After Wyss & Flynn, 1993, McKenna & Bell, 1997, Alroy, 2002, and Wesley-Hunt & Flynn, 2005
<==o †Viverravidae Wortman & Matthew, 1899 [incl. Quercygalidae Kretzoi, 1945; Didymictidae Flynn & Galiano, 1982] [Viverraroidea Wortman & Matthew 1899] |?- †Pappictidops Chiu & Li, 1977; E.Pal. EAs. |--o †Viverravinae Wortman & Matthew, 1899 [Viverravidae Wortman & Matthew, 1899; Viverravinae Matthew, 1909; Viverravini Kalandadze & Rautian, 1992; Viverravina Kalandadze & Rautian, 1992] | |-- †Pristinictis connata Fox & Youzwyshyn, 1994 | |--o †Simpsonictis MacIntyre, 1962; E.-L.Pal. NAm. | | |-- †S. jaynanneae Rigby,jr., 1980 | | |-- †S. pegus Gingerich & Winkler, 1985 | | `-- †S. tenuis (Simpson, 1935) [Didymictis tenuis Simpson, 1935; Protictis tenuis (Simpson, 1935)] | |--o †Viverravus Marsh, 1872 [Triacodon Marsh, 1871] L.Pal.-M-Eoc. NAm., ?E.Eoc.-L.Eoc. Eu. | | |-- †V. acutus Matthew & Granger, 1915 | | |-- †V. gracilis Marsh, 1872 | | |-- †V. minutus Wortman, 1901 | | |-- †V. politus Matthew & Granger, 1915 | | |-- †V. rosei Polly, 1997 | | `-- †V. sicarius Matthew, 1909b | |?- †Quergygale angustidens (Filhol, 1872) Kretzoi, 1945 [Viverra angustidens Filhol, 1872; Humbertia Beaumont, 1966] [incl. Viverra hastingsae Davies, 1884; “Cynodon” helveticus (Rütimeyer, 1862)] [a non-carnivoran carnivoromorphan (Wesley-Hunt & Werdelin, 2005)] | |-- †Simamphicyon helveticus (Pictet & Humbert 1869) Viret, 1942; M.Eoc.-?L.Eoc. Eu. | |?-o †Tapocyon Stock, 1934 [taxonomic history] [a non-carnivoran carnivoromorphan (Wesley-Hunt & Werdelin, 2005)] | | |-- †T. dawsonae Wesley & Flynn, 2003 [taxonomic history] | | `-- †T. robustus (Peterson, 1919) [taxonomic history] | `?- †Plesiomiacis progressus (Stock, 1935) Simpson, 1945 [Viverravus (Plesiomiacis) progressus Stock, 1935] [synon. of Procynodictis Wortman & Matthew 1899] [a non-carnivoran carnivoromorphan (Wesley-Hunt & Werdelin, 2005)] |--o †Ictidopappinae Van Valeb, 1969 [Ictidopappini Van Valen, 1969; Ictidopappinae McKenna & Bell, 1997] | `-- †Ictidopappus mustelinus Simpson, 1935; E.Pal. NAm. `--o †Didymictinae Flynn & Galiano, 1982 [Didymictida Flynn & Galiano, 1982; Didymictidae Flynn & Galiano, 1982; Didymictinae McKenna & Bell, 1997] |-- †Raphictis gausion Gingerich & Winkler, 1985 |--o †Didymictis Cope, 1875 | |-- †D. altidens Cope, 1880 | |-- †D. leptomylus Cope, 1880 | |-- †D. protenus (Cope, 1874) [Limnocyon protenus Cope, 1874; Didymictis protenus Cope, 1875; Didymictis curtidens Cope, 1882; Didymictis protenus Matthew, 1909; 1915; 1937; Polly, 1997; Gunnell, 1998] | |-- †D. proteus Simpson, 1937 [Didymictis protenus proteus Simpson, 1937; Didymictis dellensis Dorr, 1952; Protictis dellensis Gingerich & Winkler, 1985; Didymictis proteus Polly, 1997] | `-- †D. vancleveae Robinson, 1966 |--o †Bryanictis MacIntyre, 1966 [Bryanictis MacIntyre, 1966Protictis Matthew, 1937; Mckenna & Bell, 1997] | |-- †B. microlestes (Simpson, 1935) [Didymictis microlestes Simpson, 1935] | `-- †B. paulus Meehan & Wilson, 2002 `--o †Protictis Matthew, 1937 [Didymictis (Protictis) Matthew, 1937; Protictis MacIntyre, 1966; Protictoides Flynn & Galiano, 1982; Intyrictis Gingerich & Winkler, 1985] |-- (Protictoides) aprophatos Flynn & Galiano, 1982 |-- †P. paralus Holtzman, 1978 |-- †P. simpsoni Meehan & Wilson, 2002 |-- †P. minor Meehan & Wilson, 2002 `--+-- †P. haydenianus (Cope, 1882) [Didymictis haydenianus Cope, 1882; Didymictis primus Cope, 1884] [type] `--+-- †P. agastor `-- †P. dellensis (Dorr, 1952) Gingerich & Winkler, 1985
Reference(s):
- Alroy, J., 2002: North American fossil mammal systematics database.
–iNet: North American fossil mammal systematics database: http://www.nceas.ucsb.edu/~alroy/nafmsd.html - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Gebo, D. L. & Rose, K. D., 1993: Skeletal morphology and locomotor adaptation in Prolimnocyon atavus, an Early Eocene hyaenodontid creodont.
–Journal of Vertebrate Paleontology: Vol. 13, #1, pp. 125-144 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Polly, P. D., Wesley-Hunt, G. D., Heinrich, R. E., Davis, G. & Houde, P., 2006: Earliest known carnivoran auditory bulla and support for a recent origin of crown-group Carnivora (Eutheria, Mammalia).
–Palaeontology: Vol. 49, #5, pp. 1019-1027 - Wesley-Hunt, G. D. & Flynn, J. J., 2005: Phylogeny of the Carnivora: basal relationships among the Carnivoramorphans, and assessment of the position of ‘Miacoidea’ relative to Carnivora.
–Journal of Systematic Palaeontology: Vol. 3, #1, pp. 1-28 - Wyss, A. R. & Flynn, J. J., 1993: A phylogenetic analysis and definition of the Carnivora. 32-52.
in Szalay, F. S., Novacek, M. J. & McKenna, M. C., (eds.) 1993: Mammal Phylogeny: Placentals.
–Springer-Verlag, New York, Berlin, Heidelberg, London, Paris, Tokio, Hong Kong, Barcelona, Budapest, 1993