After McKenna & Bell, 1997
<==o Scandentia Wagner, 1855 (puupäästäiset) |?- †Nycticonodon caparti Eoc. Eu. `--o Tupaiidae Gray, 1825 (tree shrews; puupäästäiset) |-- Ptilocercus lowii Gray, 1848 [Ptilocercinae Lyon, 1913] (feather-tail tree shrew, pen-tailed treeshrew; sulkahäntätupaija) `--o Tupaiinae Gray, 1825 (true tree shrews; aitotupaijat) |-- †Eodendrogale parvum Yongsheng, 1988 [non? Tong, 1988] |--o †Prodendrogale Qiu, 1986 | |-- †P. yunnanica Qiu, 1986 | `-- †P. engesseri Ni & Qiu, 2012 |--o Tupaia Raffles, 1821 | |?- †T. miocenica Mein & Ginsburg, 1997 | |?- T. chrysogaster Miller, 1903 (Golden-bellied Treeshrew, Mentawai Treeshrew; kultavatsatupaija) | |?- T. moellendorffi Matschie, 1898 (Calamian Treeshrew | |--o (Tupaia) Raffles, 1821 | | |-- T. (T.) glis (Diard & Duvaucel, 1820) [Sorex Glis Diard & Duvaucel, 1820] (Common treeshrew; tupaii; tupaija) | | |?- T. (T.) belangeri Wagner, 1841 [ssp. T. glis] (Northern Treeshrew; belangeritupaija) | | |?- T. (T.) chinensis [ssp. T. glis] (kiinantupaija) | | |?- T. (T.) longipes Thomas, 1893 [ssp. T. glis] (Long-footed Treeshrew; pitkäjalkatupaija) | | |-- T. (T.) nicobarica Zelebor, 1869 (Nicobar Treeshrew; nikobaarientupaija) | | |-- T. (T.) splendidula Gray, 1865 (Ruddy Treeshrew; punahäntätupaija) | | |-- T. (T.) javanica Horsfield, 1822 (Horsfield's Treeshrew; indonesiantupaija) | | |-- T. (T.) gracilis Thomas, 1893 (Slender Treeshrew; hoikkatupaija) | | |-- T. (T.) picta Thomas, 1892 (Painted Treeshrew; tummaselkätupaija) | | `-- T. (T.) dorsalis Schlegel, 1857 (Striped Treeshrew; juovatupaija) | `--o (Lyonogale) Conisbee, 1953 | |-- T. (L.) montana Thomas, 1892 (Mountain Treeshrew; vuoritupaija) | |-- T. (L.) minor Günther, 1876 (Pygmy Treeshrew; kääpiötupaija) | |-- T. (L.) tana Raffles, 1821 (Large Treeshrew; maatupaija) | `-- T. (L.) palawanensis Thomas, 1894 (Palawan Treeshrew; palawanintupaija) `--+-- Anathana ellioti (Waterhouse, 1850) Lyon, 1913 (Madras Treeshrew, Indian Treeshrew; intiantupaija) `--+-- Urogale everetti (Thomas, 1892) Mearns, 1905 (Mindanao Treeshrew, Philippine treeshrew; filippiinientupaija) `--o Dendrogale Gray, 1848 (Smooth-tailed Treeshrews; pehmeähäntätupaijat) |-- D. murina (Schlegel & Muller, 1843) (Northern Smooth-tailed Treeshrew; pohjanpehmeähäntätupaija) `-- D. melanura (Thomas, 1892) (Bornean Smooth-tailed Tree Shrew; etelänpehmeähäntätupaija)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
 –W. H. Freeman and company, New York, 1988, 698.
- Carroll, R. L., 1988: Appendix. 594-648.
 in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
 –W. H. Freeman and company, New York, 1988, 698.
- Mein P. & Ginsburg L., 1997: Les mammifères du gisement miocène inférieur de Li Mae Long, Thaïlande : systématique, biostratigraphie et paléoenvironnement.
 –Geodiversitas: Vol. 19, #4, pp. 783-844
- McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
 –Columbia University Press, New York, 1997, xii-631
- Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
 –The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii
- Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
 –The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629
