After McKenna & Bell, 1997
<==o Scandentia Wagner, 1855 (puupäästäiset)
|?- †Nycticonodon caparti Eoc. Eu.
`--o Tupaiidae Gray, 1825 (tree shrews; puupäästäiset)
|-- Ptilocercus lowii Gray, 1848 [Ptilocercinae Lyon, 1913] (feather-tail tree shrew, pen-tailed treeshrew; sulkahäntätupaija)
`--o Tupaiinae Gray, 1825 (true tree shrews; aitotupaijat)
|-- †Eodendrogale parvum Yongsheng, 1988 [non? Tong, 1988]
|--o †Prodendrogale Qiu, 1986
| |-- †P. yunnanica Qiu, 1986
| `-- †P. engesseri Ni & Qiu, 2012
|--o Tupaia Raffles, 1821
| |?- †T. miocenica Mein & Ginsburg, 1997
| |?- T. chrysogaster Miller, 1903 (Golden-bellied Treeshrew, Mentawai Treeshrew; kultavatsatupaija)
| |?- T. moellendorffi Matschie, 1898 (Calamian Treeshrew
| |--o (Tupaia) Raffles, 1821
| | |-- T. (T.) glis (Diard & Duvaucel, 1820) [Sorex Glis Diard & Duvaucel, 1820] (Common treeshrew; tupaii; tupaija)
| | |?- T. (T.) belangeri Wagner, 1841 [ssp. T. glis] (Northern Treeshrew; belangeritupaija)
| | |?- T. (T.) chinensis [ssp. T. glis] (kiinantupaija)
| | |?- T. (T.) longipes Thomas, 1893 [ssp. T. glis] (Long-footed Treeshrew; pitkäjalkatupaija)
| | |-- T. (T.) nicobarica Zelebor, 1869 (Nicobar Treeshrew; nikobaarientupaija)
| | |-- T. (T.) splendidula Gray, 1865 (Ruddy Treeshrew; punahäntätupaija)
| | |-- T. (T.) javanica Horsfield, 1822 (Horsfield's Treeshrew; indonesiantupaija)
| | |-- T. (T.) gracilis Thomas, 1893 (Slender Treeshrew; hoikkatupaija)
| | |-- T. (T.) picta Thomas, 1892 (Painted Treeshrew; tummaselkätupaija)
| | `-- T. (T.) dorsalis Schlegel, 1857 (Striped Treeshrew; juovatupaija)
| `--o (Lyonogale) Conisbee, 1953
| |-- T. (L.) montana Thomas, 1892 (Mountain Treeshrew; vuoritupaija)
| |-- T. (L.) minor Günther, 1876 (Pygmy Treeshrew; kääpiötupaija)
| |-- T. (L.) tana Raffles, 1821 (Large Treeshrew; maatupaija)
| `-- T. (L.) palawanensis Thomas, 1894 (Palawan Treeshrew; palawanintupaija)
`--+-- Anathana ellioti (Waterhouse, 1850) Lyon, 1913 (Madras Treeshrew, Indian Treeshrew; intiantupaija)
`--+-- Urogale everetti (Thomas, 1892) Mearns, 1905 (Mindanao Treeshrew, Philippine treeshrew; filippiinientupaija)
`--o Dendrogale Gray, 1848 (Smooth-tailed Treeshrews; pehmeähäntätupaijat)
|-- D. murina (Schlegel & Muller, 1843) (Northern Smooth-tailed Treeshrew; pohjanpehmeähäntätupaija)
`-- D. melanura (Thomas, 1892) (Bornean Smooth-tailed Tree Shrew; etelänpehmeähäntätupaija)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Mein P. & Ginsburg L., 1997: Les mammifères du gisement miocène inférieur de Li Mae Long, Thaïlande : systématique, biostratigraphie et paléoenvironnement.
–Geodiversitas: Vol. 19, #4, pp. 783-844
- McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631
- Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii
- Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629