After McKenna & Bell, 1997

<==o Scandentia Wagner, 1855 (puupäästäiset)
   |?- †Nycticonodon caparti Eoc. Eu.
   `--o Tupaiidae Gray, 1825 (tree shrews; puupäästäiset)
      |-- Ptilocercus lowii Gray, 1848 [Ptilocercinae Lyon, 1913] (feather-tail tree shrew, pen-tailed treeshrew; sulkahäntätupaija)
      `--o Tupaiinae Gray, 1825 (true tree shrews; aitotupaijat)
         |-- †Eodendrogale parvum Yongsheng, 1988 [non? Tong, 1988]
         |--o †Prodendrogale Qiu, 1986
         |  |-- †P. yunnanica Qiu, 1986
         |  `-- †P. engesseri Ni & Qiu, 2012
         |--o Tupaia Raffles, 1821
         |  |?- †T. miocenica Mein & Ginsburg, 1997
         |  |?- T. chrysogaster Miller, 1903 (Golden-bellied Treeshrew, Mentawai Treeshrew; kultavatsatupaija)
         |  |?- T. moellendorffi Matschie, 1898 (Calamian Treeshrew
         |  |--o (Tupaia) Raffles, 1821
         |  |  |-- T. (T.) glis (Diard & Duvaucel, 1820) [Sorex Glis Diard & Duvaucel, 1820] (Common treeshrew; tupaii; tupaija)
         |  |  |?- T. (T.) belangeri Wagner, 1841 [ssp. T. glis] (Northern Treeshrew; belangeritupaija)
         |  |  |?- T. (T.) chinensis [ssp. T. glis] (kiinantupaija)
         |  |  |?- T. (T.) longipes Thomas, 1893 [ssp. T. glis] (Long-footed Treeshrew; pitkäjalkatupaija)
         |  |  |-- T. (T.) nicobarica Zelebor, 1869 (Nicobar Treeshrew; nikobaarientupaija)
         |  |  |-- T. (T.) splendidula Gray, 1865 (Ruddy Treeshrew; punahäntätupaija)
         |  |  |-- T. (T.) javanica Horsfield, 1822 (Horsfield's Treeshrew; indonesiantupaija)
         |  |  |-- T. (T.) gracilis Thomas, 1893 (Slender Treeshrew; hoikkatupaija)
         |  |  |-- T. (T.) picta Thomas, 1892 (Painted Treeshrew; tummaselkätupaija)
         |  |  `-- T. (T.) dorsalis Schlegel, 1857 (Striped Treeshrew; juovatupaija)
         |  `--o (Lyonogale) Conisbee, 1953
         |     |-- T. (L.) montana Thomas, 1892 (Mountain Treeshrew; vuoritupaija)
         |     |-- T. (L.) minor Günther, 1876 (Pygmy Treeshrew; kääpiötupaija)
         |     |-- T. (L.) tana Raffles, 1821 (Large Treeshrew; maatupaija)
         |     `-- T. (L.) palawanensis  Thomas, 1894 (Palawan Treeshrew; palawanintupaija)
         `--+-- Anathana ellioti (Waterhouse, 1850) Lyon, 1913 (Madras Treeshrew, Indian Treeshrew; intiantupaija)
            `--+-- Urogale everetti (Thomas, 1892) Mearns, 1905 (Mindanao Treeshrew, Philippine treeshrew; filippiinientupaija)
               `--o Dendrogale Gray, 1848 (Smooth-tailed Treeshrews; pehmeähäntätupaijat)
                  |-- D. murina (Schlegel & Muller, 1843) (Northern Smooth-tailed Treeshrew; pohjanpehmeähäntätupaija)
                  `-- D. melanura (Thomas, 1892) (Bornean Smooth-tailed Tree Shrew; etelänpehmeähäntätupaija)



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