Giannini, N. P. & Simmons, N. B., 2003: A phylogeny of megachiropteran bats (Mammalia: Chiroptera: Pteropodidae) based on direct optimization analysis of one nuclear and four mitochondrial genes.
–Cladistics: Vol. 19, pp. 496-511
- McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631
- Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii
- Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629